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ZOOTAXA
3082

Checklist of Nematoda associated with the fishes of Brazil


JOS L. LUQUE1, JULIO C. AGUIAR2, FABIANO M. VIEIRA3, DAVID I. GIBSON4 & CLUDIA PORTES SANTOS5
1

Departamento de Parasitologia Animal, Universidade Federal Rural do Rio de Janeiro, Caixa Postal 74508, Seropdica, RJ, 23851970, Brazil. Corresponding author. E-mail: jlluque@ufrrj.br. 2 Curso de Ps-Graduao em Cincias Veterinrias, Universidade Federal Rural do Rio de Janeiro, Seropdica, RJ, Brazil. 3 Curso de Ps-Graduao em Biologia Animal, Universidade Federal Rural do Rio de Janeiro, Seropdica, RJ, Brazil. 4 Department of Zoology, Natural History Museum, Cromwell Road, London SW7 5BD, UK 5 Laboratrio de Avaliao e Promoo da Sade Ambiental, Instituto Oswaldo Cruz, Fundao Oswaldo Cruz, Av. Brasil 4365, Rio de Janeiro, 21045-900, Brazil.

Magnolia Press Auckland, New Zealand

Accepted by G. Prez-Ponce de Len: 03 Oct. 2011; published: 28 Oct. 2011

JOS L. LUQUE, JULIO C. AGUIAR, FABIANO M. VIEIRA, DAVID I. GIBSON & CLUDIA PORTES SANTOS Checklist of Nematoda associated with the fishes of Brazil (Zootaxa 3082) 88 pp.; 30 cm. 28 Oct. 2011 ISBN 978-1-86977-813-2 (paperback) ISBN 978-1-86977-814-9 (Online edition)

FIRST PUBLISHED IN 2011 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: zootaxa@mapress.com http://www.mapress.com/zootaxa/

2011 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 ISSN 1175-5334 (Print edition) (Online edition)

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LUQUE ET AL.

Table of contents
Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 Materials and methods. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Results . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Key to Adult Nematoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Key to Larval Nematoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Parasite-Host List . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 HostParasite List . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45 Discussion . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76 Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76 References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76

Abstract
Seventy-four genera comprising approximately 140 named and 75 undetermined species of adult and larval nematodes are associated with about 330 named and 50 undetermined species of marine, brackish and freshwater fishes from Brazilian waters. These are listed with information on their hosts, habitat, distribution and records. A key to these parasites at the generic level is provided. Key words: Biodiversity, Nematoda, fish parasites, checklist, key, Neotropics, Brazil

Introduction
The phylum Nematoda represent a taxonomically, ecologically and geographically diverse helminth group, with an enormous number of species, both parasitic and freeliving forms, a huge economic and ecological impact, and a number of individuals estimated at 1022 (Hodda, 2007). Because of the enormous abundance and diversity of the group, only an estimated 210% of the world fauna has been formally described, despite its importance (Hodda 2007). Nematodes represent a conspicuous component of the parasite assemblages in fishes. According to Luque & Poulin (2007), nematodes are the second group, after digeneans, in terms of the number of parasite associations recorded in fishes from the Neotropics. Studies on nematodes parasitic in Brazilian fishes began with the description of Trichocephalus gibbosus Rudolphi [now Oncophora melanocephala (Rudolphi)] by Rudolphi (1819) based on specimens collected from Thunnus thynnus (L.) off the coast of Brazil by Pohl in 1817. The first freshwater records appear to be of Lecanocephalus spinulosus Diesing [now Goezia spinulosa (Diesing)] and Cheiracanthus gracilis Diesing [now Gnathostoma gracile (Diesing)], both of which Diesing (1839) recorded from Arapaima gigas (Schinz). However, not until the work of Travassos et al. (1928), which recorded numerous nematodes from freshwater fishes, were detailed studies undertaken. Subequently, a large number of papers by many authors, especially in recent years, have added to our knowledge of this fauna. Compilations of records have been published by Thatcher (1991, 2006), restricted to the Amazon River basin, and by Vicente et al. (1985), supplemented by Vicente & Pinto (1999). Moravec (1998), in his monograph on nematodes parasites of Neotropical freshwater fishes, also included numerous records from Brazil. Some of these compilations have included taxonomic keys, descriptions and/or aspects of the biology of the different taxa. Most recently, Muniz-Pereira et al. (2009) included records of nematode species in a checklist of helminth parasites of threatened vertebrate species from Brazil, and Eiras et al. (2010) published an account of the nematode species associated with Brazilian freshwater fishes. Here we provide a checklist of the nematode species associated with both marine and freshwater fishes from Brazil, including detailed information on locality, habitat, site of infection and life cycle stage. The information is presented as parasite-host and host-parasite lists, with a key to the generic level. We have attempted to include information from all published reports and have added a small number of original records.

NEMATODA IN BRAZILIAN FISHES

Zootaxa 3082 2011 Magnolia Press

Materials and methods

This bibliographic review of the nematode species reported from fishes in Brazilian waters was based on information collected from two main sources. Firstly, searches of the HostParasite Catalogue of the Natural History Museum in London for 1920 to 1997 and, for the years 19882003, the HostParasite Database, a partial implementation of which is now available online (Gibson et al., 2005). Secondly, searches of the Zoological Record, Biological Abstracts and Helminthological Abstracts, Web of Knowledge, Google Scholar, Aquatic Sciences and Fisheries Abstracts, Biological and Agricultural Index Plus and the Scopus databases were undertaken up to July, 2010 (with an update from Web of Knowledge in September, 2011). Original data from our own studies are also included. The checklist follows the parasite classification and systematic arrangement of Moravec (1998, 2006), Anderson et al. (2009) and Gibbons (2010), with some modifications based on recent records. The nematodes are arranged according to class, order, suborder, superfamily, family and subfamily, within which the species are presented in alphabetical order. Host species are given followed by their predominant habitat (marineMAR, freshwaterFW or brackish waterBW) and site of the infection; when possibe these are grouped. Localities are given in alphabetical order of Brazilian states (AMAmazonas, APAmap, BABahia, CECear, ESEsprito Santo, MAMaranho, MGMinas Gerais, MSMato Grosso do Sul, MTMato Grosso, PAPar, PB Paraba, PIPiau, PRParan, RJRio de Janeiro, RORondnia, RNRio Grande do Norte, RSRio Grande do Sul, and SPSo Paulo) preceded by the localities when possibly and, records in chronological sequence. The fish species names were updated according to Froese & Pauly (2010), supplemented by Eschmeyer (2010). The nematode species names have been updated to agree with the recent literature, but inclusion in the parasite or host lists does not imply that the authors necessarily agree with their validity. The species of fish in the hostparasite list are arranged according to the higher level classification of Nelson (2006).

Results
The nematodes parasitic in Brazilian fishes listed below are distributed in 74 genera comprising 142 named and 75 undetermined species of adult and larval nematodes. The total number of host-parasite associations listed is 1,688. These worms were recorded from 324 named marine, brackish and freshwater fish species, and from 50 undetermined host species. Key to Nematoda associated with Brazilian fishes (based on Moravec 1998, Anderson et al. 2009, and Gibbons 2010)

Key to Adult Nematoda


1. 2. 3. 4. 5. 6. Oesophagus cylindrical or forming stichosome or trophosome. Phasmids and deirids absent. Eggs with plug at either pole or hatching in utero .............................................................................................................Class Adenophorea, Order Enoplida (2) Oesophagus never forming stichosome. Phasmids and deirids present. Eggs without polar plugs. Third larva stage infective to final host ..................................................................................................................................................... Class Secernentea (7) Oesophagus cylindrical and well developed; stichosome or trophosome absent. Male tail modified to form ventral sucker ......... . ............................................................................. Superfamily Dioctophymatoidea, Family Dioctophymatidae, Eustrongylides Oesophagus with oesophageal glands forming stichosome or trophosome. Male tail not modified to form ventral sucker........... . ....................................................................................................................Superfamily Trichinelloidea, Family Trichuridae (3) Spicule absent. Male distal end rounded, laterally expanded ............................................................................ Freitascapillaria Spicule present. Male distal end not laterally expanded ...........................................................................................................(4) Spicular sheath with spines . ......................................................................................................................................... Capillaria Spicular sheath without spines .................................................................................................................................................. (5) Pseudobursal membrane absent or very reduced ............................................................................................... Pseudocapillaria Pseudobursal membrane well developed ...................................................................................................................................(6) Dorsolateral projection of pseudobursal margin narrow or digitiform ...................................................................Paracapillaria Dorsolateral projections of pseudobursal margin very wide (in ventral view) .............................................Capillostrongyloides

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7. 8. 9. 10. 11. 12. 13. 14. 15. 16. 17.

18. 19. 20. 21. 22. 23. 24. 25. 26. 27. 28. -

Anterior extremity not bilaterally symmetrical (triradiate). Oesophagus not divided into short anterior muscular part and long posterior glandular part ..............................................................................................................................................................(8) Anterior extremity bilaterally symmetrical. Oesophagus divided, with anterior muscular part shorter than posterior glandular part; division sometimes indistinct ............................................................................................................... Order Spirurida (35) Male with reduced number of caudal papillae. Generally only one spicule. Eggs often flattened on one side.............................. . .................................................................................... Order Oxyurida, Superfamily Oxyuroidea, Family Pharyngodonidae (9) Male with large number of caudal papillae. Generally two spicules. Eggs not often flattened on one side.................................... . .................................................................................................................................................................... Order Ascaridida (15) Oesophageal bulb without sclerotized apparatus .......................................................................................................... Travnema Oesophageal bulb with sclerotized apparatus ..........................................................................................................................(10) Corpus of oesophagus conspicuously dilate posteriorly .........................................................................................................(11) Corpus of oesophagus almost cylindrical, not markedly dilate posteriorly ........................................................................... (12) Buccal capsule present, armed with basal teeth ..................................................................................................... Cosmoxynema Buccal capsule absent........................................................................................................................................Cosmoxynemoides Vulva anterior to middle region of body ................................................................................................................................ (13) Vulva posterior or in middle region of body (excluding tail) ..................................................................................... Ichthyouris Lateral alae present. Buccal capsule present. Eggs with filaments .........................................................................................(14) Lateral alae absent. Buccal capsule absent. Eggs without filaments ................................................................ Parasynodontisia Buccal capsule small, without teeth. Male genital cone well developed. Male lateral alae terminate posterior to cloaca............... . .....................................................................................................................................................................................Spinoxyuris Buccal capsule well developed, with three teeth. Male genital cone poorly developed. Male lateral alae terminate anterior to cloaca ......................................................................................................................................................................... Brasilnema With three well-defined lips, sometimes separated by interlabia. Eggs thick-shelled and not embryonated when deposited. Precloacal sucker absent in male ....................................................................................................... Superfamily Ascaridoidea (17) Small lips present or absent, when present variable in number and form. Eggs hatch in utero or eggs with delicate shells deposited by female. Pre-cloacal sucker may be present in male ...........................................................................................(16) Oesophagus composed of a cylindrical corpus, with distinct pharyngeal part, subspherical or elongate isthmus and valvate bulb . ................................................................................................................................................ Superfamily Cosmocercoidea (26) Oesophagus short, simple or divided into two parts with or without same diameter. Isthmus and bulb absent............................... . ........................................................................................................................................................ Superfamily Seuratoidea (31) Oesophagus with oblong to cylindrical posterior ventriculus, or with small globular ventriculus giving off one or two posteriorly directed appendices. Excretory pore situated between base of subventral lips or near nerve ring ................................ . ..................................................................................................................................................................Family Anisakidae (18) Oesophagus lacking ventriculus or with globular to ellipsoidal ventriculus from which two anteriorly and three posteriorly directed appendices may arise. Excretory pore situated near nerve ring. Family Ascarididae, Multicaecinae, Brevimulticaecum Cuticular annules with posterior spines ......................................................................................... Subfamily Goeziinae, Goezia Cuticular annules without spines .............................................................................................................................................(19) Excretory pore usually between subventral lips or at base of ventral interlabium ............................ Subfamily Anisakinae (20) Excretory pore opens at the level of nerve ring .......................................................................................................................(21) Ventriculus very elongate ........................................................................................................................................ Pulchrascaris Ventriculus slightly elongate .........................................................................................................................................Terranova Head with low lips. In elasmobranchs .......................................................................Subfamily Pseudanisakinae, Pseudanisakis Head with well-developed lips. In teleosts...............................................................................Subfamily Raphidascaridinae (22) Lips surrounded posteriorly by ring with small anteriorly directed curves between lips .......................................Lappetascaris Lips not surrounded posteriorly by ring ...................................................................................................................................(23) Interlabia absent ..................................................................................................................................................... Raphidascaris Interlabia present .............................................................................................................................................................(24) Intestinal caecum absent ................................................................................................................................Raphidascaroides Instestinal caecum present ...................................................................................................................................................... (25) Cuticle with overlapping annulations giving serrate appearance but without spination ........................................ Iheringascaris Cuticle without overlapping annulations ........................................................................................................... Hysterothylacium Oviparous. Didelphic ............................................................................................................................. Family Kathlaniidae (27) Viviparous with larvae in utero at advanced stage of development and capable of endogenous development. Monodelphic....... . ...................................................................................................................................................................Family Atractidae (30) Pharyngeal part of oesophagus armed with anterior, hexagonal sclerotized ring ................................................. Chabaudinema Pharyngeal part of oesophagus simple or armed with three small teeth .................................................................................(28) Anterior extremity of oesophagus differentiated into pharyngeal region ................................................................... Falcaustra Anterior extremity of oesophagus simple, not differentiated into pharyngeal region ............................................................(29)

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29. 30. 31. 32. 33. 34. 35. 36. 37. 38. 39. 40. 41.

42. 43. 44. 45. 46.

Cephalic extremity with six lips. Cephalic region continuous with body proper. Oesophageal isthmus not expanded. Postcloacal lobe-like formation with horns absent .......................................................................................................... Spectatus Cephalic extremith with three lips. Cephalic region separated by transverse cuticular fold from wider portion of body. Oesophageal isthmus expanded. Postcloacal lobe-like formation with two horns present ........................................ Myleusnema Cephalic end armed with eight Y-shaped sclerotized pieces. .................................................................................. Klossinemella Cephalic end without sclerotized pieces ........................................................................................................................ Rondonia Buccal capsule absent. Buccal cavity, when present, lined by cuticle (cheilostome) similar to that of body wall......................... . ..............................................................................................................................................................Family Quimperiidae (32) Pseudobuccal capsule present, formed as modification of oesophagus (oesophastome) with wall surrounded by oesophageal tissue ..................................................................................................................................................... Family Cucullanidae (33) Cephalic region simple, without longitudinal bands of inflated cuticle. Anterior end of oesophagus with bulbous inflation. Male without precloacal muscle bands .....................................................................................................................Paraseuratum Cephalic region with numerous longitudinal bands of inflated cuticle. Anterior end of oesophagus not inflated. Male with precloacal muscle bands .....................................................................................................................................Neoparaseuratum Intestinal caecum present ............................................................................................................................................... Dichelyne Intestinal caecum absent ..........................................................................................................................................................(34) Mouth not surrounded by membranous collarette with denticles. Male tail conical, without small terminal caudal alae............... . ..................................................................................................................................................................................... Cucullanus Mouth surrounded by membranous collarette with denticles. Male tail short and rounded, with small caudal alae. .................... .................................................................................................................................................................................Neocucullanus Pseudolabia absent. Buccal capsule present or not. Oesophagus divided into muscular and glandular portions or muscular throughout. Larvae without cephalic hooks ....................................................................................... Suborder Camallanina (36) Pseudolabia present. Buccal capsule present, usually tubular. Muscular and glandular portions of oesophagus well differentiated. Larvae usually with cephalic hooks or spines.................................................................. Suborder Spirurina (47) Buccal capsule and buccal cavity well developed. Buccal capsule orange-brown, simple or with two valves; tridents may be present. Parasites of digestive tract.........................................................Superfamily Camallanoidea, Family Camallanidae (37) Buccal capsule reduced or absent; buccal cavity poorly developed; Buccal capsule colourless, rounded, not divided into valves; tridents absent. Not usually parasites of digestive tract ................................................Superfamily Dracunculoidea (40) Buccal capsule divided into two lateral valves ........................................................................................................................(38) Buccal capsule rounded, continuous, not separated into two valves .......................Subfamily Procamallaninae, Procamallanus Buccal cavity reduced or absent ..................................................................................................... Subfamily Camallaninae (39) Buccal capsule of adults subdivided into two almost equal parts, with large buccal cavity present................................................ . ..............................................................................................................................Subfamily Paracamallaninae, Paracamallanus Longitudinal bands supporting buccal valves transformed posteriorly into two rows of spines or several combs .... Oncophora Longitudinal bands supporting buccal valves continuous .......................................................................................... Camallanus Vulva in gravid females atrophied or poorly developed (except Alinema), situated in middle to posterior part of body. Oesophagus without free posterior glandular appendix .......................................................................................................... (41) Vulva in gravid females well developed, situated in anterior part of body. Oesophagus with free posterior glandular appendix. .. ............................................................................................................................................................ Family Guyanemidae (42) Oesophagus divided into anterior muscular and posterior glandular portions or undivided. Oesophageal gland multinucleate. Anterior end of oesophagus never inflated. Tail of both sexes conical, pointed............................................................................. ....................................................................................................Family Micropleuridae, Subfamily Micropleurinae, Philonema Oesophagus short, undivided or with large unicellular dorsal oesophageal gland containing large cell nucleus; anterior end of oesophagus often bulbously inflated. Tail of both sexes blunt, sometimes rounded ........................... Family Philometridae (43) Oesophagus with very long glandular portion, several times longer than muscular portion, without free posterior appendage..... . ............................................................................................................................................ Subfamily Guyaneminae, Guyanema Oesophagus with short glandular portion, not longer than muscular portion, with free posterior appendage.................................. . ............................................................................................................................. Subfamily Travassosneminae, Travassosnema Gravid females with atrophied vagina and vulva. Oral aperture unarmed......................................Subfamily Philometrinae (44) Gravid females with well-developed vagina and vulva situated near posterior extremity. Oral aperture with small, sclerotized formations supporting peribucal rim ...........................................................................................Subfamily Alineminae, Alinema Body surface with cuticular ornamentations ........................................................................................................................... (45) Body surface without cuticular ornamentations ......................................................................................................................(46) Cuticle with cone-shaped protuberances. Oesophagus cylindrical, not swollen at anterior end. Caudal end pointed ... Nilonema Cuticle without cone-shaped protuberances, but with small bosses often present in oesophageal region of body. Oesophagus usually bulbous or somewhat swollen near mouth. Caudal end rounded or blunt ................................................. Philometroides Caudal end conical. Cephalic end with apical outgrowths. Oesophagus cylindrical anteriorly, not swollen near mouth, markedly expanded posteriorly ........................................................................................................................................... Rumai

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47. 48. 49.

50. 51. 52. 53. 54. 55. 56. 57. -

Caudal end rounded or blunt, sometimes with pair of papilla-like caudal projections. Cephalic end without apical outgrowths. Oesophagus usually has bulbous inflation at anterior end ............................................................................................Philometra Buccal cavity well cuticularized ............................................................................................................................................. (48) Buccal cavity poorly cuticularized .......................................................................................................................................... (49) Oral opening hexagonal or oval. Pseudolabia absent. Internal labial papillae well developed........................................................ . ............................................................................................................. Superfamily Thelazioidea, Family Rhabdochonidae (50) Oral opening with well-defined bilateral symmetry. Pseudolabia present. Internal labial papillae small or absent........................ .. ..............................................................................................................................................Superfamily Habronematoidea (51) Pseudolabia trilobed. Cephalic and outer labial papillae separate. Cuticle on inner face of each pseudolabium thick, generally folded into rounded, tooth-like formations which fit into corresponding folds on adjacent pseudolabium..................................... . .................................................................................................Superfamily Gnathostomatoidea, Family Gnathostomatidae (55) Pseudolabia unlobed. Cephalic and outer labial papillae fused. Variable number of teeth on free borders of pseudolabia. Body cuticle immediately posterior to pseudolabia, often expanded to form collarete.... Superfamily Physalopteroidea, Family Physalopteridae, Subfamily Proleptinae ....................................................................(57) Buccal capsule armed with longitudinal sclerotized rods projecting anteriorly as teeth. Pre-anal papillae numerous................... .. ................................................................................................................................................................................ Rhabdochona Buccal capsule without sclerotized rods. Oral opening unarmed. Pre-anal papillae not numerous.................Johnstonmawsonia Median lips (ventral and dorsal) present, extending over lateral pseudolabia. Cephalic papillae on lateral pseudolabia. Oesophagus simple. Tail of female modified as attachment organ ..................................................Family Hedruridae, Hedruris Median lips (ventral and dorsal) not extending over lateral pseudolabia. Cephalic papillae never on lateral pseudolabia. Oesophagus divided. Tail of female not in form of attachment organ ................................................ Family Cystidicolidae (52) Pseudolabia relatively large, without teeth, but with enlarged anterior border covering greater part of oral opening............(53) Pseudolabia very small, each sometimes armed with one or two teeth; anterior border slightly enlarged and not covering greater part of oral opening ......................................................................................................................................................(54) Body without rows of spines. Pharynx generally long...........................................................................................Cystidicoloides Anterior region of body with rows of spines or combs. Pharynx generally short ......................................................... Spinitectus Pseudolabia each with single tooth. Vulva post-equatorial.......................................................................................... Ascarophis Pseudolabia each with armed with two teeth. Vulva anterior ................................................................................. Parascarophis Cephalic bulb absent ................................................................................................Subfamily Ancyracanthinae, Ancyracanthus Cephalic bulb present ............................................................................................................... Subfamily Gnathostomatinae (56) Body unarmed ...................................................................................................................................................... Echinocephalus Body partly or entriely armed with posteriorly directed spines ............................................................................... Gnathostoma Vulva near anus .............................................................................................................................................................. Proleptus Vulva slightly pre-equatorial ....................................................................................................................................... Heliconema

Key to Larval Nematoda


1. 2. 3. 4. 5. 6. 7. 8. 9. 10. 11. Oral aperture surrounded by 12 or 18 papillae arranged in two or three circles of six papillae each . . . . . . . . . . . Eustrongylides Oral aperture without these characteristics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . (2) Oral aperture without pseudolabia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . (3) Oral aperture provided with pseudolabia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .(11) Ventriculus with several short anterior and posterior appendices . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Brevimulticaecum Ventriculus without appendices or with only one, usually long, posterior appendix . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .(4) Anterior intestinal caecum present and posterior ventricular appendix absent. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . (5) Anterior intestinal caecum and posterior ventricular appendix present, or both caecum and appendix absent . . . . . . . . . . . . .(7) Excretory pore at about level of nerve ring or just posterior to it . . . . . . . . . . . . . . . . . . . . . . . . . . . . Porrocaecum-like larvae Excretory pore near ventral interlabium . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . (6) Tail with mucron . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseudoterranova Tail without mucron . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .Terranova Cuticular rings of older larvae provided with numerous small posterior spines . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Goezia Cuticular rings always lack spines . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .(8) Both posterior ventricular appendix and anterior intestinal caecum present . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .(9) Only posterior ventricular appendix or only anterior intestinal caecum present, or both absent . . . . . . . . . . . . . . . . . . . . . . .(10) Excretory pore at level of nerve ring or just posterior . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hysterothylacium Excretory pore at base of ventral interlabium . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Contracaecum Posterior ventricular appendix and anterior intestinal caecum absent. Excretory pore at base of ventral interlabium . . Anisakis Posterior ventricular appendix present; anterior intestinal caecum absent. Excretory pore at or posterior to level of nerve ring . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Raphidascaris Mouth provided with two large, lateral, triangular pseudolabia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Spiroxys Cephalic end bears helmet-like cuticular structure with thickened, free posterior region . . . . . . . . . . . . . . . . . Pseudoproleptus

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Parasite-Host List
Nematoda gen. sp.adult and larva Hosts, and site of infection: Ageneiosus inermis (FWadultintestine), Aphyocharax anisitsi (FW adultintestine), Astronotus ocellatus (FWunspecifiedunspecified), Astyanax bimaculatus (FW unspecifiedunspecified), A. eigenmanniorum (FWunspecifiedunspecified), Brycon microleps (= B. hilarii) (FWunspecifiedunspecified), B. orbignyanus (FWlarva), Brycon sp. (FWunspecified unspecified), Catathyridium jenynsii (FWunspecifiedunspecified), Choanoscolex abscissus [Cestoda] in Pseudoplatystoma corruscans (FWlarvaintestine), Coryphaena hippurus (MARunspecified unspecified), Crenicichla britskii (FWadult, larvaintestine), C. niederleinii (FWadultintestine), Cyphocharax gilbert (FWunspecifiedunspecified), C. nagelii (FWunspecifiedunspecified), Cynopotamus kincaidi (FWunspecifiedunspecified), Cynoscion leiarchus (MARunspecified unspecified), Diplodus argenteus argenteus (MARunspecifiedunspecified), Epinephelus sp. (MAR unspecifiedunspecified), Galeocerdo cuvier (MARunspecifiedstomach), Genypterus brasiliensis (MARunspecifiedunspecified), Geophagus brasiliensis (FWunspecifiedunspecified), Gymnotus carapo (FWunspecifiedunspecified), Hemisorubim platyrhynchos (FWlarvaunspecified), Hoplerythrinus unitaeniatus (FWadultintestine), Hoplias malabaricus (FWunspecified unspecified), Hypophthalmus edentatus (FWadultintestine), Hypostomus regani (FWunspecified unspecified), Iheringichthys labrosus (FWadultintestine), Leporellus vittatus (FWadultintestine), Leporinus elongatus (FWunspecifiedunspecified), L. friderici (FWadultintestine), L. lacustris (FWadultintestine), L. obtusidens (FWadultintestine), L. octofasciatus (FWunspecified unspecified), Leporinus sp. (FWadultintestine), Lophiosilurus alexandri (FWadultintestine), Lophius gastrophysus (MARunspecifiedunspecified), Loricariichthys platymetopon (FWadult intestine), Megalonema platanum (FWunspecifiedunspecified), Menticirrhus americanus (MAR unspecifiedunspecified), Micropogonias furnieri (MARunspecifiedunspecified), Myctoperca sp. (MARunspecifiedunspecified), Myletes sp. (FW unspecifiedunspecified), Myleus levis (FW unspecifiedunspecified), Oreochromis niloticus niloticus (FWlarvaintestine), Pachyurus squamipennis (MARunspecifiedunspecified), Pagrus pagrus (MARunspecifiedunspecified), Pellona castelnaeana (FWunspecifiedunspecified), Piaractus brachypomus (FWunspecified unspecified), Pimelodus maculatus (FWunspecifiedunspecified), Pinirampus pirinampu (FWadult, larvaintestine), Plagioscion squamosissimus (FWlarvamesentery), Pomatomus saltator (MAR unspecifiedunspecified), Potamorhina squamoralevis (FWunspecifiedunspecified), Potamotrygon motoro (FWunspecifiedunspecified), Priacanthus arenatus (MARunspecifiedunspecified), Proloricaria prolixa (FWunspecifiedunspecified), Pseudopercis numida (MARunspecified unspecified), Pseudoplatystoma sp. (FWunspecifiedunspecified), Pseudupeneus maculatus (MAR unspecifiedunspecified), Rhamphichthys hahni (FW adultintestine), Rhaphiodon vulpinus (FW adultintestine), Rhinodoras dorbignyi (FWadultintestine), Roeboides paranensis (FW unspecifiedunspecified), Satanoperca pappaterra (FW adultintestine), Serrasalmus spilopleura (FWunspecifiedunspecified), Synbranchus marmoratus (FW adultintestine), Tetragonopterus argenteus (FWunspecifiedunspecified), Thyrsitops lepidopoides (MARunspecifiedunspecified), Triportheus angulatus (FWunspecifiedunspecified), T. nematurus, T. paranaensis (FW unspecifiedunspecified). Localities: PentecosteCE; off ES; So Francisco RiverMG; MT; Itaip reservoir, Maring, Saraiva Lagoon, Upper Paran RiverPR. References: Travassos (1940a, b, 1945), Travassos & Freitas (1941, 1942, 1943, 1948), , Rgo & Gibson (1989), Pavanelli et al. (1997, 2004), Brasil-Sato (2003), Kohn et al. (2003, 2004), Arci et al. (2006), Graa & Machado (2007), Takemoto et al. (2009). Remark: Choanoscolex abscissus [Cestoda] in Pseudoplatystoma corruscans (larvaintestine). This record from a cestode represents a case of hyperparasitism.

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Class Adenophorea Linstow Order Enoplida Schuurmans, Stekhoven & Deconinck Superfamily Dioctophymatoidea Railliet Family Dioctophymatidae Castellani & Chalmers
Eustrongylides ignotus Jgerskildadult and larvaFW Hosts and site of infection: Auchenipterus nigripinnis (larvabody cavity), Brycon hilarii (larvabody cavity), Galeocharax humeralis (larvabody cavity), Hemisorubim platyrhynchos (larvamesentery, muscle), Hoplias aff. malabaricus, H. malabaricus (larvabody cavity), Hoplias sp., Loricariidae gen. sp., Paratrygon sp. (larvabody cavity), Pinirampus pirinampu (larvabody cavity), Poeciliidae gen. sp. (larvabody cavity), Pseudoplatystoma corruscans (adult and larvabody cavity, intestine), P. fasciatum (larvabody cavity), Pygocentrus nattereri (larvabody cavity), Rhaphiodon vulpinus (larvabody cavity), Salminus brasiliensis (larvabody cavity), Serrasalmus maculatus (larvaunspecified), S. marginatus (larvabody cavity), S. spilopleura (larvabody cavity), Synbranchus marmoratus (larva body cavity). Localities: Cuiab RiverMT; CachimboPA; Upper Paran RiverPR; Rio de JaneiroRJ; PirassunungaSP. References: Travassos (1925), Rgo & Eiras (1988), Rgo & Vicente (1988a, b), Eiras & Rgo (1988, 1989), Pavanelli et al. (1997, 2004), Moravec (1998), Vicente & Pinto (1999), Guidelli et al. (2003), Takemoto et al. (2009), Eiras et al. (2010), Rocha (2011). Eustrongylides tubifex Nitzsch in RudolphilarvaFW Syn.: E. perpapillatus Jgerskild Hosts and site of infection: Pseudoplatystoma corruscans (mesenteries). Locality: Piracicaba RiverSP. Reference: Lordello & Monteiro (1959). Eustrongylides sp.larvaFW Hosts and site of infection: Acestrorhynchus lacustris (intestine), Arapaima gigas (tumour on skin), Auchenipterus nigripinnis (muscle), Brycon hilarii (body cavity), B. microleps, (mesentery, muscle), Callichthys callichthys, Calophysus macropterus, Charax gibbosus, Cichla kelberi, C. monoculus, C. ocellaris (body cavity), C. piquiti, Galeocharax humeralis (mesenteries), Gymnotus carapo, Hoplerythrinus unitaeniatus (mesentery, muscle, stomach wall), Hoplias malabaricus (body cavity, liver serosa, mesentery, muscle, stomach wall), Leporinus copelandii (mesenteries), L. obtusidens (body cavity), Lepthoplosternum pectorale, Oligosarcus sp. (body cavity), Paratrygon sp., Pimelodus maculatus (body cavity), Pirinampus pirinampu (mesentery, muscle), Plagioscion squamosissimus, Poecilia vivipara, Pseudoplatystoma corruscans (intestine, mesenteries), P. fasciatum (mesenteries), Pseudoplatystoma sp., Pygocentrus nattereri (body cavity, mesentery, muscle, swim bladder), Rhaphiodon vulpinus, Salminus brasiliensis (body cavity, mesenteries), Steindachneridion parahybae (mesenteries), Synbranchus marmoratus, Tetragonopterinae gen. sp. Localities: So Francisco RiverMG; Cuiab River, PantanalMS, MT; Maicur, Mexiana Island, Amazonas River BasinPA; Upper Paran RiverPR; Agua, Paran River, PirassunungaSP. References: Travassos et al. (1928), Lordello & Monteiro (1959), Travassos & Freitas (1964), Travassos & Kohn (1965), Vicente et al. (1985), Kohn & Fernandes (1987), Rgo & Vicente (1988b), Thatcher (1991), Brasil-Sato (2003), Carvalho et al. (2003), Santos et al. (2003), Adriano et al. (2005), Barros et al. (2006, 2007, 2009a, 2010), Saraiva et al. (2006a), Martins et al. (2009), Muniz-Pereira et al. (2009), Santos & Moravec (2009b), Takemoto et al. (2009), Eiras et al. (2010). Eustrongylides sp. 1larva FW Hosts and site of infection: Gymnotus sp. (mesenteries). Locality: Baa RiverMS. Reference: Isaac et al. (2004).

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Eustrongylides sp. 2larva FW Hosts and site of infection: Gymnotus sp. (mesenteries). Locality: Baa RiverMS. Reference: Isaac et al. (2004).

Superfamily Trichinelloidea Railliet Family Trichuridae (Ransom)


Capillaria (Neocapillaria) carioca Freitas & LentadultMAR Syn.: Echinocephalus carioca Lopes Neyra; Thominx carioca Skrjabin & Shikhobalova Hosts and site of infection: Sphoeroides testudineus (intestine). Locality: Rio de JaneiroRJ. References: Freitas & Lent (1935), Yamaguti (1961), Vicente & Pinto (1981), Vicente et al. (1985), Moravec (2001). Capillaria sp.adult Hosts and site of infection: Astyanax fasciatus (FWintestine), Hypostomus derbyi (FWintestine), Leporellus vittatus (FWintestine), Leporinus copelandii (FWintestine), Menticirrhus americanus (MARintestine), Pseudupeneus maculatus (MARmusculature) Pseudopercis numida (MAR intestine), P. semifasciata (MARintestine), Serrasalmus marginatus (FWintestine). Localities: Igua River Basin, Upper Paran RiverPR; off Rio de JaneiroRJ; PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985), Kohn & Fernandes (1987), Kohn et al. (1988), Chaves & Luque (1999), Vicente & Pinto (1999), Pavanelli et al. (2004), Luque et al. (2006), Lima et al. (2008). Capillariidae gen. sp.FW Hosts: Serrasalmus maculatus, S. marginatus, S. spilopleura. Locality: Upper Paran RiverPR. References: Pavanelli et al. (2004), Takemoto et al. (2009). Capillariidae gen. sp. 1adultFW Hosts and site of infection: Schizodon fasciatus (intestine). Locality: Paran River, GuaraPR. References: Moravec et al. (1992a), Vicente & Pinto (1999), Moravec (2001). Remark: Vicente & Pinto (1999) referred this taxon as Trichuridae sp. Moravec (2001) considered this taxon to be a species inquirenda. Capillariidae gen. sp. 2adultFW Hosts and site of infection: Pseudoplatystoma corruscans (intestine). Localitiy: Paran River, GuaraPR. References: Moravec et al. (1992a), Vicente & Pinto (1999), Moravec (2001). Remark: Vicente & Pinto (1999) referred this taxon as Trichuridae sp. Whereas, Moravec (2001) considered it to be a species inquirenda. Capillariinae gen. sp.adult Hosts and site of infection: Gymnotus carapo (FW), Hoplosternum littorale (FWintestine), Rhamdia quelen (FW unspecified). Locality: Guandu RiverRJ. References: Abdallah et al. (2006), Azevedo et al. (2010). Capillostrongyloides arapaimae Santos, Moravec & VenturieriadultFW Host and site of infection: Arapaima gigas (intestine, pyloric caeca). Locality: Mexiana Island, Amazonas River BasinPA. References: Santos et al. (2008), Eiras et al. (2010). Capillostrongyloides sentinosa (Travassos)adultFW Syn.: Capillaria minima Travassos, Artigas & Pereira; C. zederi Freitas & Lent; C. sentinosa Travassos

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Hosts and site of infection: Anostominae gen. sp. (stomach), Astyanax fasciatus (stomach), Hoplias malabaricus (stomach), Leporinus elongatus (intestine, stomach), Leporinus sp. (stomach), Schizodon nasutus (intestine, stomach). Localities: Upper Paran RiverPR; Grande River, Parnaba, PirassunungaSP. References: Travassos (1927), Travassos et al. (1928), Freitas & Lent (1935), Yamaguti (1961), Travassos & Kohn (1965), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernades (1987), Thatcher (1991), Moravec (1998, 2001), Takemoto et al. (2009), Eiras et al. (2010). Freitascapillaria maxillosa (Vaz & Pereira)adultFW Syn.: Capillaria maxillosa Vaz & Pereira; Pseudocapillaria nuda Mendona Host and site of infection: Salminus brasiliensis (intestine, stomach, pyloric caeca). Locality: Piracicaba River, PirassunungaSP. References: Vaz & Pereira (1934), Freitas & Lent (1935), Yamaguti (1961), Mendona (1963), Travassos & Kohn (1965), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Moravec (1998, 2001), Thatcher (2006), Eiras et al. (2010). Paracapillaria (Paracapillaria) argentinensis Timi, Rossin, Lanfranchi & EtchegoinadultMARintestine Hosts: Pinguipes brasilianus, Pseudopercis numida, P. semifasciata. Localities: Cabo Frio, Rio de JaneiroRJ. References: Luque et al. (2008), Timi et al. (2010). Paracapillaria (Paracapillaria) piscicola (Travassos, Artigas & Pereira)adultFW Syn.: Capillaria piscicola Travassos, Artigas & Pereira Hosts and site of infection: Acestrorhynchus falcatus (stomach), Acestrorhynchus sp. (intestine), Galeocharax humeralis (intestine, stomach), G. knerii (stomach), Hoplias malabaricus (intestine, stomach), Hypostomus affinis (unspecified), Leporellus vittatus (stomach), Leporinus copelandii (stomach), Oligosarcus sp. (stomach), Salminus brasiliensis (stomach), S. hilarii (stomach), Trachelyopterus striatulus (stomach). Localities: Amazonas River Basin; Foz do Igua, Paran and Upper Paran RiverPR; PirassunungaSP. References: Travassos et al. (1928), Freitas & Lent (1935), Yamaguti (1961), Mendona (1963), Travassos & Kohn (1965), Buhrheim (1976), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Moravec et al. (1992a), Moravec (1998, 2001), Pavanelli et al. (2004), Thatcher (2006), Takemoto et al. (2009), Azevedo et al. (2010), Eiras et al. (2010), Mesquita et al. (2011). Pseudocapillaria (Pseudocapillaria) magalhaesi (Lent & Freitas)adultMARbody cavity and intestine Syn.: Trichosoma da corvina of Lent & Freitas (1937); Capillaria magalhaesi Lent & Freitas; Pseudocapillaria (Ictiocapillaria) magalhaesi (Lent & Freitas) Hosts: Micropogonias furnieri, M. undulatus. Localities: Rio de JaneiroRJ; Cassino BeachRS. References: Lent & Freitas (1937), Vicente et al. (1985), Alves & Luque (1999, 2000, 2001a, 2001b), Moravec (2001), Almeida et al. (2008), Luque et al. (2010).

Class Secernentea Dougherty Order Ascaridida Skrjabin & Schulz Superfamily Ascaridoidea Baird
Ascaridoidea fam. gen. sp.larvaFW Hosts: Electrophorus electricus, Hoplias malabaricus, Pseudoplatystoma sp., Salminus brasiliensis. Localities: SalobraMG; MaicurPA. Reference: Travassos (1940a), Travassos & Freitas (1964), Vicente et al. (1985). Filocapsulariinae gen. sp.adultFWswim bladder Hosts: Galeocharax knerii, Hoplias malabaricus, Leporellus vittatus, Leporinus copelandii, L. octofasciatus, L. striatus, Leporinus sp., Salminus hilarii.
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Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985).

Family Anisakidae Railliet & Henry


Anisakidae gen. sp.larva Hosts and site of infection: Acestrorhynchus lacustris (FWunspecified), Auchenipterus nigripinnis (FWmesentery), Brycon orbignyanus (FWmesentery), Conorynchus conirostris (FWmesentery), Genypterus brasiliensis (MARunspecified), Katsuwonus pelamis (MAR unspecified), Micropogonias furnieri (MARunspecified), Pagrus pagrus (MARlateral muscle), Prochilodus lineatus (FW unspecified), Pseudopercis numida (MAR unspecified), Pseudoplatystoma corruscans (FW unspecified), P. fasciatum (FWunspecified), Salminus brasiliensis (FWmesentery). Localities: So Franciso RiverMG; PantanalMT; Upper Paran RiverPR; off Rio de JaneiroRJ. References: Rgo & Vicente (1988b), Pavanelli et al. (1997, 2004), Brasil-Sato & Santos (2005), Pereira Cardoso et al. (2006), Saad & Luque (2009), Takemoto et al. (2009).

Subfamily Anisakinae Railliet & Henry


Anisakis physeteris BaylislarvaMARmesentery Hosts: Auxis thazard thazard, Genypterus brasiliensis. Localities: Brazilian coast; Niteri, Rio de JaneiroRJ. References: Knoff et al. (2007), Iiguez et al. (2009). Anisakis typica (Diesing)larvaMARintestine Syn.: Cercocephalis typicus Diesing Hosts: Auxis thazard thazard, Thunnus thynnus. Locality: Brazilian coast. References: Mattiucci et al. (2002), Mattiucci & Nascetti (2006), Iiguez et al. (2009). Anisakis simplex (Rudolphi)larvaMAR Hosts and site of infection: Genypterus brasiliensis (mesentery, stomach serosa, intestinal serosa, ovary, musculature), Paralichthys isosceles (abdominal cavity, intestine, stomach serosa). Locality: Niteri, Rio de JaneiroRJ. References: Knoff et al. (2007), Felizardo et al. (2009a). Anisakis pegreffii Campana-Rouget & BioccalarvaMAR Host: Thunnus thynnus. Locality: Brazilian coast. References: Mattiucci & Nascetti (2006). Anisakis sp.larvaMAR Hosts and site of infection: Aluterus monoceros (mesentery), Bagre bagre (mesentery), Balistes vetula (mesentery), Brevoortia aurea (mesentery), Caranx latus (mesentery), Carcharhinus signatus (spiral valve), Dipturus trachydermus (spiral valve), Euthynnus alletteratus (mesentery), Galeorhinus galeus (spiral valve), Genypterus brasiliensis (intestinal serosa, mesentery, muscles, stomach), Heptranchias perlo (spiral valve), Hexanchus griseus (spiral valve), Katsuwonus pelamis, Merluccius hubbsi (mesentery), Micropogonias furnieri (mesentery), Mullus argentinae (mesentery), Mustelus canis (spiral valve), Nebris microps (mesentery), Pagrus pagrus (liver, mesentery, lateral muscle), Paralichthys isosceles (mesentery), Percophis brasiliensis (mesentery), Pomatomus saltatrix (intestine), Prionotus punctatus (liver, mesentery), Pseudopercis numida (mesentery), Scomber japonicus (mesentery), S. scombrus (mesentery),

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Scomberomorus cavalla (stomach serosa), Selene setapinnis (mesentery), Sphyraena guachancho (mesentery), Squalus megalops (spiral valve), Squatina sp. (spiral valve), Trichiurus lepturus (coelomic serosa, mesentery), Urophycis mystacea (mesentery). Localities: FortalezaCE; PR; Rio de JaneiroRJ; RS. References: Rgo et al. (1983), Rgo & Santos (1983), Vicente et al. (1985), Barros & Amato (1993), SoClemente et al. (1994), Marques et al. (1995), Luque & Chaves (1999), Luque et al. (2000a, 2002, 2008, 2010), Silva et al. (2000a,b), Paraguass et al. (2000, 2002), Knoff et al. (2001, 2003a,b, 2007), Luque & Alves (2001), Abdallah et al. (2002), Alves et al. (2002c, 2003, 2005), , Cordeiro & Luque (2004), Luque & Poulin (2004), Tavares et al. (2004b), Bicudo et al. (2005a,b), Padovani et al. (2005), Alves & Luque (2006), Tavares & Luque (2006), Muniz-Pereira et al. (2009), Saad & Luque (2009), Dias et al. (2010, 2011). Anisakis sp. 1larvaMARmesentery Host: Katsuwonus pelamis. Locality: Rio de JaneiroRJ. References: Alves & Luque (2006). Anisakis sp. 2larvaMARmesentery Host: Katsuwonus pelamis. Locality: Rio de JaneiroRJ. References: Alves & Luque (2006). Contracaecum sp.adult and larva Hosts and site of infection: Oligosarcus macrolepis (FWlarvabody cavity), Acestrorhampus sp. (= Oligosarcus sp.) (FWunspecifiedunspecified), Acestrorhynchus lacustris (FWlarvamesentery), Ageneiosus inermis, Aluterus monoceros (MARlarvamesentery), Astronotus ocellatus (FWlarva mesentery), Astyanax altiparanae (FWlarvabody cavity, mesentery), A. bimaculatus (FWlarva body cavity, mesentery), A. fasciatus (FWlarvabody cavity, mesentery), A. schubarti (FWlarva body cavity, mesentery), Balistes capriscus (MARlarvamesentery), B. vetula (MARlarva mesentery), Bergiaria sp. (FWlarvabody cavity), Brevoortia aurea (MARlarvamesentery), Brycon microleps (= B. hilarii) (FWlarvamesentery), Caranx hippos (MARlarvamesentery), C. latus (MARlarvamesentery), Carcharhinus brachyurus (MARlarvaspiral valve), C. signatus (MARlarvaspiral valve), Centropomus undecimalis (MARlarvamesentery), Cephalopholis fulva (MARlarvamesentery), Cichla kelberi (FWlarvaunspecified), C. monoculus (FWlarva mesentery), C. ocellaris (FWlarvamesentery), Crenicichla lepidota (FWlarvabody cavity), Curimata sp. (FWlarvabody cavity), Dactylopterus volitans (MARlarvamesentery), Diapterus rhombeus (MARlarvamesentery), Dipturus trachydermus (MARlarvastomach, spiral valve), Euthynnus alletteratus (MARlarvamesentery), Galeocharax humeralis, G. knerii (FWlarvabody cavity), Galeorhinus galeus (MARlarvaspiral valve), Genidens barbus (MARlarvamesentery), Genypterus brasiliensis (MARlarvamesentery), Geophagus brasiliensis (FWlarvabody cavity, mesentery), Gymnotus carapo (FWlarva), Heptranchias perlo (MARlarvaspiral valve), Hemisorubim platyrhynchos (FWlarvaunspecified), Hexanchus griseus (MARlarvaspiral valve), Hoplerythrinus unitaeniatus (FWadultmesentery, muscle), Hoplias malabaricus (FWlarvabody cavity, intestinal, liver, stomach serosa, mesentery, muscle), Iheringichthys labrosus (FWlarva unspecified), Leporellus vittatus (FWlarvabody cavity), Leporinus copelandii (FWlarvabody cavity), L. elongatus (FWlarvamesentery), L. friderici (FWlarvamesentery), L. lacustris (FW larvabody cavity, mesentery), L. obtusidens (FWlarvamesentery), Loricariichthys castaneus (FW unspecifiedunspecified), Macrodon ancylodon (MARlarvamesentery), Menticirrhus americanus (MARlarvaintestine), Metynnis lippincottianus (FW unspecifiedunspecified), Micropogonias furnieri (MARlarvamesentery), Mullus argentinae (MARlarvamesentery), Mustelus canis (MARlarvaspiral valve), M. schmitti (MARlarvaspiral valve), Oligoplites palometa (MAR

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larvagills), Oligosarcus paranensis, Oligosarcus sp. (FWlarvabody cavity), Oligoplites saliens (MARlarvagills), O. saurus (MARlarvagills), Pagrus pagrus (MARlarvalateral muscle, liver and mesentery), Paralichthys isosceles (MARlarvamesentery and intestine), Paralonchurus brasiliensis (MARlarvamesentery), Parona signata (MARlarvamesentery), Paulicea luetkeni (= Zungaro zungaro) (FWlarvamesentery), Peprilus paru (MARunspecifiedunspecified), Percophis brasiliensis (MARlarvamesentery), Pimelodella lateristriga (FW unspecifiedunspecified), Pimelodus maculatus (FWlarvamesentery), P. ortmanni (FWlarvabody cavity), Pirinampus pirinampu (FWlarvamesentery), Plagioscion squamosissimus (FWlarvamesentery), Pomatomus saltatrix (MARlarvaintestine), Priacanthus arenatus (MARlarvamesentery), Prochilodus lineatus (FWlarvabody cavity, mesentery), Pseudoplatystoma corruscans (FWlarvamesentery), P. fasciatum (FWlarvamesentery), Psellogrammus kennedyi (FWlarvaunspecified), Pygocentrus nattereri (FWlarvabody cavity, mesentery, swim bladder), Rhamdia quelen (FWlarvamesentery), Rhaphiodon vulpinus (FWlarvaunspecified), Salminus brasiliensis (FWlarvabody cavity, mesentery), S. hilarii (FWlarvamesentery), Sarda sarda (MARlarvamesentery), Schizodon nasutus (FWlarvabody cavity), Scomber japonicus (MARlarvaintestine, mesentery, pyloric caeca, swim bladder), Scomberomorus brasiliensis (MARlarvamesentery) S. cavalla (FWunspecified mesentery), Scyliorhinus haeckelii (MARlarvaspiral valve), Selene setapinnis (MARlarva mesentery), Serrasalmus altuvei (FWunspecifiedunspecified), S. marginatus (FWlarva mesentery), Sphyrna zygaena (MARlarvaspiral valve), Squatina sp. (MARlarvaspiral valve), Tetragonopterus argenteus (FW unspecifiedunspecified), Trachelyopterus striatulus (FW unspecifiedunspecified), Trachurus lathami (MARlarvamesentery), Trichiurus lepturus (MAR larvacoelomic serosa, mesentery), Urophycis brasiliensis (MARlarvamesentery), Urophycis mystacea (MARlarvamesentery). Localities: Amazonas River Basin; off So LuizMA; Cuiab River, PantanalMT; Igua River, Upper Paran RiverPR; Campos, Guandu River, Lajes Reservoir, Pira, off Rio de JaneiroRJ; Campinas, Jaguari Reservoir, Paran RiverSP; MS; Paran River, PirassunungaSP; of RS. References: Travassos & Freitas (1940), Kloss (1966), Vicente & Fernandes (1978), Fbio (1982), Rgo et al. (1983), Rgo & Santos (1983), Vicente et al. (1985), Kohn et al. (1988, 1989), Rgo & Vicente (1988b), Eiras & Rgo (1989), Leo et al. (1991), Thatcher (1991), Barros & Amato (1993), Barros (1994), SoClemente et al. (1994, 1995), Marques et al. (1995), Takemoto et al. (1996, 2009), Pavanelli et al. (1997, 2004), Alves & Luque (1999, 2000, 2001a, 2001b, 2006), Chaves & Luque (1999), Luque & Chaves (1999), Vicente & Pinto (1999), Luque et al. (2000a, 2002, 2003, 2010), Machado et al. (2000), Paraguass et al. (2000, 2002, 2005), Silva et al. (2000a,b), Knoff et al. (2001, 2007), Luque & Alves (2001), Tavares et al. (2001, 2004b), Abdallah et al. (2002), Alves et al. (2002a,b,c, 2003, 2004, 2005), Ribeiro et al. (2002), Brasil-Sato (2003), Carvalho et al. (2003, 2010), Martins et al. (2003, 2005), Cordeiro & Luque (2004, 2005b), Luque & Poulin (2004), Tavares & Luque (2004a,b, 2006), Lizama et al. (2005), Madi & Silva (2005), Azevedo et al. (2006, 2007a,c, 2010), Paraguass & Luque (2006), Barros et al. (2007, 2009a,b, 2010), Felizardo et al. (2009a), Moreira et al. (2009, 2010), Muniz-Pereira et al. (2009), Saad & Luque (2009), Dias et al. (2010, 2011), Eiras et al. (2010), Mesquita et al. (2011), Vicentin et al. (2011), Rocha (2011). Contracaecum sp. 1LarvaFW Hosts and site of infection: Gymnotus sp. (mesentery), Hemisorubim platyrhynchos (mesentery), Pseudoplatystoma corruscans (unspecified). Localities: Upper Paran RiverPR; Baia RiverMS. References: Machado et al. (1996), Isaac et al. (2004), Pavanelli et al. (2004). Contracaecum sp. 2LarvaFWmesentery Hosts: Gymnotus sp., Hemisorubim platyrhynchos, Pseudoplatystoma corruscans. Localities: Upper Paran RiverPR; Baia RiverMS.

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References: Machado et al. (1996), Isaac et al. (2004), Pavanelli et al. (2004). Contracaecum sp. Type AFW Unspecified site of infection Hosts: Astyanax bimaculatus, A. schubarti, Galeocharax knerii, Salminus brasiliensis, S. hilarii. Locality: PirassunungaSP. Reference: Bhrnheim (1976). Contracaecum sp. Type BFW Unspecified site of infection Hosts: Astyanax bimaculatus, Galeocharax knerii, Hoplias malabaricus, Leporinus copelandii, L. lacustris, Salminus brasiliensis. Locality: PirassunungaSP. Reference: Bhrnheim (1976). Contracaecum sp. Type 1LarvaFW Hosts and site of infection: Acestrorhynchus lacustris (intestine), Astyanax altiparanae (mesentery), A. fasciatus (body cavity), Brycon hilarii (liver, mesentery), Cichla kelberi, Cichla piquiti, Crenicichla lepidota (body cavity), Galeocharax humeralis (intestine, body cavity), G. knerii (intestine, body cavity), Gymnotus carapo (body cavity), Hemisorubim platyrhynchos, Hoplerythrinus unitaeniatus (mesentery), Hoplias malabaricus (body cavity and mesentery), Leporinus friderici (gonads, mesentery, musculature of intestine), L. lacustris (gonads, mesentery, musculature of intestine), Plagioscion squamosissimus (body cavity), Pseudoplatystoma corruscans, P. fasciatum (intestine), Rhaphiodon vulpinus (body cavity). Localities: Juba RiverMT; Aquidauna RiverMG; PantanalMS; Foz do Igua, Guaira, Itaip Reservoir, Paran River, Upper Paran RiverPR; AguaSP. References: Moravec et al. (1993a), Moravec (1998), Carvalho et al. (2003), Guidelli et al. (2003, 2006), Saraiva et al. (2006a,b), Campos et al. (2008, 2009b,c), Eiras et al. (2010). Contracaecum sp. Type 2LarvaFW Hosts and site of infection: Acestrorhynchus lacustris (intestine), Astronotus ocellatus, Astyanax fasciatus (body cavity), Brycon hilarii (liver, mesentery), Gymnotus carapo (body cavity), Hemisorubim platyrhynchos (mesentery), Hoplerythrinus unitaeniatus (body cavity), Hoplias malabaricus (body cavity), Markiana geayi (body cavity), Pachyurus bonariensis (mesentery), Plagioscion ternetzi (mesentery), Prochilodus lineatus (body cavity), Pseudoplatystoma corruscans (body cavity), P. fasciatum (body cavity), Pygocentrus cariba (body cavity), Rhamdia quelen, Rhaphiodon vulpinus, Salminus hilarii, S. brasiliensis (unspecified). Localities: MA; Juba RiverMT; Foz do Igua, Guaira, Itaip Reservoir, Paran RiverPR; Sinha Mariana LagoonMG. References: Moravec et al. (1993a), Moravec (1998), Carvalho et al. (2003), Guidelli et al. (2003), Martins et al. (2005), Saraiva et al. (2006b), Lacerda et al. (2009), Eiras et al. (2010). Pseudoterranova decipiens (Krabbe)larvaMAR Host and site of infection: Genypterus brasiliensis (mesentery, stomach serosa, stomach). Locality: Niteri, Rio de JaneiroRJ. Reference: Knoff et al. (2007). Pseudoterranova sp. larvaMAR Hosts and site of infection: Galeorhinus galeus (spiral valve), Genypterus brasiliensis (intestine, mesentery), Mustelus canis (spiral valve), M. schmitti (spiral valve), Pagrus pagrus (mesentery), Squalus megalops (spiral valve), Trichiurus lepturus (mesentery). Locality: Niteri, Rio de JaneiroRJ; RS. References: Paraguass et al. (2000, 2002), Silva et al. (2000a,b), Knoff et al. (2001, 2007), Luque & Alves (2001), Muniz-Pereira et al. (2009). Pulchrascaris caballeroi Vicente & SantosadultMARstomach Host: Squatina squatina. Locality: MacaRJ.

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References: Vicente & Santos (1972, 1973), Vicente et al. (1985). Terranova rochalimai (Pereira)adultMARstomach Host: Cao Panan (Carcharhinus longimanus ?). Locality: NatalRN . References: Pereira (1935), Yamaguti (1961), Vicente et al. (1985). Terranova serrata (Drasche)adultFWintestine Syn.: Ascaris serrata Drasche; A. draschei Stossich; Porrocaecum draschei (Stossich) Host: Arapaima gigas. Localities: Amazon fish farmAM; Amazonas RiverPA. References: Drasche (1884a), Baylis (1927), Travasos et al. (1928), Yamaguti (1961), Santos et al. (1979), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Araujo et al. (2009b), Eiras et al. (2010). Terranova trichiuri ChandleradultMAR Hosts and site of infection: Bagre bagre (intestine), Macrodon ancylodon (intestine), Oligoplites saurus (intestine), Squatina squatina (intestine, stomach). Localities: So LuizMA; MacaRJ. References: Vicente & Santos (1973), Vicente & Fernandes (1978), Vicente et al. (1985). Terranova sp.adult and larva Hosts and site of infection: Anchoa tricolor (MARlarvamesentery), Balistes capriscus (MARlarva mesentery), Brevoortia aurea (MARlarvamesentery), Caranx hippos (MARlarvamesentery), C. latus (MARlarvamesentery), Cynoscion guatucupa (MARlarvamesentery), Genypterus brasiliensis (MARadult and larvamesentery and intestine), Gymnothorax moringa (MARlarva mesentery), Micropogonias furnieri (MARadult and larvamesentery), Pagrus pagrus (MARadult liver, mesentery), Paralichthys isosceles (MARadultintestine, liver, mesentery, stomach mucosa), Percophis brasiliensis (MARadultmesentery), Plagioscion squamosissimus (FWlarvamesentery), Pomatomus saltatrix (MARadultintestine), Priacanthus arenatus (MARadultmesentery), Prionotus punctatus (MARadultmesentery), Pseudopercis numida (MARadult, larvamesentery), P. semifasciata (MARadult, larvamesentery), Sarda sarda (MARadultmesentery), Scomber japonicus (MARLarvaintestine, gonads and pyloric caeca), Selene setapinnis (MARadult mesentery), Trachurus lathami (MARadultmesentery), Trichiurus lepturus (MARadult mesentery), Tylosurus acus acus (MARadultmesentery), Umbrina canosai (MARadult mesentery). Localities: off IlhusBA; off FortalezaCE; off Cabo Frio, Rio de JaneiroRJ; off ES; off Cassino BeachRS; off FlorianpolisSC; Araguaia RiverTO. References: Rgo & Santos (1983), Rgo et al. (1983, 1985), Vicente et al. (1985), So-Clemente et al. (1994), Marques et al. (1995), Alves & Luque (1999, 2000, 2001a,b, 2006), Luque & Chaves (1999), Vicente & Pinto (1999), Luque et al. (2000a, 2008, 2010), Silva et al. (2000a), Luque & Alves (2001), Tavares et al. (2001, 2005, 2007), Alves et al. (2002a,b, 2005), Paraguass et al. (2002), Sabas & Luque (2003), Cordeiro & Luque (2004), Luque & Poulin (2004), Bicudo et al. (2005a,b), Timi et al. (2005), Tavares & Luque (2006), Knoff et al. (2007), Felizardo et al. (2009a), Muniz-Pereira et al. (2009), Eiras et al. (2010). Remark: These larval records could represent species of Pseudoterranova, Terranova or other anisakine genera, as is the case with those listed below as Porrocaecum larvae.

Subfamily Goeziinae Travassos


Goezia brasiliensis Moravec, Kohn & FernandesadultFW

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Hosts and site of infection: Brycon hilarii (stomach), Pseudoplatystoma corruscans (intestine). Localities: Foz do Igua, Itaip Reservoir, Paran RiverPR. References: Moravec et al. (1994b), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Eiras et al. (2010). Goezia brevicaeca Moravec, Kohn & FernandesadultFWstomach Hosts: Brycon hilarii, Leporinus friderici. Locality: Foz do Igua, Itaip Reservoir, Paran RiverPR. References: Moravec et al. (1994b), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Guidelli et al. (2006), Takemoto et al. (2009), Eiras et al. (2010). Goezia leporini Martins & YoshitoshiadultFWintestine Hosts: Leporinus macrocephalus, Piaractus mesopotamicus. Locality: BatataisSP. References: Martins & Yoshitoshi (2003), Martins et al. (2004), Eiras et al. (2010). Goezia spinulosa (Diesing)adult and larvaFW Syn.: Lecanocephalus spinulosus Diesing Hosts and site of infection: Acar-a, Arapaima gigas (adult and larvastomach), Astronotus ocellatus (adultstomach), Astyanax schubarti* (adultstomach), Galeocharax knerii* (adultstomach), Hoplias malabaricus* (adultintestine, stomach), Leporellus vittatus* (adultstomach), Leporinus copelandii (adultstomach), L. elongatus, L. friderici (adultintestine, caecum), L. obtusidens (adultintestine), Micropterus salmoides (adultstomach), Pimelodus maculatus* (adultstomach), Pseudoplatystoma corruscans (unspecifiedunspecified), Salminus brasiliensis* (adultstomach). Localities: Amazonas River Basin; Amazon fish farmAM; Araguaia River MT; Posto de Piscicultura Lima CamposCE; MG; Mexiana IslandPA; Upper Paran RiverPR; Rio de JaneiroRJ; Guapor RiverRO; PirassunungaSP. References: Diesing (1839), Travassos et al. (1928), Freitas & Lent (1946), Yamaguti (1961), Rasheed (1965), Travassos & Kohn (1965), Bhrnheim (1976), Santos et al. (1979), Thatcher (1981, 1991, 2006), Vicente et al. (1985), Costa et al. (1995), Moravec (1998), Pavanelli et al. (2004), Guidelli et al. (2006), Tavares & Luque (2006), Santos, C.P. et al. (2008), Araujo et al. (2009b), Santos & Moravec (2009), Takemoto et al. (2009), Eiras et al. (2010), Menezes et al. (2011). Remark: According to Bhrnheim (1976), larval specimens were found in the mesentery, wall of the stomach and intestine in hosts marked with an asterisk. Goezia sp.adult and larvaFW Hosts and site of infection: Ageneiosus militaris (larvastomach), Hemisorubim platyrhynchos (adult intestine), Hoplosternum littorale (adultintestine), Mylossoma duriventre (larvastomach), Pimelodus maculatus (adultintestine), Rhaphiodon vulpinus (larvastomach), Serrasalmus marginatus (larva stomach), Trachelyopterus galeatus (adult, intestinelarva, stomach). Localities: So Francisco RiverMG; Foz do Igua, Guaira, Paran River, Upper Paran RiverPR; Guandu RiverRJ. References: Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999), Brasil-Sato (2003), Guidelli et al. (2003), Pavanelli et al. (2004), Abdallah et al. (2006), Takemoto et al. (2009), Azevedo et al. (2010), Eiras et al. (2010).

Subfamily Pseudanisakinae Petter, Paradiznik, Radujkovic & Cassone


Pseudanisakis sulamericana Santos, Lent & GibsonadultMARspiral valve Hosts: Psammobatis extenta, Rioraja agassizii Locality: UbatubaSP. References: Santos et al. (2004), Muniz-Pereira et al. (2009).
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Subfamily Raphidascaridinae Hartwich


Hysterothylacium fortalezae (Klein)adultMARintestine Syn.: Contracaecum fortalezae Klein Hosts: Harengula clupeola, Scomberomorus cavalla, S. maculatus. Localities: SalvadorBA; FortalezaCE. References: Klein (1973), Guimares & Cristfaro (1974), Vicente et al. (1985). Hysterothylacium eurycheilum (Olsen)adultMARintestine Syn.: Heterotyphlum eurycheilum Olsen Host: Epinephelus guttatus. Locality: Guanabara BayRJ. References: Rodrigues et al. (1971), Vicente et al. (1985). Hysterothylacium sp.larva Syn.: Thynnascaris sp. Hosts and site of infection: Anchoa marinii (MARmesentery), A. tricolor (MARmesentery), Archosargus rhomboidalis (MARintestine), Balistes vetula (MARmesentery), Caranx latus (MAR mesentery), Cichla kelberi (FWunspecified), Crenicichla lepidota (FWbody cavity), Cynoscion guatucupa (MARmesentery), Dactylopterus volitans (MARmesentery), Galeocharax knerii (FW body cavity), Genidens barbus (MARmesentery), Genypterus brasiliensis (MARmesentery), Gymnotus carapo (FWunspecified), Gymnotus sp. (FWmesentery), Gymnothorax moringa (MAR mesentery), Hypophthalmus edentatus (FWbody cavity), Leporinus friderici (FWmesentery, swim bladder), Lophius gastrophysus (MARmesentery), Loricarichthys sp. (FWbody cavity), Macrodon ancylodon (MARmesentery), Menticirrhus americanus (MARmesentery), Merluccius hubbsi (MAR mesentery), Micropogonias furnieri (MARintestine, intestinal caecum, stomach, coelomatic cavity, liver, spleen), Mugil liza (FWunspecified), Pagrus pagrus (MARlateral and ventral muscle, mesentery), Paralichthys isosceles (MARabdominal cavity, abdominal musculature, intestine, heart serosa, kidney serosa, liver serosa, mesentery, ovary, ovarian serosa, stomach, stomach mucosa, spleen serosa), Parona signata (MARmesentery), Peprilus paru (MARmesentery), Percophis brasiliensis (MAR mesentery), Pinguipes brasilianus (MARmesentery), Plagioscion squamosissimus (FWbody cavity), Prionotus punctatus (MARliver, mesentery), Pseudopercis numida (MARmesentery), P. semifasciata (MARmesentery), Pterodoras granulosus (FWbody cavity), Pygocentrus piraya (FWunspecified), Rhaphiodon vulpinus (FWbody cavity), Salminus brasiliensis (FWbody cavity), Selene setapinnis (MARmesentery), Serrasalmus brandtii (FWunspecified), Scomber japonicus (MARunspecified), S. scombrus (MARmesentery), Trachelyopterus striatulus (FWunspecified), Tetragonopterus chalceus (FWunspecified), Trichiurus lepturus (MARmesentery), Triportheus guentheri (FWunspecified), Tylosurus acus acus (MARmesentery), Umbrina canosai (MARmesentery), Uraspis secunda (MAR mesentery), Urophycis brasiliensis (MARunspecified), U. mystacea (MARunspecified). Localities: off IlhusBA; off FortalezaCE; off Angra dos Reis, Cabo Frio, Rio de JaneiroRJ; off Cassino BeachRS; off FlorianpolisSC; off RS; Foz do Igua, Itaip Reservoir, Upper Paran River PR; Volta Grande ReservoirMG. References: Rgo et al. (1985), Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999), Luque et al. (2000a, 2008, 2010), Martins et al. (2000), Silva et al. (2000a,b), Luque & Alves (2001), Alves et al. (2002a,b, 2005), Sabas & Luque (2003), Isaac et al. (2004), Cordeiro & Luque (2004, 2005a,b), Luque & Poulin (2004), Pereira Jr. et al. (2004), Tavares et al. (2004a. 2005), Bicudo et al. (2005a,b), Timi et al. (2005, 2010), Alves & Luque (2006), Azevedo et al. (2007b, 2010), Guidelli et al. (2006), Tavares & Luque (2006, 2008), Knoff et al. (2007), Felizardo et al. (2009a,b), Muniz-Pereira et al. (2009), Saad & Luque (2009), Takemoto et al. (2009), Eiras et al. (2010), Mesquita et al. (2011). Iheringascaris inquies (Linton)adultMARintestine, stomach and pyloric caeca

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Syn.: Ascaris inquies Linton; Contracaecum inquies (Linton); C. (Thynnascaris) inquies (Linton); T. inquies (Linton); I. iheringascaris Pereira; C. iheringascaris (Pereira); Thynnascaris iheringascaris (Pereira) Host: Rachycentron canadum. Locality: NatalRN. References: Pereira (1935), Yamaguti (1961), Travassos et al. (1963), Vicente et al. (1985). Lappetascaris lutjani RasheedadultMARstomach Host: Trachipterus jacksonensis. Locality: Atlantic coastRS. References: Vicente et al. (2002). Raphidascaris (Ichthyascaris) vicentei SantosadultMAR Syn.: R. atlanticus Rodrigues; R. yamagutii Vicente & Santos; R. camura Deardorff & Overstreet Hosts and site of infection: Lutjanus synagris (intestine), Harengula clupeola (intestine), Nebris microps (intestine), Sardinella sp. (intestine, stomach). Localities: SalvadorBA; Guanabara Bay, Maca, MaricRJ. References: Santos (1970), Rodrigues (1974), Vicente & Santos (1974), Guimares et al. (1976), Feij et al. (1979), Vicente et al. (1985), Rodrigues et al. (1990), Luque et al. (2000b). Raphidascaris (Sprentascaris) hypostomi (Petter & Cassone)adultFW Syn: Sprentascaris hypostomi Petter & Cassone Hosts and site of infection: Ancistrus cirrhosus (intestine, stomach), Hypostomus sp. (intestine, stomach), H. albopunctatus (intestine, stomach), H. cochliodon (unspecified), H. commersoni (intestine, stomach), H. derbyi (intestine, stomach), Metynnis lippincottianus (unspecified). Localities: Salto OsrioPR; Passo FundoRS. References: Moravec et al. (1990), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Takemoto et al. (2009), Eiras et al. (2010). Raphidascaris (Sprentascaris) lanfrediae Melo, Giese, Santos, Santos & Portes SantosadultFW Host and site of infection: Satanoperca jurupari (intestine). Localities: Guam RiverPA. References: Melo et al. (2011b). Raphidascaris (Sprentascaris) mahnerti (Petter & Cassone)adultFW Syn: Sprentascaris mahnerti Petter & Cassone Hosts and site of infection: Pseudohemiodon laticeps (intestine), Loricariichthys brunneus, L. labialis, L. platymetopon, Loricariichthys sp. (intestine), Metynnis lippincottianus (unspecified). Localities: Paran RiverPR; Uruguai RiverRS. References: Moravec et al. (1990, 1993a), Fortes et al. (1999), Vicente & Pinto (1999), Thatcher (2006), Moreira et al. (2009, 2010), Takemoto et al. (2009), Eiras et al. (2010). Raphidascaris (Sprentascaris) sp. of Moravec, Kohn & Fernandes (1993)FW Hosts and site of infection: Geophagus brasiliensis (larvaintestine), G. proximus (adultunspecified). Locality: Paran RiverPR. References: Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999), Eiras et al. (2010). Menezes et al. (2011). Raphidascaris sp.adult and larva Hosts and site of infection: Anchoa marinii (MARlarvamesentery), Ancistrus cirrhosus (FWlarva body cavity, intestine), Atlantoraja castelnaui (MARlarvaspiral valve), Balistes capriscus (MAR larvamesentery) Caranx hippos (MARlarvamesentery), Cephalopholis fulva (MARlarva mesentery), Cyphocharax gilbert (FWlarvaintestine), Dactylopterus volitans (MARlarva mesentery), Diapterus rhombeus (MARlarvamesentery), Dipturus trachydermus (MARlarvaspiral valve), Euthynnus alletteratus (MARlarvamesentery), Galeorhinus galeus (MARlarvaspiral valve), Genypterus brasiliensis (MARlarvamesentery), Geophagus proximus (FW unspecified

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unspecified), Gymnothorax moringa (MARlarvamesentery), Harengula clupeola (MARlarva mesentery), Hypostomus albopunctatus (FWunspecifiedintestine, stomach), H. commersoni (FW unspecifiedintestine, stomach), H. derbyi (FWunspecifiedintestine, stomach), Lophius gastrophysus (MARlarvamesentery), Merluccius hubbsi (MARlarvamesentery), Micropogonias furnieri (MARadult and larvamesentery), Mullus argentinae (MARlarvamesentery), Nebris microps (MARlarvamesentery, pyloric caeca), Pagrus pagrus (MARlarvalateral and ventral musculature, liver, mesentery), Paralichthys isosceles (MARlarvaabdominal cavity, intestine, liver, mesentery, stomach, stomach serosa), Parona signata (MARlarvamesentery), Peprilus paru (MARlarva mesentery, pyloric caeca), Percophis brasiliensis (MARlarvaintestine, pyloric caeca, mesentery), Pinguipes brasilianus (MARadultgills), Pomatomus saltatrix (MARlarvamesentery), Priacanthus arenatus (MARlarvamesentery), Prionotus punctatus (MARlarvaliver, mesentery), Prochilodus lineatus (FWlarvaintestine), Pseudopercis numida (MARunspecifiedmesentery), P. semifasciata (MARunspecifiedmesentery), Sarda sarda (MARlarvamesentery), Sardinha Boca Larga (MARintestine, pyloric caeca), Sardinha Maromba (MARunspecifiedintestine), Scomber colias (MARlarvaintestine), Scomber japonicus (MARlarvaintestine, mesentery, pyloric caeca, stomach), Scomber scombrus (MARlarvamesentery), Scombroides occidentalis* (MARlarva intestine), Scyliorhinus haeckelii (MARlarvaspiral valve), Selene setapinnis (MARlarva mesentery), S. vomer (FWlarvaintestine, pyloric caeca), Sphyraena guachancho (MARlarva mesentery), Thyrsitops lepidopoides (MARlarvaintestine, pyloric caeca), Trichiurus lepturus (MAR larvamesentery), Umbrina canosai (MARlarvamesentery), Uraspis secunda (MARlarva mesentery), Urophycis brasiliensis (MARlarvamesentery). Localities: Amazonas River Basin; IlhusBA; FortalezaCE; Igua River Basin, Upper Paran River PR; Cabo Frio, Guandu River, Maca, Niteri, Rio de JaneiroRJ; FlorianpolisSC; coastPR; RS. References: Vicente & Santos (1973, 1974), Rgo & Santos (1983), Rgo et al. (1983, 1985), Vicente et al. (1985), Kohn et al. (1988, 1989), Thatcher (1991), Vicente & Pinto (1999), Luque et al. (2000a, 2002, 2008), Silva et al. (2000a,b), Knoff et al. (2001, 2007), Luque & Alves (2001), Tavares et al. (2001), Abdallah et al. (2002, 2005), Paraguass et al. (2000, 2002), Alves et al. (2003, 2004, 2005), Cordeiro & Luque (2004, 2005b), Luque & Poulin (2004), Bicudo et al. (2005a,b), Lizama et al. (2005), Alves & Luque (2006), Tavares & Luque (2006), Azevedo et al. (2007b, 2010), Felizardo et al. (2009a), Muniz-Pereira et al. (2009), Saad & Luque (2009), Eiras et al. (2010), Timi et al. (2010). Remark: Asterisk indicates a host not listed in FishBase (Froese & Pauly, 2010); junior synonym of Oligoplites saurus (Berg). Raphidascaroides brasiliensis Moravec & ThatcheradultFWintestine Host: Pterodoras granulosus. Locality: Janauac Lake, AM. References: Moravec & Thatcher (1997c), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Eiras et al. (2010).

Family Ascarididae Baird


Ascarididae gen. sp.FW - Unspecified site of infection Hosts: Astyanax fasciatus (larva), Brachyplatystoma sp. (unspecified), Potamotrygon motoro (unspecified), Hoplias malabaricus (unspecified), Loricaria sp. (larva), Rhaphiodon vulpinus (unspecified), Pygocentrus nattereri (unspecified), Salminus brasiliensis (unspecified), Siluriformes fam. gen. sp. (unspecified) Localities: Barra Seca River, Cupido River, Juruparan LagoonES; SalobraMT; MaicurPA. References: Travassos & Freitas (1942, 1943, 1948, 1964), Vicente et al. (1985).

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Subfamily Angusticaecinae Skrjabin & Karokhin


Amplicaecum sp.FW Hosts and site of infection: Brycon orthotaenia (larvaintestine, swim bladder wall), Hoplias malabaricus (unspecifiedintestine), Leporinus copelandii (unspecifiedintestine), Pseudoplatystoma sp. (larva swim bladder wall), Salminus hilarii (unspecifiedintestine), Steindachneridion parahybae (larva intestine, swim bladder). Locality: PirassunungaSP. References: Travassos et al. (1928), Travassos & Kohn (1965), Vicente et al. (1985), Kohn & Fernandes (1987), Vicente & Pinto (1999), Muniz-Pereira et al. (2009). Angusticaecum sp.larvaMARintestine Host: Centropomus parallelus Locality: Rio de JaneiroRJ References: Lamothe-Argumedo et al. (1996)

Subfamily Multicaecinae Hartwich


Brevimulticaecum regoi SprentadultFWintestine Host: Potamotrygon motoro. Locality: SalobraMS, MT. References: Rgo (1979), Sprent (1990), Bruce et al. (1994), Moravec (1998), Eiras et al. (2010). Brevimulticaecum sp.adult and larvaFW Hosts and site of infection: Gymnotus inaequilabiatus*, Hemisorubim platyrhynchos*, Hoplias malabaricus*, Leporinus elongatus (larvamesentery), L. friderici (larvamesentery), L. lacustris (larvamesentery), L. obtusidens (larvamesentery), Myleus levis*, Platydoras costatus (adult intestine), Potamotrygon falkneri (larvastomach), P. motoro (stomach), Pseudoplatystoma corruscans*, Pseudoplatystoma sp. (adultintestine), Pygocentrus nattereri*, P. piraya (adultintestine), Serrasalmus marginatus (larva - stomach tissue, mesentery), Sorubim lima*. Localities: Miranda River, Pantanal, SalobraMS; Upper Paran RiverPR. References: Travassos et al. (1939), Rgo (1979), Vicente et al. (1985), Lacerda et al. (2009), Takemoto et al. (2009), Eiras et al. (2010), Vieira et al. (2010), Vicentin et al. (2011). Remarks: Asterisk indicates that the parasites were in the body cavity, liver, mesentery, muscles or stomach wall. Brevimulticaecum sp. larva of Moravec, Prouza & Royero (1997)FW Hosts and site of infection: Leporinus friderici (musculature of stomach and intestine, mesentery, adipose tissue), L. lacustris (musculature of stomach and intestine, mesentery, adipose tissue), Potamotrygon falkneri (external wall of stomach). Locality: Upper Paran RiverPR. References: Guidelli et al. (2006), Lacerda et al. (2008). Subfamily Toxocarinae (Hartwich) Remark: Toxocarines are typically parasites of terrestrial higher vertebrates and the Brazilian records of Porrocaecum spp. are doubtful (Moravec, 1994, 1998). So, all records of Porrocaecum spp. from fishes are likely to be misidentifications of other ascaridiod, probably anisakid taxa, and, thus, can be considered as Porrocaecum-like species. Porrocaecum jardimfreire ForteslarvaFWintestine Hosts: Cathorops agassizii, Genidens barbus, Genidens genidens.

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Locality: Guaba LakeRS. References: Fortes (1981), Vicente et al. (1985), Fortes & Hoffmann (1995). Remark: Considered a species inquirenda by Moravec (1998). Porrocaecum paivai Motta & GomesadultMARintestine Host: Scomberomorus cavalla. Locality: FortalezaCE. References: Mota & Gomes (1968), Vicente et al. (1985). Porrocaecum secundum ChandlerlarvaMARintestine Host: Oligoplites saurus, Lanceta. Locality: MacaRJ. References: Vicente & Santos (1973). Porrocaecum sp.larva Hosts and site of infection: Chaetodipterus faber (MARintestine), Franciscodoras marmoratus (FW unspecified), Hoplias malabaricus (FWunspecified), Lanceta(MAR?unspecified), Leporinus elongatus (FWmesentery), Piaractus mesopotamicus (FWmesentery), Platydoras costatus (FW intestine), Selene vomer (MARintestine). Localities: UberabaMG; Miranda RiverMS; Upper Paran RiverPR; off MacaRJ. References: Travassos et al. (1939), Vicente & Santos (1973), , Vicente et al. (1985), Martins & Onaka (2004), Pavanelli et al. (2004), Takemoto et al. (2009), Eiras et al. (2010).

Superfamily Cosmocercoidea Railliet Family Atractidae Railliet


Klossinemella iheringi (Travassos, Artigas & Pereira)adultFWintestine, pyloric caeca Syn.: Monhysterides iheringi Travassos, Artigar & Pereira; Klossiella iheringi (Travassos, Artigas & Pereira) Hosts: Hoplias malabaricus, Leporinus copelandii, L. fasciatus, Mylesinus paraschomburgkii, Myleus asterias, Myleus sp., Piaractus brachypomus, Pterodoras granulosus, Salminus hilarii, Schizodon nasutus, Tetragonopterinae gen. sp. Localities: Capucapu, Pitinga, Uatuma RiversAM; Araguari RiverAP; Cuiab River, Paraguai River Basin, So Loureno RiverMT; Trombetas River, Jari RiverPA; PirassunungaSP. References: Travassos et al. (1928), Costa (1960, 1961), Yamaguti (1961), Travassos & Kohn (1965), Costa et al. (1968), Vicente & Pinto (1981, 1999), Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006), Moravec & Thatcher (1997b), Moravec (1998), Eiras et al. (2010). Klossinemella sp.adultFW Syn.: Monhysterides sp. Host: Pimelodus maculatus. Locality: Upper Paran RiverPR. References: Takemoto et al. (2009), Eiras et al. (2010). Rondonia rondoni Travassosadult and larvaFWintestine Hosts: Acanthodoras spinosissimus (unspecified), Luciopimelodus pati (unspecified), Mylesinus paraschomburgkii (unspecified), Myletes sp. (adult), Myleus asterias (adult), M. micans (adult), M. torquatus (adult), Myleus sp. (adult), Oxydoras kneri (unspecified), Piaractus brachypomus (adult), P. mesopotamicus (adult and larva), Pimelodus maculatus (unspecified), Pterodoras granulosus (adult), Salminus sp. (unspecified), Synodontis clarias (unspecified), Trachydoras paraguayensis (adult), Zungaro zungaro (unspecified). Localities: Barra and Grande RiversBA; So Francisco River, Trs Marias ReservoirMG; Aquidauana, Miranda and Paran River, Porto CabralMS; Paraguai and So Loureno Rivers, PantanalMT;

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Amazonas, Cachimbo and Trombetas RiversPA; Porto Rico, Paran River, Upper Paran RiverPR; Porto Cabral, Paran River, Pirassununga, Tiet RiverSP. References: Travassos (1920, 1923, 1945), Travassos et al. (1928, 1939), Baylis (1936), GllegoBerenger (1947), Yamaguti (1961), Costa (1962, 1963), Travassos & Kohn (1965), Santos et al. (1979, 2003), Vicente & Pinto (1981, 1999), Kohn et al. (1985), Kohn & Fernandes (1987), Rgo & Vicente (1988b), Thatcher (1991, 2006), Moravec et al. (1992a), Martins & Urbinati (1993), Parra et al. (1997), Pavanelli et al. (1997, 2004), Moravec & Thatcher (1997b), Moravec (1998), Brasil-Sato & Santos (2003), Brasil-Sato (2003), Dias et al. (2004), Martins et al. (2004), Takemoto et al. (2004, 2009), Campos et al. (2007, 2009a), Eiras et al. (2010). Rondonia sp.FW Hosts and site of infection: Myletes sp. (unspecifiedunspecified), Myleus sp. (unspecifiedintestine), Piaractus brachypomus (unspecifiedunspecified). Localities: MaicurPA; Mato Grosso; SalobraMS. References: Travassos (1940b), Travassos & Freitas (1943, 1964), Vicente et al. (1985).

Family Cosmocercidae Railliet


Cosmocercinae gen. sp.adultMARintestine Host: Sardinella sp. Locality: Rio de JaneiroRJ. References: Feij et al. (1979), Vicente et al. (1985).

Family Kathlaniidae Lane


Chabaudinema americanum Daz-UngraadultFW - intestine Host: Piaractus mesopotamicus Locality: Aquidauana and Miranda River, Porto CabralMS. References: Campos et al. (2009a), Eiras et al. (2010). Falcaustra mirandafroesi (Fortes)adultBWintestine Syn.: Spironoura mirandafroesi Fortes Hosts: Cathorops agassizii, Genidens barbus, Genidens genidens. Locality: Guaba LakeRS. References: Fortes (1981), Vicente et al. (1985), Fortes & Hoffmann (1995), Moravec (1998). Remark: This species was considered to be a species inquirenda by Moravec (1998). Myleusnema brasiliense Moravec & ThatcheradultFWintestine Host: Myleus sp. Locality: Tocantins RiverPA. References: Moravec & Thatcher (1999), Thatcher (2006), Eiras et al. (2010). Spectatus spectatus TravassosadultFWintestine Hosts: Myletes sp., Piaractus brachypomus, P. mesopotamicus, Pterodoras granulosus. Localities: Porto Esperana, Porto So Joo, So Loureno River and PantanalMT; Miranda River, Salobra and PantanalMS; Paran River and Porto CabralSP. References: Travassos (1923, 1945), Travassos et al. (1928, 1939), Yamaguti (1961), Vicente & Pinto (1981), Vicente et al. (1985), Rgo & Vicente (1988b), Moravec (1998), Santos et al. (2003), Thatcher (2006), Campos et al. (2009a), Eiras et al. (2010). Spectatus sp.FWintestine

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Hosts and site of infection: Mylossoma duriventre (adult), Piaractus mesopotamicus (adult, juvenile). Locality: Foz do Igua, Paran RiverPR. References: Moravec et al. (1997), Moravec (1998), Vicente & Pinto (1999).

Superfamily Seuratoidea Hall


Seuratoidea gen. sp. larva of Moravec, Kohn & Fernandes (1997)FWstomach Host: Zungaro zungaro. Locality: Guara, Paran RiverPR. References: Moravec et al. (1997), Moravec (1998), Vicente & Pinto (1999).

Family Cucullanidae Cobbold


Cucullanus (Cucullanus) brevispiculus Moravec, Kohn & FernandesFWintestine Host: Auchenipterus nuchalis, Leporinus copelandii. Locality: Itaip ReservoirPR, Guandu River - RJ. References: Moravec et al. (1993a, 1997), Moravec (1998), Vicente & Pinto (1999), Moreira et al. (2000), Kohn et al. (2003), Thatcher (2006), Azevedo et al. (2010), Eiras et al. (2010). Cucullanus (Cucullanus) grandistomis (Ferraz & Thatcher)adultFWintestine Syn.: Bacudacnitis grandistomis Ferraz & Thatcher Host: Oxydoras niger, Mugil liza. Locality: Amazonas RiverAM, Guandu River - RJ, Solimes River AM. References: Ferraz & Thatcher (1988), Thatcher (1991, 2006), Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999), Azevedo et al. (2010), Eiras et al. (2010), Silva et al.(2011) Cucullanus (Cucullanus) mogi TravassosadultFWintestine Hosts: Leporinus copelandii, Leporinus sp. Locality: PirassunungaSP. References: Travassos (1947, 1948), Yamaguti (1961), Travassos & Kohn (1965), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Moravec (1998), Thatcher (2006), Eiras et al. (2010). Cucullanus (Cucullanus) oswaldocruzi Santos, Vicente & JardimadultFWintestine Host: Zungaro zungaro. Locality: Curu, Cachimbo RiversPA. References: Santos et al. (1979), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Cucullanus (Cucullanus) pauliceae Vaz & PereiraadultFWintestine Host: Zungaro zungaro. Locality: Piracicaba RiverSP. References: Vaz & Pereira (1934), Yamaguti (1961), Vicente & Pinto (1981), Vicente et al. (1985), Moravec (1998), Thatcher (1991, 2006), Eiras et al. (2010). Cucullanus (Cucullanus) pimelodellae Moravec, Kohn & FernandesadultFWintestine Host: Pimelodella lateristriga. Locality: Guaira, Paran RiverPR. References: Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Eiras et al. (2010). Cucullanus (Cucullanus) pinnai pinnai Travassos, Artigas & PereiraadultFW

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Hosts and site of infection: Ageneiosus militaris (intestine), Galeocharax knerii (unspecified), Leporinus copelandii (intestine), Loricaria sp. (intestine), Luciopimelodus pati (unspecified), Megalonema platanum (unspecified), Pimelodella gracilis (intestine), Pimelodus albicans (unspecified), P. maculatus (intestine), P. ornatus (intestine), Pimelodus sp. (unspecified), Pseudopimelodus mangurus (unspecified), Pseudoplatystoma corruscans (unspecified), P. fasciatum (unspecified), Pseudoplatystoma sp. (unspecified), Rhamdia quelen (intestine) Schizodon borelli (unspecified), Steindachneridion parahybae (intestine), Synodontis clarias (unspecified), Zungaro zungaro (unspecified). Localities: PantanalMS; Paran River, Upper Paran RiverPR; PirassunungaSP; Aquidauna River MG; Guandu River - RJ. References: Travassos et al. (1928), Yamaguti (1961), Travassos & Kohn (1965), Bhrnheim (1976), Vicente & Pinto (1981, 1999), Kohn et al. (1985), Vicente et al. (1985), Kohn & Fernandes (1987), Moravec et al. (1993a, 1997), Moreira et al. (1994), Machado et al. (1994, 1995, 1996), Pavanelli et al. (1997, 2004), Moravec (1998), , Brasil-Sato (2003), Thatcher (2006), Santos et al. (2007), Albuquerque et al. (2008), Campos et. al. (2008, 2009c), Muniz-Pereira et al. (2009), Takemoto et al. (2009), Azevedo et al. (2010), Eiras et al. (2010), Venancio et al. (2010). Cucullanus (Cucullanus) pinnai pterodorasi Moravec, Kohn & FernandesadultFWintestine Host: Pterodoras granulosus. Locality: Guaira, Itaip ReservoirPR. References: Moravec et al. (1997), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Eiras et al. (2010), Silva et al. (2011). Cucullanus (Cucullanus) pseudoplatystomae Moravec, Kohn & FernandesadultFWintestine Hosts: Pseudoplatystoma corruscans, Pseudoplatystoma fasciatum. Localities: PantanalMS; Upper Paran RiverPR; Aquidauna RiverMG. References: Moravec et al. (1993a), Machado et al. (1994, 1995, 1996), Pavanelli et al. (1997, 2004), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Campos et al. (2008, 2009c), Takemoto et al. (2009), Eiras et al. (2010) Cucullanus (Cucullanus) zungaro Vaz & PereiraadultFWintestine Hosts: Hemisorubim platyrhynchos, Luciopimelodus pati, Zungaro zungaro. Localities: Foz do Igua, Upper Paran RiverPR; Piracicaba and Tiet RiversSP. References: Vaz & Pereira (1934), Yamaguti (1961), Vicente & Pinto (1981, 1999), Thatcher (1991,2006), Moravec et al. (1997), Moravec (1998), Guidelli et al. (2003), Pavanelli et al. (2004), Takemoto et al. (2009). Cucullanus (Cucullanus) sp.adultFWspiral valve Host: Potamotrygon falkneri. Locality: Upper Paran RiverPR. Reference: Lacerda et al. (2009). Cucullanus ageneiosus Giese, Furtado, Lanfredi & SantosadultFWintestine Host: Ageneiosus ucayalensis. Locality: Guajar Bay, BelmPA. Reference: Giese et al. (2010). Cucullanus carioca Vicente & FernandesadultMAR intestine Hosts: Pinguipes brasilianus, Pseudopercis numida, P. semifasciata. Locality: Guanabara Bay, Rio de JaneiroRJ. References: Vicente & Fernandes (1978), Vicente et al. (1985), Luque et al. (2008), Timi et al. (2010). Cucullanus cassinensis Pereira Jr. & CostaadultMARintestine Host: Micropogonias furnieri. Locality: Coast, Patos LagoonRS. References: Pereira Jr. & Costa (1996), Vicente & Pinto (1999), Pereira Jr. et al. (2002).

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Remark: Considered to be a species inquirenda by Timi et al. (2009). Cucullanus cirratus MlleradultMARintestine Host: Micropogonias undulatus, Urophycis mystacea. Locality: Maca, Rio de JaneiroRJ. References: Vicente & Santos (1973), Vicente et al. (1985), Alves et al. (2002c). Cucullanus debacoi Sarmento, Fortes & HoffmannadultFWintestine Host: Pimelodus maculatus. Locality: Guaba Lake, Porto AlegreRS. References: Fortes & Hoffmann (1995), Sarmento et al. (1995), Vicente & Pinto (1999). Remark: Considered to be a species inquirenda by Moravec (1998). Cucullanus dodsworthi BarretoadultMARintestine Host: Sphoeroides testudineus. Locality: Guanabara Bay, Rio de JaneiroRJ. References: Barreto (1922), Yamaguti (1961), Vicente & Pinto (1981), Vicente et al. (1985). Cucullanus fabregasi Fortes, Hoffmann & SarmentoadultFWintestine Host: Pimelodus maculatus. Locality: Guaba Lake, Porto AlegreRS. References: Fortes et al. (1993b), Fortes & Hoffmann (1995), Vicente & Pinto (1999). Remark: Considered to be a species inquirenda by Moravec (1998). Cucullanus genypteri Sardella, Navone & TimiadultMARintestine Host: Genypterus brasiliensis. Locality: Rio de JaneiroRJ. References: Alves et al. (2001a, 2002a,b). Cucullanus heliomartinsi Moreira, Rocha & CostaadultFWintestine Host: Trachelyopterus striatulus. Locality: Central Lake, Doce River State ParkMG. References: Moreira et al. (2000), Eiras et al. (2010). Cucullanus patoi Fortes, Hoffmann & SarmentoadultFWintestine Host: Pimelodus maculatus. Locality: Guaba LakeRS. References: Fortes et al. (1992), Fortes & Hoffmann (1995), Vicente & Pinto (1999). Remark: Considered to be a species inquirenda by Moravec (1998). Cucullanus pulcherrimus BarretoadultMARintestine Hosts: Caranx hippos, C. lugubris, Micropogonias furnieri, Mugil platanus, Paralonchurus brasiliensis. Localities: Trindade IslandES; Marambaia Island, Rio de JaneiroRJ; Coast and Patos LagoonRS. References: Barreto (1918, 1922), Yamaguti (1961), Vicente et al. (1985), Pinto et al. (1992), Pereira Jr. & Costa (1996), Knoff et al. (1997), Vicente & Pinto (1999), Alves et al. (2000), Luque & Alves (2001), Pereira Jr. et al. (2002). Cucullanus riograndensis Fortes, Hoffmann & SarmentoadultFWintestine Host: Pimelodus maculatus. Locality: Guaba LakeRS. References: Fortes et al. (1993a), Fortes & Hoffmann (1995), Vicente & Pinto (1999). Renark: Considered to be a species inquirenda by Moravec (1998). Cucullanus rougetae Vicente & SantosadultFWintestine Host: Plagioscion auratus. Locality: MacaRJ. References: Vicente & Santos (1974). Cucullanus schubarti TravassosadultFWintestine

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Host: Zungaro zungaro. Localities: Amazonas River Basin; PirassunungaSP. References: Travassos (1947, 1948), Yamaguti (1961), Travassos & Kohn (1965), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006). Cucullanus sp.adult Host: Ariidae gen. sp. (MARintestine), Astyanax bimaculatus (FWintestine), Brachyplatystoma sp. (FWintestine), Brycon hilarii (FWintestine), Centropomus undecimalis (BWpyloric caeca), Cynoscion guatucupa (MARintestine), Geophagus brasiliensis (FWintestine), Leporellus vittatus (FWintestine), Leporinus copelandii (FWintestine), Macrodon ancylodon (MARintestine), Micropogonias furnieri (MARintestine), Mugil curema (BWintestine), Pimelodus blochii (FW intestine), P. maculatus (FWintestine), Pimelodus sp. (FWintestine), Potamotrygon falkneri (FW spiral valve), Prionotus punctatus (MARintestine), Pseudoplatystoma fasciatum (FWintestine), Pseudoplatystoma sp. (FWintestine), Pterodoras granulosus (FWintestine), Rhamdia quelen (FW unspecified), Serrasalmus maculatus (FWunspecified), S. marginatus (FWintestine), S. spilopleura (FWintestine), Sorubim lima (FWintestine), Trachelyopterus galeatus (FWintestine), T. striatulus (FWunspecified), Urophycis brasiliensis (MARintestine), Zungaro zungaro (FWintestine). Localities: off IlhusBA; off FortalezaCE; off So LuisMA; So Francisco River, Aquidauana RiverMG; Bodoquena, Miranda River, SalobraMS; MT; BraganaPA; Paranagu Bay, Upper Paran RiverPR; Guand River, off Rio de JaneiroRJ; off Cassino BeachRS; off FlorianpolisSC; Canania, PirassunungaSP. References: Travassos et al. (1939), Travassos & Freitas (1942), Kloss (1966), Bhrnheim (1976), Vicente & Fernandes (1978), Conroy & Conroy (1984), Vicente et al. (1985), Kohn & Fernandes (1987), Pavanelli et al. (1997, 2004), Vicente & Pinto (1999), Brasil-Sato (2003), Sabas & Luque (2003), Santos et al. (2003), Alves et al. (2004), Luque & Poulin (2004), Bicudo et al. (2005a), Timi et al. (2005), Lacerda et al. (2008), Fujimoto et al. (2009), Takemoto et al. (2009), Campos et al. (2009b), Azevedo et al. (2010), Carvalho et al. (2010), Mesquita et al. (2011). Cucullanus sp. A of Bhrnheim (1976)adultFWintestine Hosts: Leporellus vittatus, Schizodon nasutus. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985). Cucullanus sp. B of Bhrnheim(1976)Unspecified stage of developmentFWintestine Host: Pimelodus maculatus. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985) Cucullanus sp. 1adult and larvaFWintestine Host: Zungaro zungaro. Locality: Paran River, GuairaPR. Reference: Moravec et al. (1993a). Remark: This species is C. pinna according to Moravec et al. (1997). Cucullanus sp. 2adultFWintestine Host: Pterodoras granulosus. Locality: Paran River, GuairaPR. Reference: Moravec et al. (1993a). Cucullanus sp. 3adultFWintestine Host: Serrasalmus marginatus. Locality: Paran River, GuairaPR. Reference: Moravec et al. (1993a). Cucullanus (Truttaedacnitis) rhamphichthydis Moravec, Kohn & FernandesadultFWintestine

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Host: Rhamphichthys rostratus. Locality: Itaip ReservoirPR. References: Moravec et al. (1997), Moravec (1998), Vicente & Pinto (1999), Moreira et al. (2000), Kohn et al. (2003), Thatcher (2006), Eiras et al. (2010). Dichelyne (Cucullanellus) sciaenidicola Timi, Lanfranchi, Tavares & LuqueMARintestine Hosts: Micropogonias furnieri, Paralonchurus brasiliensis, Umbrina canosai. Localities: IlhusBA; FortalezaCE; Rio de JaneiroRJ; Patos Lagoon and Cassino BeachRS; FlorianpolisSC. References: Pinto et al. (1970), Vicente et al. (1985, 1989), Vicente & Pinto (1999), Pereira Jr. & Costa (1996), Pereira Jr. et al. (2002), Alves & Luque (1999, 2000, 2001a,b), Timi et al. (2009), Luque et al. (2010). Remark: Previously reported as Dichelyne (Cucullanellus) elongatus (Tornquist, 1931), D. (C.) amaruincai (Freitas, Vicente & Ibaez, 1969), D. (C.) rodriguesi (Pinto, Fbio & Noronha, 1970) and D. (Dichelyne) micropogonii Pereira Jr. & Costa, 1996. Dichelyne (Cucullanellus) travassosi (Guimares & Cristofaro)adultMARstomach, intestine Syn.: Cucullanus travassosi Guimares & Cristofaro Hosts: Balistes vetula, Halichoeres radiatus. Locality: SalvadorBA. References: Guimares & Cristofaro (1974), Vicente & Santos (1974), Vicente et al. (1985), Muniz-Pereira et al. (2009). Dichelyne (Cucullanellus) sp. Hosts and site of infection: Paralonchurus brasiliensis (MARintestine), Trachelyopterus galeatus (FW adult). Localities: Upper Paran RiverPR; Marambaia IslandRJ. References: Pinto et al. (1992), Pavanelli et al. (1997, 2004), Vicente & Pinto (1999), Takemoto et al. (2009). Dichelyne (Dichelyne) leporini PetteradultFWintestine Hosts: Leporinus friderici, Leporinus lacustris, Schizodon fasciatus. Locality: Upper Paran RiverPR. References: Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999), Guidelli et al. (2006), Takemoto (2009), Eiras et al. (2010). Dichelyne (Dichelyne) pimelodi Moravec, Kohn & FernandesadultFWintestine Host: Pimelodus maculatus, Pimelodus sp. Locality: Guara, Itaip ReservoirPR. References: Moravec et al. (1997), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Eiras et al. (2010). Dichelyne sp.adult Hosts and site of infection: Balistes capriscus (MARintestine), B. vetula (MARintestine), Pseudopercis numida (MARintestine), P. semifasciata (MARintestine). Locality: Rio de JaneiroRJ. References: Alves et al. (2005), Luque et al. (2008), Muniz-Pereira et al. (2009). Neocucullanus neocucullanus Travassos, Artigas & PereiraadultFW Syn: Cucullanus interrogativus Travassos, Artigas & Pereira; C. neocucullanus (Travassos, Artigas & Pereira); Neocucullanus multipapilatus Petter Hosts and site of infection: Brycon sp., Brycon hilarii (intestine, pyloric caeca), Characidae gen. sp. (intestine), Salminus brasiliensis (intestine), S. hilarii (liver). Localities: Das Velhas River BasinMG; Juba RiverMT; PirassunungaSP.

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References: Travassos et al. (1928), Yamaguti (1961), Travassos & Kohn (1965), Bhrnheim (1976), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006), Moravec (1998), Rodrigues et al. (2002), Brasil-Sato (2003), Saraiva et al. (2006b,c), Eiras et al. (2010). Cucullanidae gen. sp.adultFWintestine Host: Galeocharax humeralis. Locality: PirassunungaSP. Reference: Kohn & Fernandes (1987).

Family Quimperiidae Gendre


Neoparaseuratum travassosi Moravec, Kohn & FernandesadultFWintestine Hosts: Pterodoras granulosus, Trachydoras paraguayensis. Locality: Itaip Reservoir, Paran RiverPR. References: Moravec et al. (1992c, 1993a, 1997), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Eiras et al. (2010). Paraseuratum albidum KlossadultFWintestine Hosts: Astyanax bimaculatus, A. fasciatus, Erythrinus erythrinus, Tetragonopterus argenteus. Locality: PirassunungaSP. References: Kloss (1966), Bhrnheim (1976), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Paraseuratum soaresi FbioadultFW Host: Hoplias malabaricus. Locality: CamposRJ. References: Fbio (1982), Vicente et al. (1985), Moravec (1998), Eiras et al. (2010). Paraseuratum sp.Unspecified stage of developmentFW Syn.: Paraseuratum luroris of Bhrnheim (1976) Host: Hoplias malabaricus. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985).

Order Oxyurida Chabaud Superfamily Oxyuroidea Cobbold Family Pharyngodonidae Travassos


Brasilnema pimelodellae Moravec, Kohn & FernandesadultFWintestine Host: Pimelodella lateristriga. Locality: Guara, Paran RiverPR. References: Moravec et al. (1992a,b), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Eiras et al. (2010). Cosmoxynema vianai TravassosadultFW Hosts and site of infection: Cyphocharax gilbert (intestine), Pseudocurimata sp. (intestine), Steindachnerina brevipinna, S. elegans (intestine), S. insculpta (unspecified). Localities: Barra Seca River and Juparan LagoonES; Guara, Low Paranapanema river, Paran River, Upper Paran RiverPR; PirassunungaSP.

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References: Travassos (1948), Yamaguti (1961), Bhrnheim (1976), Vicente et al. (1985), Thatcher (1991, 2006), Moravec et al. (1992a), , Moravec (1998), Vicente & Pinto (1999), Takemoto et al. (2009), Ceschini et al. (2010a,b), Eiras et al. (2010). Cosmoxynemoides aguirrei TravassosadultFWintestine Hosts: Astyanax bimaculatus, Curimatella lepidura, Cyphocharax gilbert, Geophagus brasiliensis, G. steindachneri, Pseudocurimata sp., Steindachnerina elegans. Localities: Barra Seca River and Juparan LagoonES; Guara, Paran RiverPR; Guandu RiverRJ; PirassunungaSP. References: Travassos (1948), Yamaguti (1961), Bhrnheim (1976), Vicente & Pinto (1981, 1999), Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006), Moravec et al. (1992a, 1994a), Moravec (1998), Abdallah et al. (2005), Azevedo et al. (2010), Eiras et al. (2010). Cosmoxynemoides sp.adultFWintestine Host: Trachydoras paraguayensis. Locality: Upper Paran RiverPR. References: Pavanelli et al. (1997, 2004), Takemoto et al. (2009). Ichthyouris brasiliensis Moravec, Kohn & FernandesadultFWintestine Host: Megalancistrus aculeatus. Locality: Guara, Paran RiverPR. References: Moravec et al. (1992a,b), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Eiras et al. (2010). Ichthyouris bursata Moravec & ProuzaadultFWintestine Host: Symphysodon aequifasciatus. Locality: Brazil. References: Moravec & Laoprasert (2008). Ichthyouris laterifilamenta Moravec, Kohn & FernandesadultFWintestine Host: Trachydoras paraguayensis. Locality: Guara, Paran RiverPR. References: Moravec et al. (1992a), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Thatcher (2006), Eiras et al. (2010). Ichthyouris ovofilamentosa Moravec & ThatcheradultFWintestine Host: Cichlasoma sp. Locality: Negro RiverAM. References: Moravec & Thatcher (2001), Thatcher (2006), Eiras et al. (2010). Ichthyouris voltagrandensis Martins, Yoshitoshi & UmekitaadultFWintestine Host: Myleus tiete. Locality: Volta Grande ReservoirMG. References: Martins et al. (2001, 2003), Eiras et al. (2010). Ichthyouris sp. of Moravec & Thatcher (2001)adultFWintestine Host: Cichlasoma sp. Locality: Negro RiverAM. References: Moravec & Thatcher (2001), Eiras et al. (2010). Parasynodontisia petterae Moravec, Kohn & FernandesadultFWintestine Host: Rhinelepis aspera. Locality: Guara, Paran RiverPR. References: Moravec et al. (1992a,b), Moravec (1998), Vicente & Pinto (1999), Thatcher (2006), Eiras et al. (2010). Spinoxyuris annulata Moravec & ThatcheradultFWUnspecified site of infection Host: Mylossoma aureum.

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Locality: Guandu RiverRJ. Reference: Azevedo et al. (2010). Spinoxyuris oxydoras PetteradultFWUnspecified site of infection Host: Metynnis lippincottianus, Oxydoras kneri. Locality: Upper Paran RiverPR. References: Moreira et al. (2009, 2010), Takemoto et al. (2009), Eiras et al. (2010). Spinoxyuris sp.adultFWintestine Host: Steindachnerina brevipinna. Locality: Low Paranapanema riverPR. Reference: Ceschini et al. (2010a,b). Travnema araujoi Fernandes, Campos & ArtigasadultFWintestine Hosts: Cyphocharax gilbert, Salminus sp., Steindachnerina elegans. Localities: Amazonas River; Guandu River RJ; Paran RiverPR; Ilha Solteira DamSP. References: Travassos (1920, 1945), Travassos et al. (1928, 1939), Travassos & Kohn (1965), Santos et al. (1979), Fernandes et al. (1983), Kohn et al. (1985), Vicente et al. (1985), Kohn & Fernandes (1987), Rgo & Vicente (1988), Thatcher (1991, 2006), Moravec et al. (1992a), Martins & Urbinati (1993), Moravec & Thatcher (1997b), Moravec (1998), Vicente & Pinto (1999), Abdallah et al. (2005), Azevedo et al. (2010), Eiras et al. (2010). Travnema travnema PereiraadultFW intestine Hosts and site of infection: Astyanax bimaculatus, Cyphocharax gilbert, C. plumbeus, Steindachnerina elegans, S. insculpta, S. brevipinna. Localities: FortalezaCE; Guara, Itaip Reservoir, Lower Paranapanema River, Paran and Upper Paran RiverPR; PirassunungaSP. References: Pereira (1938), Yamaguti (1961), Bhrnheim (1976), Vicente & Pinto (1981, 1999), Kohn et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006), Moravec et al. (1992a, 1994a), Moravec (1998), Takemoto et al. (2009), Ceschini et al. (2010a,b), Eiras et al. (2010). Travnema sp.FWpyloric caeca and intestine Syn.: Travnema incoptus of Bhrnheim (1976, unpublished) Hosts: Cyphocharax gilbert, Steindachnerina elegans, Pseudocurimata sp. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985). Remark: According to Bhrnheim (1976), this parasite differs from T. travnema.

Order Spirurida Chitwood Suborder Camallanina Chitwood Superfamily Camallanoidea Railliet & Henry Family Camallanidae Railliet & Henry Camallanidae gen. sp.adultFW Hosts: Astyanax bimaculatus (intestine), Brycon hilarii (unspecified), Cyphocharax gilbert (intestine), Moenkhausia doceana (intestine), Pimelodus fur (immatureunspecified), Pygocentrus nattereri (unspecified). Localities: SooteramaES; SalobraMT; Ilha SecaSP. References: Travassos (1940a), Travassos & Feitas (1942), Travassos et al. (1964), Vicente et al. (1985).

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Subfamily Camallaninae Railliet & Henry


Camallanus (Camallanus) acaudatus Ferraz & ThatcheradultFWintestine Host: Osteoglossum bicirrhosum. Locality: Amazonas River BasinAM. References: Ferraz & Thatcher (1990), Thatcher (1991, 2006), Moravec (1998), Vicente & Pinto (1999), Eiras et al. (2010). Camallanus cotti FujitaadultFW Hosts and site of infection: Apistogramma cacatuoides (unspecified), Betta splendens (rectal lumen), Poecilia latipinna (unspecified), P. reticulata (intestine), Pterophylum scalare (unspecified), Xiphophorus helleri (unspecified). Locality: Itagua, NiteriRJ. References: Alves et al. (2000), Menezes et al. (2006), Eiras et al. (2010). Camallanus maculatus Martins, Garcia, Piazza & Ghiraldelliadult and larvaFWintestine Host: Xiphophorus maculatus, Poecilia sphenops. Locality: AraraquaraSP. References: Martins et al. (2007), Eiras et al. (2010). Camallanus (Camallanus) tridentatus (Drasche)adultFW Syn.: Cucullanus tridentatus Drasche Hosts and site of infection: Arapaima gigas (stomach, pyloric caeca, intestine), Osteoglossum bicirrhosum (intestine). Localities: Amazon fish farmAM; Amazonas River Basin, Mexiana IslandPA. References: Drasche (1884a), Baylis (1927), Travassos et al. (1928), Yamaguti (1961), Vicente & Pinto (1981, 1999), Vicente et al.(1985), Ferraz & Thatcher (1990), Thatcher (1991, 2006), Moravec (1998), Araujo et al. (2009b), Santos & Moravec (2009a), Eiras et al. (2010). Camallanus sp.adultFWintestine Hosts: Astronotus ocellatus, Charax gibbosus, Poecilia sphenops. Locality: Bodoquena, SalobraMS. References: Travassos & Freitas (1942), Vicente et al. (1985), Eiras et al. (2010). Oncophora melanocephala (Rudolphi)adultMARintestine Syn.: Trichocephalus gibbosus Rudolphi; Cucullanus melanocephalus Rudolphi Hosts: Auxis thazard thazard, Priacanthus arenatus, Sarda sarda, Thunnus albacores, T. atlanticus, T. thynnus. Locality: Cabo Frio, Rio de JaneiroRJ. References: Rudolphi (1819), Diesing, 1851, Pinto et al. (1988), Moravec et al. (1999), Vicente & Pinto (1999), Tavares et al. (2001), Alves & Luque (2006).

Subfamily Paracamallaninae Stromberg & Crites


Paracamallanus amazonensis Ferraz & ThatcheradultFWintestine Hosts: Hypophthalmus edentatus, Pterodoras granulosus. Localities: Marchantaria Island, Solimes RiverAM; Itaip Reservoir, Paran RiverPR References: Ferraz & Thatcher (1992), Moravec et al. (1993b), Moravec (1998), Vicente & Pinto (1999), Eiras et al. (2010). Paracamallanus sp.Unspecified stage of developmentFWintestine Host: Hypophthalmus edentates. Locality: Amazonas River Basin. References: Thatcher (1991), Vicente & Pinto (1999).

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Subfamily Procamallaninae Yeh


Procamallanus (Denticamallanus) dentatus Moravec & ThatcheradultFWintestine Host: Bryconops alburnoides. Locality: Amazonas River BasinAM. References: Moravec & Thatcher (1997a), Moravec (1998), Thatcher (2006), Eiras et al. (2010). Procamallanus (Procamallanus) annipetterae Kohn & FernandesadultFWintestine Syn.: Procamallanus petterae Kohn & Fernandes Hosts: Hypostomus albopunctatus, H. cochliodon. Locality: Iguau River BasinPR. References: Kohn et al. (1988), Kohn & Fernandes (1988a,b), Moravec & Sey (1988), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Vicente & Pinto (1999), Eiras et al. (2010). Procamallanus (Procamallanus) peraccuratus Pinto, Fbio, Noronha & RolasadultFWintestine Hosts: Australoheros facetus, Cichla ocellaris, Crenicichla lepidota, Geophagus brasiliensis, Gymnotus carapo, Hoplias malabaricus, Pimelodus ortmanni, Potamotrygon motoro, Schizodon nasutus, Trachelyopterus striatulus. Localities: Cupido, Engano, Itaunas Rivers and Juparan Lake, Santa TerezaES; Guand RiverRJ; Iguau River BasinPR; Paran; PirassunungaSP; Upper Parana River. References: Pinto et al. (1976), Vicente & Pinto (1981, 1999), Kohn et al. (1985), Vicente et al. (1985), Kohn et al. (1988, 1989), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec et al. (1993b), Pavanelli et al. (2004), Takemoto et al. (2009), Azevedo et al. (2010), Carvalho et al. (2010), Eiras et al. (2010), Mesquita et al. (2011). Procamallanus (Spirocamallanus) amarali Vaz & PereiraadultFWintestine Hosts: Leporinus copelandii, L. elongatus, L. friderici, L. obtusidens, Leporinus sp. Localities: Upper Paran RiverPR; Pirassununga, Tiet River, SaltoSP. References: Vaz & Pereira (1934), Yamaguti (1961), Pinto et al. (1974, 1975), Bhrnheim (1976), Vicente et al. (1985), Kohn & Fernandes (1987), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Guidelli et al. (2006), Takemoto et al. (2009), Eiras et al. (2010). Procamallanus (Spirocamallanus) barroslimai PereiraUnspecified stage of developmentFWintestine Hosts: Clupeidae gen. sp., Sardinha, Triportheus sp. Localities: Amazonas River BasinAM; CruzetaRN. References: Pereira (1935), Yamaguti (1961), Pinto & Noronha (1976), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Procamallanus (Spirocamallanus) belenensis Giese, Santos & LanfrediadultFWintestine Host: Ageneiosus ucayalensis. Locality: Guajar Bay BelmPA. References: Giese et al. (2009). Procamallanus (Spirocamallanus) caballeroi BashirullahadultFWintestine Host: Astyanax altiparanae. Locality: Upper Paran RiverPR. References: Pavanelli et al. (2004), Takemoto et al. (2009), Eiras et al. (2010). Procamallanus (Spirocamallanus) cruzi Guimares, Cristofaro & RodriguesadultMAR Hosts and site of infection: Mullus argentinae (intestine), Polydactylus virginicus (intestine and stomach). Localities: SalvadorBA; Rio de JaneiroRJ. References: Guimares et al. (1976), Vicente et al. (1985), Rodrigues et al. (1991), Luque et al. (2002). Procamallanus (Spirocamallanus) freitasi (Moreira, Oliveira & Costa)adultFWintestine Hosts: Bergiaria westermanni, Pimelodus maculatus, Pimelodus sp.. Locality: So Francisco RiverMG.

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References: Moreira et al. (1991), Rodrigues et al. (1991), Moravec (1998), Vicente & Pinto (1999), BrasilSato (2003), Eiras et al. (2010). Procamallanus (Spirocamallanus) halitrophus (Fusco & Overstreet)adultMARintestine Hosts: Citharichthys macrops, Syacium papillosum. Locality: Rio de JaneiroRJ. References: Crdenas & Lanfredi (2005), Crdenas et al. (2005). Procamallanus (Spirocamallanus) hilarii Vaz & Pereiraadult and larvaFWintestine Syn.: Spirocamallanus hilarii Vaz & Pereira; Procamallanus (S.) cearensis Pereira, Dias & Azevedo; P. cearensis Pereira, Dias & Azevedo; P. (Isospiculus) cearensis (Pereira, Dias & Azevedo); S. incarocai Freitas & Ibes Hosts: Acestrorhynchus microlepis (unspecified), Astyanax bimaculatus (adult), A. fasciatus (adult), A. parahybae (adult), Hoplias lacerdae (adult), H. malabaricus (adult), Oligosarcus macrolepis (adult), Rhamdia quelen (adult), Salminus hilarii (adult), Steindachnerina elegans (adult and larva), Trichomycterus piurae (unspecified). Localities: Fortaleza, Porangaba, Soure, TauapCE; Atuba, Santo Amaro, Curitiba, Igua River, ParanPR; Guandu RiverRJ; Passo Fundo Reservoir, Uruguay RiverRS; PirassunungaSP. References: Vaz & Pereira (1934), Pereira et al. (1936), Yamaguti (1961), Pinto et al. (1974), Bhrnheim (1976), Pinto & Noronha (1976), Vicente & Pinto (1981, 1999), Vicente et al. (1985), Kohn & Fernandes (1987), Kohn et al. (1988, 1989), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Abdallah et al. (2004), Azevedo et al. (2010), Eiras et al. (2010). Procamallanus (Spirocamallanus) iheringi Travassos, Artigas & PereiraadultFW Hosts and site of infection: Anostominae gen. sp. (intestine, pyloric caeca), Astyanax fasciatus (intestine), Characidae gen. sp. (unspecified), Hoplias malabaricus (intestine), Hoplias sp. (intestine, pyloric caeca), Leporinus copelandii (intestine), L. elongatus (intestine), L. fasciatus (intestine), L. friderici (intestine), L. obtusidens (intestine), L. octofasciatus (intestine), Leporinus sp. (intestine), Salminus hilarii (intestine), Schizodon borellii (intestine), S. fasciatus (intestine), S. nasutus (intestine), Tetragonopterinae gen. sp. (intestine, pyloric caeca), Tetragonopterus sp. (intestine), Zungaro zungaro (intestine). Localities: LinharesES; Nova Ponte Dam, PerdizesMG; Paran, Upper Paran RiverPR; Paran River, Pirassununga, Porto CabralSP. References: Travassos et al. (1928), Travassos & Freitas (1948), Yamaguti (1961), Travassos & Kohn (1965), Pinto et al. (1974, 1975), Pinto & Noronha (1976), Vicente & Pinto (1981, 1999), Kohn et al. (1985), Vicente et al. (1985), Kohn & Fernandes (1987), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec et al. (1993b), Pavanelli et al. (1997, 2004), Feltran et al. (2004), Guidelli et al. (2006), Takemoto et al. (2009), Eiras et al. (2010).

Procamallanus (Spirocamallanus) inopinatus (Travassos, Artigas & Pereira)adultFW Syn.: Spirocamallanus inopinatus (Travassos, Artigas & Pereira); Procamallanus fariasi Pereira; P. wrighti Pereira; S. wrighti Olsen; P. (Isospiculus) wrighti Ali; P. probus Pinto & Fernandes; P. (S.) wrighti Pereira Hosts and site of infection: Acestrorhynchus falcatus (unspecified), Ageneiosus inermis (unspecified), Astronotus ocellatus (intestine), Astyanax altiparanae (intestine), A.bimaculatus (intestine), A. fasciatus (intestine), A. lacustris (unspecified), A. schubarti (intestine), Astyanax sp. (intestine), Brycon amazonicus (intestine), B. cephalus (intestine), B. falcatus (intestine), B. hilarii (intestine, pyloric caeca), B. orthotaenia (intestine), Brycon sp. (intestine), Calophysus macropterus (intestine), Charax gibbosus, Cichla keberi (intestine), Crenicichla haroldoi (intestine), Galeocharax humeralis (intestine), Hoplias malabaricus (intestine, pyloric caecum), Leporellus vittatus (unspecified), Leporinus agassizii (intestine), L. copelandii (intestine), L. elongatus (intestine), L. fasciatus (intestine), L. friderici (intestine), L. lacustris (intestine), L. obtusidens (intestine), L. octofasciatus (intestine), L. piau (intestine), L. reinhardti (intestine), L. striatus (intestine), L. taeniatus (intestine), Leporinus sp. (intestine), Metynnis lippincottianus (unspecified), Myleus

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asterias (intestine), M. schomburgkii (unspecified), Pimelodus blochii (intestine), Potamotrygon motoro (intestine), Prochilodus lineatus (intestine), Pterodoras granulosus (intestine), Pygocentrus nattereri (intestine), P. piraya (intestine), Pygocentrus sp. (intestine), Salminus brasiliensis (intestine), S. hilarii (intestine), Schizodon borellii (intestine), S. knerii (intestine), S. nasutus (intestine), Serrasalmus brandtii (intestine), S. gouldingi (unspecified), S. maculatus (unspecified), S. manueli (unspecified), S. marginatus (intestine), S. spilopleura (intestine), Trachydoras paraguayensis (intestine), Triportheus paranensis (unspecified). Localities: Manaus, Negro and Solimes RiversAM; Juruparan LagoonES; Alfenas, Harmonia, Perdizes, So Francisco River, Trs Marias ReservoirMG; Juba River, Pantanal, SalobraMT; Cachimbo, Curu RiverPA; Areia, PatosPB; PaulistanaPI; Paran, Upper Paran RiverPR; Campos, Rio de Janeiro, Guandu RiverRJ; Cruzeta, CaicRN; Machado River, Porto VelhoRO; Agua, Campinas, Paran River, Pirassununga, Porto CabralSP. References: Travassos et al. (1928), Pereira (1935), Yamaguti (1961), Travassos & Kohn (1965), Kloss (1966), Pinto & Fernandes (1972), Pinto & Noronha (1972, 1976), Pinto et al. (1974, 1975, 1976), Santos et al. (1979), Vicente & Pinto (1981, 1999), Fbio (1982), Kohn et al. (1985), Vicente et al. (1985), Kohn & Fernandes (1987), Petter & Thatcher (1988), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec et al. (1993b), Moreira et al. (1994, 2009, 2010), Pavanelli et al. (1997, 2004), Andrade et al. (2001), Brasil-Sato (2003), Feltran et al. (2004), Adriano et al. (2005), Guidelli et al. (2006), Saraiva et al. (2006a,b), Andrade & Malta (2006), Azevedo et al. (2007c, 2010), Araujo et al. (2009a), Takemoto et al. (2009), Eiras et al. (2010), Takemoto & Lizama (2010), Vicentin et al. (2011). Procamallanus (Spirocamallanus) macaensis Vicente & Santosadult Syn.: P. macaensis Vicente & Santos Hosts and site of infection: Cara suja (MARintestine), Chaetodipterus faber (MARintestine), Dactylopterus volitans (MARintestine), Menticirrhus americanus (MARintestine), Micropogonias undulatus (MARintestine), Paralonchurus brasiliensis (MARintestine), Plagioscion auratus (FW intestine), Thyrsitops lepidopoides (MARintestine), Urophycis brasiliensis (MAR), Urophycis sp. (MARintestine). Locality: MacaRJ. References: Vicente & Santos (1972, 1973), Pinto et al. (1974), Pinto & Noronha (1976), Vicente et al. (1985), Rodrigues et al. (1991), Alves et al. (2004), Cordeiro & Luque (2005b). Procamallanus (Spirocamallanus) paraensis Pinto & NoronhaadultFWintestine Hosts: Erythrinus erythrinus, Jeju. Localities: Amazonas River BasinAM; CachimboPA. References: Pinto & Noronha (1976), Vicente & Pinto (1981), Vicente et al. (1985), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Remark: Thatcher (2006) suggested that the Jeju may be an erythrinid. Procamallanus (Spirocamallanus) pereirai AnnereauxadultMAR Hosts: Atlantoraja castelnaui (spiral valve), Menticirrhus americanus (intestine), Micropogonias furnieri (intestine), Nebris microps (intestine), Paralonchurus brasiliensis (intestine), Porichthys porosissimus (intestine), Stellifer brasiliensis (intestine), Symphurus tessellatus (intestine). Localities: Araruama, Rio de JaneiroRJ; CoastRS. References: Pinto et al. (1984), Rodrigues et al. (1991), Santos et al. (1999), Knoff et al. (2001), Ribeiro et al. (2002), Luque et al. (2003). Procamallanus (Spirocamallanus) pexatus Pinto, Fbio, Noronha & RolasadultFWintestine Host: Pygidium brasiliensis. Locality: Santa TerezaES.

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References: Pinto et al. (1976), Vicente & Pinto (1981), Vicente et al. (1985), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Remark: The host genus has been considered as a genus inquirendum (Pinna & Wosiacki, 2003; Ferraris, 2007; Eschmeyer, 2010). Procamallanus (Spirocamallanus) pimelodus Pinto, Fbio, Noronha & RolasadultFWintestine Syn.: P. intermedius Pinto, Fbio, Noronha &. Rolas Hosts: Iheringichthys labrosus, Pimelodella lateristriga, P. blochii, P. maculatus, P. ortmanni, Pimelodus sp., Synodontis clarias. Localities: Porto EsperanaMT; PantanalMT; Paran, Upper Paran RiverPR; Paran River, PirassunungaSP. References: Pinto et al. (1974, 1976), Vicente et al. (1985), Kohn & Fernandes (1987), Kohn et al. (1988), Rgo & Vicente (1988b), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec et al. (1993b), Moravec (1998), Vicente & Pinto (1999), Moreira et al. (2005), Takemoto et al. (2009), Eiras et al. (2010). Procamallanus (Spirocamallanus) pintoi (Kohn & Fernandes)adultFWintestine Syn. S. pintoi Kohn & Fernandes Hosts: Corydoras aeneus, C. paleatus. Locality: Paran, Upper Paran RiverPR. References: Kohn et al. (1988), Kohn & Fernandes (1988a), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Vicente & Pinto (1999), Ito et al. (2005), Eiras et al. (2010). Procamallanus (Spirocamallanus) rarus Travassos, Artigas & PereiraadultFWintestine Hosts: Ageneiosus ucayalensis, Bagre, Pimelodella lateristriga, Pimelodus albicans, P. maculatus, Rhinodoras dorbignyi, Serrasalmus sp., Synodontis clarias. Localities: Belm, Guajar BayPA; PirassunungaSP. References: Travassos et al. (1928), Yamagutti (1961), Travassos & Kohn (1965), Pinto et al. (1974), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Rodrigues et al. (1991), Thatcher (1991, 2006), Giese et al. (2009), Eiras et al. (2010). Procamallanus (Spirocamallanus) saofranciscensis (Moreira, Oliveira & Costa)adultFWintestine Syn.: S. saofranciscensis Moreira, Oliveira & Costa Hosts: Acestrorhynchus lacustris, Astyanax fasciatus, Tetragonopterus chalceus, Triportheus guentheri. Locality: So Francisco RiverMG. References: Moreira et al. (1994), Moravec (1998), Vicente & Pinto (1999), Brasil-Sato (2003), Eiras et al. (2010). Procamallanus (Spirocamallanus) solani Pinto, Fbio, Noronha & RolasadultFWintestine Host: Siluriformes fam. gen. sp. Locality: Amazonas River, MaicurPA. References: Pinto et al. (1975), Vicente & Pinto (1981), Vicente et al. (1985), Rodrigues et al. (1991), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Procamallanus (Spirocamallanus) sp.adult and larvaFW Hosts: Astyanax bimaculatus (intestine), A. fasciatus (intestine), Auchenipterus osteomystax (unspecified), Bivibranchia fowleri (unspecified), Bivibranchia velox (unspecified), Car-cachimbo (intestine), Chimbor (intestine), Colossoma macropomum (adultintestine), Conorynchus conirostris (larva intestine), Ferreirinha (intestine), Hemiodus sp. (intestine), Leporinus friderici (unspecified), L. obtusidens (adultintestine), Pachyurus squamipennis (adultintestine), Peixe-cachorro (intestine), Paracheirodon axelrodi (intestine), Pimelodus maculatus (adultunspecified), Prochilodus lineatus (adultintestine), Pseudoplatystoma corruscans (adultintestine), Rhinodoras dorbignyi (intestine), Salminus hilarii (adultintestine), Triportheus nematurus (intestine). Localities: Amazonas River, Maicur, Xingu RiverPA; So Francisco RiverMG; Igua River, Upper Paran RiverPR; PirassunungaSP.

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References: Pinto et al. (1976), Kohn & Fernandes (1987), Kohn et al. (1988), Machado et al. (1996), Pavanelli et al. (1997, 2004), Fischer et al. (2003), Brasil-Sato & Santos (2005), Takemoto et al. (2009), Tavernari et al. (2009), Costa & Camargo (2009). Procamallanus (Spirocamallanus) sp.adultFWintestine Syn.: "S. furcospiculum" of Bhrnheim (1976) Hosts: Pimelodus maculatus, Pimelodus sp. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985). Procamallanus (Spirocamallanus) sp.adultFWintestine Syn.: "S. laticonchus" of Bhrnheim (1976) Hosts: Acestrorhynchus lacustris, Astyanax bimaculatus, A. fasciatus, A. schubarti, Hoplias malabaricus, Salminus brasiliensis, S. hilarii. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985). Procamallanus (Spirocamallanus) sp. A of Bhrnheim (1976)FWintestine Hosts: Leporinus copelandii, L. lacustris, L. striatus, Pimelodus maculatus Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985). Remark: Bhrnheim (1976) claimed to have found only larva specimens. Procamallanus (Spirocamallanus) sp. B of Bhrnheim (1976)adultFWintestine Hosts: Leporinus lacustris, L. octofasciatus, Leporinus sp. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985). Procamallanus sp.adult and larva Hosts and site of infection: Acestrorhynchus lacustris (FWadultintestine), Astyanax bimaculatus (FWadultintestine), A. fasciatus (FWadultintestine), Astyanax sp. (FW unspecified unspecified), Australoheros facetus (FW unspecifiedunspecified), Brevoortia aurea (MARadult intestine), Brycon insignis (FWadultintestine, stomach), Calophysus macropterus (FWunspecified intestine), Car-cachimbo (FWunspecifiedintestine), Colossoma macropomum (FWadult intestine), Genypterus brasiliensis (MARadultintestine), Hemiodus sp. (FWunspecifiedintestine), Hoplias malabaricus (FWunspecifiedunspecified), Leporinus macrocephalus (FWadultintestine), L. octofasciatus (FWunspecifiedintestine), L. striatus (FWunspecifiedintestine), Leporinus sp. (FWunspecifiedintestine), Menticirrhus americanus (MARadultintestine), Paracheirodon axelrodi (FWunspecifiedintestine), Peixe-cachorro (FWunspecifiedintestine), Percophis brasiliensis (MARadultintestine), Pimelodus blochii (FWunspecifiedunspecified), P. maculatus (FWunspecifiedunspecified), Prionotus punctatus (MARadultintestine), Prochilodus lineatus (FWlarvaintestine), Pseudopercis numida (MARadultintestine), P. semifasciata (MARadult intestine), Pygocentrus nattereri (FWbody cavity), Salminus hilarii (FWlarvaintestine), Sarda sarda (MARadultintestine), Schizodon altoparanae (FWadultintestine), Serrasalmus marginatus (FW unspecifiedunspecified), S. spilopleura (FWunspecifiedunspecified), Siluriformes fam. gen. sp. (FWunspecifiedintestine), Synodontis clarias (FWunspecifiedunspecified), Trachydoras paraguayensis (FWadultintestine), Trichiurus lepturus (MARadultintestine). Localities: ManausAM; off SoteramaES; Cuiab River, Porto EsperanaMS, MT; Amazonas River, MaicurPA; Upper Paran RiverPR; off Cabo Frio, Itaocara, Maca, Rio de Janeiro; Guandu River RJ; Porto Cabral, Paran River, PirasunungaSP. References: Travassos & Freitas (1940, 1948), Vicente & Santos (1973), Pinto & Noronha (1976), Kohn et al. (1985), Vicente et al. (1985), Pavanelli et al. (1997, 2004), Vicente & Pinto (1999), Silva et al. (2000b), Alves et al. (2002a,b), Carvalho et al. (2003), Fischer et al. (2003), Santos et al. (2003, 2007), Tavares et al.

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(2004b), Bicudo et al. (2005a), Alves & Luque (2006), Fernandes et al. (2006), Luque et al. (2008), Takemoto et al. (2009), Azevedo et al. (2010), Tavares-Dias et al. (2009, 2010), Eiras et al. (2010).

Superfamily Dracunculoidea Stiles


Filariidae gen. sp.adultFW Host: Hoplias malabaricus. Localities: SalobraMT; Ilha SecaSP. Reference: Travassos (1940a). Remark: Travassos (1940a) named these worms, collected during the third expedition to northwest of Brazil, as filariid nematodes; however, the actual nature of this record is not known.

Family Guyanemidae Petter Subfamily Guyaneminae Petter


Guyanema raphiodoni Moravec, Kohn & FernandesadultFWintestine Host: Rhaphiodon vulpinus. Locality: Foz do Igua, Itaip ReservoirPR. References: Moravec et al. (1993b,c), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Thatcher (2006), Eiras et al. (2010). Guyanema sp.adultFWintestine Host: Cyphocharax gilbert. Locality: Foz do Igua, Itaip ReservoirPR. References: Moravec et al. (1993b,c), Moravec (1998), Vicente & Pinto (1999), Eiras et al. (2010).

Subfamily Travassosneminae Moravec


Travassosnema travassosi paranaensis Moravec, Kohn & FernandesadultFWbody cavity Host: Acestrorhynchus lacustris. Locality: Guara, Paran RiverPR. References: Moravec et al. (1993b,c), Moravec (1998), Vicente & Pinto (1999), Kohn et al. (2003), Eiras et al. (2010). Travassosnema travassosi travassosi Costa, Moreira & OliveiraadultFWeyes Host: Acestrorhynchus lacustris. Localities: So Francisco River, Trs Marias ReservoirMG; Sertanpolis, Tibagi RiverPR. References: Costa et al. (1991), Moravec (1998), Vicente & Pinto (1999), Silva-Souza & Saraiva (2002), Brasil-Sato (2003), Thatcher (2006), Eiras et al. (2010).

Family Micropleuridae Baylis & Daubney Subfamily Micropleurinae Baylis & Daubney
Philonema sp. AadultFWbody cavity Host: Acestrorhynchus lacustris.

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Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985), Moravec (1998), Eiras et al. (2010). Philonema sp. BadultFWbody cavity Hosts: Cyphocharax gilbert, Pseudocurimata sp. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985), Moravec (1998), Eiras et al. (2010). Philonema sp. CadultFWbody cavity Host: Hoplias malabaricus. Locality: PirassunungaSP. References: Bhrnheim (1976), Vicente et al. (1985).

Family Philometridae Baylis & Daubney


Philometridae gen. sp.Unspecified stage of developmentFW Hosts and site of infection: Acestrorhynchus lacustris (unspecified), Pseudoplatystoma corruscans (intestine), Serrasalmus maculatus (unspecified), S. marginatus(unspecified), S. spilopleura (unspecified). Locality: Upper Paran RiverPR. References: Pavanelli et al. (1997, 2004), Takemoto et al. (2009).

Subfamily Alineminae Moravec


Alinema amazonicum (Travassos)adultFWbody cavity Syn.: Philometra amazonica Travassos; P. alli Rasheed; P. (Alinema) alli Rasheed; A. alii Rasheed Hosts: Calophysus macropterus, Luciopimelodus pati, Pinirampus pirinampu. Locality: Amazonas River, MaicuruPA. References: Travassos (1960), Rasheed (1963), Inglis & Ogden (1964), Santos et al. (1979), Vicente & Jardim (1980), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Moravec et al. (2006), Eiras et al. (2010).

Subfamily Philometrinae Baylis & Daubney


Nilonema senticosum (Baylis)adultFWbody cavity, swim bladder Syn.: Philometra senticosa Baylis Host: Arapaima gigas. Localities: Araguaia RiverMT; Amazonas RiverPA. References: Baylis (1927), Travassos et al. (1928), Travassos (1960),Yamaguti (1961), Santos et al. (1979), Vicente & Jardim (1980), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Santos & Gibson (2007), Santos & Moravec (2009c), Eiras et al. (2010). Philometra baylisi Vaz & PereiraadultFWabdominal cavity Host: Pimelodus maculatus, Synodontis clarias. Locality: Tiet River, SaltoSP. References: Vaz & Pereira (1934), Yamaguti (1961), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Philometra fariaslimai FortesadultBWbody cavity Hosts: Cathorops agassizii, Genidens barbus, G. genidens. Localities: Angra dos ReisRJ; Guaba LakeRS.
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References: Fortes (1981), Vicente et al. (1985), Fortes & Hoffmann (1995), Tavares & Luque (2008). Remark: This species was considered a species inquirenda by Moravec (1998). Philometra katsuwoni Petter & BaudinLaurencinadultMARgonads Host: Katsuwonus pelamis. Locality: Cabo FrioRJ. Reference: Crdenas et al. (2009). Philometra lateolabracis (Yamaguti)adultMARgonads Host: Haemulon plumierii. Locality: FortalezaCE. References: Crisp & Klein (1973), Vicente et al. (1985). Philometra sp.adult Hosts and site of infection: Caranx hippos (MARintestine), Lutjanus synagris (MARovary), Paralonchurus brasiliensis (MARgonads), Pimelodus maculatus (FW), Pomatomus saltatrix (MAR gonads). Localities: Rio de JaneiroRJ; Upper Paran RiverPR; off coastRio Grande do Norte RN References: Rgo et al. (1983), Vicente et al. (1985), Luque & Chaves (1999), Luque & Alves (2001), Ribeiro et al. (2002), Luque et al. (2003), Pavanelli et al. (2004), Takemoto et al. (2009), Cavalacanti et al. (2010). Philometroides maplestoni (Travassos, Artigas & Pereira)adultFWbody cavity Syn.: Philometra maplestoni Travassos, Artigas & Pereira Host: Salminus hilarii. Locality: PirassunungaSP. References: Travassos et al. (1928), Yamaguti (1961), Travassos & Kohn (1965), Vicente & Jardim (1980), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Rumai rumai Travassosadult and larvaFWbody cavity, encapsulated in tissues of mouth, tongue, operculum and head Host: Arapaima gigas. Locality: Amazonas Basin River, Mexiana IslandPA. References: Travassos (1960), Vicente & Pinto (1981), Vicente et al. (1985), Moravec (1998), Santos & Moravec (2009c), Eiras et al. (2010).

Suborder Spirurina Railliet & Henry Superfamily Acuaroidea Railliet, Henry & Sisoff Family Acuariidae Railliet, Henry & Sisoff

Acuariidae gen. sp.larvaFWstomach Host: Salminus brasiliensis. Locality: Foz do Igua, Paran RiverPR. References: Moravec et al. (1993a), Moravec (1998), Vicente & Pinto (1999) Remark: Moravec et al. (1993a) refered to this taxon as Acuariinae gen. sp.

Superfamily Habronematoidea Railliet & Henry Family Cystidicolidae Skrjabin

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Ascarophis brasiliensis Pinto, Vicente & NoronhaadultMARstomach Host: Trachinotus carolinus. Locality: AraruamaRJ. References: Pinto et al. (1984), Vicente et al. (1985). Ascarophis marina (Szidat)adultMARintestine Host: Urophycis brasiliensis. Locality: Rio de JaneiroRJ. Reference: Alves et al. (2004). Cystidicoloides fischeri (Travassos, Artigas & Pereira)adultFWstomach Syn.: Cystidicola fisheri Travassos, Artigas & Pereira Hosts: Galeocharax knerii, Pygocentrus piraya, Salminus brasiliensis, S. hilarii, Serrasalmus brandtii. Localities: Upper So Francisco RiverMG; PirassunungaSP. References: Travassos et al. (1928), Yamaguti (1961), Travassos & Kohn (1965), Bhrnheim (1976), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991), Moravec (1998), Moravec et al. (2008), Eiras et al. (2010). Cystidicoloides izecksohni (Fbio)adultFWintestine Syn.: Heliconema izecksohni Fbio Host: Hoplias malabaricus. Locality: CamposRJ. References: Fbio (1982), Vicente et al. (1985), Moravec et al. (2008), Eiras et al. (2010). Cystidicoloides sp.adult and larvaFW Host: Brycon orthotaenia (adultintestine), Leporinus friderici (larvamesentery). Localities: Upper Paran RiverPR; PirassunungaSP. References: Kohn & Fernandes (1987), Guidelli et al. (2006), Takemoto et al. (2009), Eiras et al. (2010). Parascarophis sphyrnae Campana-RougetadultMARspiral valve Host: Sphyrna zygaena. Locality: CoastRS. Reference: Knoff et al. (2001). Pseudoproleptus sp.larvaFWmesentery Hosts: Satanoperca jurupari, Ageneiosus ucayalensis. Locality: Guam RiverPA. Reference: Melo et al. (2011a). Spinitectus asperus Travassos, Artigas & PereiraadultFWintestine, stomach Syn.: S. jamundensis Thatcher & Padilha Hosts: Leporinus striatus, Prochilodus argenteus, P. lineatus. Localities: Upper So Francisco RiverMG; PantanalMS; Guara, Paran RiverPR; Pirassununga SP. References: Travassos et al. (1928), Yamaguti (1961), Travassos & Kohn (1965), Bhrnheim (1976), Thatcher & Padilla (1977), Vicente & Pinto (1981), Fernandes et al. (1982), , Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006), Moravec et al. (1993a), Moravec (1998), Santos et al. (2003), Monteiro et al. (2009), Eiras et al. (2010). Spinitectus rodolphiheringi Vaz & PereiraadultFWintestine Syn.: S. sternopygi Petter, 1984 Hosts: Astyanax bimaculatus, Franciscodoras marmoratus, Pimelodella lateristriga, Pimelodus maculatus, Salminus hilarii, Sternopygus macrurus, Triportheus paranensis. Localities: Grande & Mogi Guau Rivers, PirassunungaSP. References: Vaz & Pereira (1934), Yamaguti (1961), Bhrnheim (1976), Vicente & Pinto (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010).

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Spinitectus yorkei Travassos, Artigas & PereiraadultFWintestine Host: Pimelodella lateristriga. Locality: PirassunungaSP. References: Travassos et al. (1928), Yamaguti (1961), Travassos & Kohn (1965), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Spinitectus sp.adultFW Hosts and site of infection: Franciscodoras marmoratus (intestine), Potamotrygon falkneri (spiral valve). Localities: Trs MariasMG; Upper Paran RiverPR. References: Santos & Brasil-Sato (2004), Lacerda et al. (2008, 2009), Takemoto et al. (2009), Eiras et al. (2010).

Family Hedruridae Railliet


Hedruris iheringi Pereira & VazadultFWintestine Host: Austrolebias bellottii Locality: SP. Reference: Pereira & Vaz (1933).

Superfamily Gnathostomatoidea Railliet Family Gnathostomatidae Railliet Subfamily Ancyracanthinae Yorke & Maplestone
Ancyracanthus schubarti (Kohn, Gomes & Motta)adultFW Syn.: Piavussunema schubarti Kohn, Gomes & Motta Hosts and site of infection: Leporellus vittatus (stomach), Leporinus copelandii (stomach), L. lacustris (intestine), L. octofasciatus (intestine, stomach), Schizodon borellii (intestine). Localities: Upper Paran RiverPR; PirassunungaSP. References: Kohn et al. (1968), Gomes & Kohn (1970), Bhrnheim (1976), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Machado et al. (1996), Pavanelli et al. (1997, 2004), Moravec (1998), Guidelli et al. (2006), Takemoto et al. (2009), Eiras et al. (2010).

Subfamily Gnathostomatinae Railliet


Echinocephalus daileyi Deardoff, Brooks & ThorsonadultFW intestine Hosts: Potamotrygon hystrix, P. motoro. Locality: Itacuai RiverAM. References: Deardorff et al. (1981), Vicente et al. (1985), Thatcher (1991, 2006), Moravec (1998), Eiras et al. (2010). Echinocephalus sp.adultFWspiral valve Host: Potamotrygon falkneri. Locality: Upper Paran RiverPR. References: Lacerda et al. (2008, 2009), Takemoto et al. (2009), Eiras et al. (2010).

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Gnathostoma gracile (Diesing)adultFWintestine Syn.: Cheiracanthus gracilis Diesing Host: Arapaima gigas. Locality: Amazonas River Basin. References: Diesing (1839), Travassos et al. (1928), Vicente et al. (1985), Moravec (1998), Eiras et al. (2010).

Subfamily Spiroxyinae Baylis & Lane


Spiroxys sp.larvaFW Hosts and site of infection: Astyanax altiparanae (intestine), Cichla kelberi (body cavity, intestine), Gymnotus carapo, Gymnotus sp. (intestine), Pygocentrus piraya (body cavity, intestinal caecum, intestine, gall bladder), Serrasalmus brandtii (body cavity, stomach, intestine). Localities: Trs Marias Reservoir, So Francisco RiverMG; Baa RiverMS; Upper Paran RiverPR. References: Isaac et al. (2004), Pavanelli et al. (2004), Santos et al. (2009), Takemoto et al. (2009), Eiras et al. (2010).

Superfamily Physalopteroidea Railliet Family Physalopteridae Railliet Subfamily Proleptinae Schulz


Heliconema heliconema TravassosadultMARintestine and stomach Hosts: Echidna catenata, Ophichthus gomesii, O. ophis. Localities: Campos, Copacabana and Urcha BeachRJ; SalvadorBA; Trindade IslandES. References: Travassos (1919), Yamaguti (1961), Guimares et al. (1976), Vicente et al. (1985), Fernandes et al. (2002). Proleptus acutus DujardinadultMARstomach, intestine Host: Zapteryx brevirostris. Locality: UbatubaSP. References: Silva et al. (2008), Muniz-Pereira et al. (2009).

Superfamily Thelazioidea Oerley Family Rhabdochonidae Skrjabin


Johnstonmawsonia sp.adultFWUnspecified site of infection Host: Auchenipterus osteomystax. Locality: Upper Paran RiverPR. References: Takemoto et al. (2009), Tavernari et al. (2009), Eiras et al. (2010). Rhabdochona acuminata (Molin)adultFW Syn.: Spiroptera acuminata Molin; Rhabdochona elegans Travassos, Artigas & Pereira; R. fasciata Kloss; R. australis Kloss; R. siluriformes Kloss Hosts and site of infection: Astyanax bimaculatus (intestine), A. fasciatus (intestine), A. schubarti (intestine), Auchenipterus osteomystax, Brycon falcatus (intestine), Geophagus brasiliensis (intestine),

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Leporellus vittatus (intestine), Pimelodella lateristriga (intestine, gall bladder), Tatia neivai (intestine), Tetragonopterinae gen. sp. (intestine), Tetragonopterus argenteus, Tetragonopterus sp. (intestine). Localities: MT; Lajes Reservoir, PiraRJ; Grande, Tiet, PirassunungaSP References: Molin (1860), Drasche (1884b), Travassos et al. (1928), Vaz & Pereira (1934), Yamaguti (1961), Travassos & Kohn (1965), Kloss (1966), Bhrnheim (1976), Vicente & Pinto (1981), Vicente et al. (1985), Kohn & Fernandes (1987), Moravec (1972, 1998), Thatcher (1991, 2006), Paraguass et al. (2005), Paraguass & Luque (2006), Takemoto et al. (2009), Tavernari et al. (2009), Eiras et al. (2010), Pinto et al. (2010). Remark: Drasche (1884b) did not mention a host and locality, but redescribed Molins type-material. Rhabdochona obtusus Dujardin Host: Tetragonopterus argenteus. Locality: PantanalMT. Reference: Rgo & Vicente (1988b). Rhabdochona uruyeni Diaz-UngriaadultFW - intestine Hosts: Pachyurus squamipennis, Pimelodus maculatus. Locality: Lagoa Juparan, LinharesES, Guandu RiverRJ. Reference: Azevedo et al. (2010), Pinto et al. (2010). Rhabdochona sp.adult and larvaFWintestine Hosts and site of infection: Brycon orthotaenia (adultintestine), Conorynchus conirostris (larva intestine), Centropomus undecimalis, Leporinus octofasciatus (adultintestine), Tetragonopterus argenteus (stomach). Localities: PantanalMT; PirassunungaSP; So Francisco RiverMG; Guandu RiverRJ. References: Kloss (1966), Bhrnheim (1976), Kohn & Fernandes (1987), Brasil-Sato & Santos (2005), Azevedo et al. (2010).

Final remarks:
Two records founded in our bibliographic searches are clearly misidentified or could be accidental associations. Kohn & Macedo (1984) recorded the oxyurid Aspiculurus tetraptera (Nitzsch, 1821), a common parasite of rodents, in Haemulon sciurus off Rio de Janeiro. Another record, that of Raphidascaris (Raphidascaris) acus (Bloch, 1779) recorded by Guimares (1975) in Lutjanus vivanus off the Salvador (Bahia), is considered a misidentification, because this is a Holarctic freshwater species (Anderson, 2000).

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HostParasite List

Hosts Class Actinopterygii Cara suja Lanceta Sardinha Sardinha Boca Larga Sardinha Maromba Order Anguilliformes Family Muraenidae Echidna catenata (Bloch) Gymnothorax moringa (Cuvier)

Parasite species Procamallanus (S.) macaenses Porrocaecum secundum Porrocaecum sp. Procamallanus (S.) barroslimai Raphidascaris sp. Raphidascaris sp.

Heliconema heliconema Hysterothylacium sp. Raphidascaris sp. Terranova sp.

Family Ophichthidae Ophichthus ophis (Linnaeus) O. gomesii (Castelnau) Order Batrachoidiformes Family Batrachoididae Porichthys porosissimus (Cuvier) Order Beloniformes Family Belonidae Tylosurus acus acus (Lacpde) Hysterothylacium sp. Terranova sp. Procamallanus (S.) pereirai Heliconema heliconema Heliconema heliconema

Order Clupeiformes Family Clupeidae Clupeidae gen. sp. Brevoortia aurea (Spix & Agassiz) Procamallanus (S.) barroslimai Anisakis sp. Contracaecum sp. Procamallanus sp. Terranova sp. Hysterothylacium fortalezae Raphidascaris (I.) vicentei Raphidascaris sp. Raphidascaris (I.) vicentei Cosmocercinae gen. sp.

Harengula clupeola (Cuvier)

Sardinella sp.

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Family Engraulidae Anchoa marinii Hildebrand Anchoa tricolor (Spix & Agassiz) Hysterothylacium sp. Raphidascaris sp. Hysterothylacium sp. Terranova sp.

Family Pristigasteridae Pellona castelnaeana Valenciennes Order Characiformes Family Acestrorhynchidae Acestrorhynchus falcatus (Bloch) Acestrorhynchus lacustris (Ltken) Paracapillaria (P.) piscicola Procamallanus (S.) inopinatus Anisakidae gen. sp. Contracaecum sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Eustrongylides sp. Philometridae gen. sp. Philonema sp. A Procamallanus (S.) sp. Procamallanus (S.) saofranciscensis Procamallanus sp. Travassosnema travassosi paranaensis Travassosnema travassosi travassosi Procamallanus (S.) hilarii Paracapillaria (P.) piscicola Procamallanus sp. Procamallanus (S.) sp. Nematoda gen. sp.

Acestrorhynchus microlepis (Jardine) Acestrorhynchus sp. Peixe-cachorro

Family Anostomidae Anostominae gen. sp. Chimbor Ferreirinha Leporellus vittatus (Valenciennes) Capillostrongyloides sentinosa Procamallanus (S.) iheringi Procamallanus (S.) sp. Procamallanus (S.) sp. Ancyracanthus schubarti Capillaria sp. Contracaecum sp. Cucullanus sp. Cucullanus sp. A Filocapsulariinae gen. sp. Goezia spinulosa Nematoda gen. sp. Paracapillaria (P.) piscicola Procamallanus (S.) inopinatus Rhabdochona acuminata

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Leporinus agassizii Steindachner Leporinus copelandii Steindachner

Procamallanus (S.) inopinatus Amplicaecum sp. Ancyracanthus schubarti Capillaria sp. Contracaecum sp. Contracaecum sp. Type B Cucullanus (C.) brevispiculus Cucullanus (C.) mogi Cucullanus (C.) pinnai pinnai Cucullanus sp. Eustrongylides sp. Filocapsulariinae gen. sp. Goezia spinulosa Klossinemella iheringi Paracapillaria (P.) piscicola Procamallanus (S.) amarali Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. A Brevimulticaecum sp. Capillostrongyloides sentinosa Contracaecum sp. Goezia spinulosa Nematoda gen. sp. Porrocaecum sp. Procamallanus (S.) amarali Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Klossinemella iheringi Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Brevimulticaecum sp. Contracaecum sp. Contracaecum sp. Type 1 Cystidicoloides sp. Dichelyne (D.) leporini Goezia brevicaeca Goezia spinulosa Hysterothylacium sp. Nematoda gen. sp. Procamallanus (S.) amarali Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. Ancyracanthus schubarti Brevimulticaecum sp. Brevimulticaecum sp. of Moravec et al. (1997) Contracaecum sp. Contracaecum sp. Type B Contracaecum sp. Type 1

Leporinus elongatus Valenciennes

Leporinus fasciatus (Bloch)

Leporinus friderici (Bloch)

Leporinus lacustris Amaral Campos

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Leporinus lacustris Amaral Campos

Dichelyne (D.) leporini Nematoda gen. sp. Procamallanus (S.) inopinatus Procamallanus (S.) sp. A of Bhrnheim (1976) Procamallanus (S.) sp. B of Bhrnheim (1976) Goezia leporini Procamallanus sp. Brevimulticaecum sp. Contracaecum sp. Eustrongylides sp. Goezia spinulosa Procamallanus (S.) amarali Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. Nematoda gen. sp. Ancyracanthus schubarti Filocapsulariinae gen. sp. Nematoda gen. sp. Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. B of Bhrnheim (1976) Procamallanus sp. Rhabdochona sp. Procamallanus (S.) inopinatus Procamallanus (S.) inopinatus Capillostrongyloides sentinosa Cucullanus (C.) mogi Filocapsulariinae gen. sp. Nematoda gen. sp. Procamallanus sp. Procamallanus (S.) amarali Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. B of Bhrnheim (1976) Procamallanus sp. Procamallanus (S.) inopinatus Procamallanus (S.) sp. A Spinitectus asperus Filocapsulariinae gen. sp. Contracaecum sp. Procamallanus (S.) inopinatus Procamallanus sp. Ancyracanthus schubarti Cucullanus (C.) pinnai pinnai Procamallanus (S.) iheringi Procamallanus (S.) inopinatus

Leporinus macrocephalus Garavello & Britski Leporinus obtusidens (Valenciennes)

Leporinus octofasciatus Steindachner

Leporinus piau Fowler Leporinus reinhardti Ltken Leporinus sp.

Leporinus striatus Kner

Leporinus taeniatus Ltken Schizodon altoparanae Garavello & Britski Schizodon borellii (Boulenger)

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Schizodon fasciatus Spix & Agassiz

Capillariidae gen. sp. 1 Dichelyne (D.) leporini Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Capillostrongyloides sentinosa Contracaecum sp. Cucullanus sp. A Klossinemella iheringi Procamallanus (P.) peraccuratus Procamallanus (S.) iheringi Procamallanus (S.) inopinatus

Schizodon knerii (Steindachner) Schizodon nasutus Kner

Family Characidae Aphyocharax anisitsi Eigenmann & Kennedy Astyanax altiparanae Garutti & Britski Nematoda gen. sp. Contracaecum sp. Contracaecum sp. Type 1 Procamallanus (S.) caballeroi Procamallanus (S.) inopinatus Spiroxys sp. Camallanidae gen. sp. Contracaecum sp. Contracaecum sp. Type A Contracaecum sp. Type B Cosmoxynemoides aguirrei Cucullanus sp. Nematoda gen. sp. Paraseuratum albidum Procamallanus (S.) hilarii Procamallanus (S.) inopinatus Procamallanus (S.) sp. Procamallanus (S.) sp. (= "P. laticonchus") Procamallanus sp. Rhabdochona acuminata Spinitectus rodolphiheringi Travnema travnema Nematoda gen. sp. Ascarididae gen. sp. Capillaria sp. Capillostrongyloides sentinosa Contracaecum sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Paraseuratum albidum Procamallanus (S.) hilarii Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) saofranciscensis Procamallanus (S.) sp.

Astyanax bimaculatus (Linnaeus)

Astyanax eigenmanniorum (Cope) Astyanax fasciatus (Cuvier)

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Astyanax fasciatus (Cuvier)

Procamallanus (S.) sp.(= "P. laticonchus") Procamallanus sp. Rhabdochona acuminata Procamallanus (S.) inopinatus Procamallanus (S.) hilarii Contracaecum sp. Contracaecum sp. Type A Goezia spinulosa Procamallanus (S.) hilarii Procamallanus (S.) inopinatus Procamallanus (S.) sp. Rhabdochona acuminata Procamallanus (S.) inopinatus Procamallanus sp. Procamallanus (S.) inopinatus Procamallanus (S.) inopinatus Procamallanus (S.) inopinatus Rhabdochona acuminata Camallanidae gen. sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Cucullanus sp. Eustrongylides ignotus Goezia brasiliensis Goezia brevicaeca Neocucullanus neocucullanus Procamallanus (S.) inopinatus Contracaecum sp. Eustrongylides sp. Nematoda gen. sp. Anisakidae gen. sp. Nematoda gen. sp. Amplicaecum sp. Cystidicoloides sp. Procamallanus (S.) inopinatus Rhabdochona sp. Procamallanus sp. Nematoda gen. sp. Procamallanus (S.) inopinatus Procamallanus (D.) dentatus Procamallanus (S.) sp. Nematoda gen. sp. Camallanus sp. Eustrongylides sp. Procamallanus (S.) inopinatus

Astyanax lacustris (Ltken) Astyanax parahybae Eigenmann Astyanax schubarti Britski

Astyanax sp. Brycon amazonicus (Spix & Agassiz) Brycon cephalus (Gnther) Brycon falcatus Mller & Troschel Brycon hilarii (Valenciennes)

Brycon microleps (Valenciennes)

Brycon orbignyanus (Valenciennes) Brycon orthotaenia Gnther

Brycon insignis Steindachner Brycon sp. Bryconops alburnoides Kner Triportheus nematurus (Kner) Charax gibbosus (Linnaeus)

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Colossoma macropomum (Cuvier) Cynopotamus kincaidi (Schultz) Galeocharax humeralis (Valenciennes)

Procamallanus (S.) sp. Procamallanus sp. Nematoda gen. sp. Contracaecum sp. Contracaecum sp. Type 1 Eustrongylides ignotus Eustrongylides sp. Paracapillaria (P.) piscicola Procamallanus (S.) inopinatus Cucullanidae gen. sp. Contracaecum sp. Contracaecum sp. Type A Contracaecum sp. Type B Contracaecum sp. Type 1 Cucullanus (C.) pinnai pinnai Cystidocoloides fischeri Goezia spinulosa Hysterothylacium sp. Paracapillaria (P.) piscicola Filocapsulariinae gen. sp. Contracaecum sp. Type 2 Contracaecum sp. Procamallanus (S.) inopinatus Raphidascaris (S.) mahnerti Spinoxyuris oxydoras Camallanidae gen. sp. Klossinemella iheringi Rondonia rondoni Rondonia rondoni Rondonia sp. Spectatus spectatus Nematoda gen.sp. Klossinemella iheringi Procamallanus (S.) inopinatus Rondonia rondoni Brevimulticaecum sp. Nematoda gen. sp. Rondonia rondoni Procamallanus (S.) inopinatus Ichthyouris voltagrandensis Rondonia rondoni Klossinemella iheringi Myleusnema brasiliense Rondonia rondoni Rondonia sp. Spinoxyuris annulata

Galeocharax knerii (Steindachner)

Markiana geayi (Pellegrin) Metynnis lippincottianus (Cope)

Moenkhausia doceana (Steindachner) Mylesinus paraschomburgkii Jgu, Santos & Ferreira Myletes sp.

Myleus asterias (Mller & Troschel)

Myleus levis Eigenmann & McAtee Myleus micans (Ltken) Myleus schomburgkii (Jardine) Myleus tiete (Eigenmann & Norris) Myleus torquatus (Kner) Myleus sp.

Mylossoma aureum (Spix & Agassiz)

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Mylossoma duriventre (Cuvier) Oligosarcus macrolepis (Steindachner) Oligosarcus paranensis Menezes & Gry Oligosarcus sp.

Goezia sp. Spectatus sp. Contracaecum sp. Procamallanus (S.) hilarii Contracaecum sp. Contracaecum sp. Eustrongylides sp. Paracapillaria (P.) piscicola Procamallanus sp. Klossinemella iheringi Rondonia rondoni Rondonia sp. Spectatus spectatus Nematoda gen. sp. Goezia leporine Porrocaecum sp. Rondonia rondoni Spectatus spectatus Spectatus sp. Chabaudinema americanum Contracaecum sp. Contracaecum sp. Type 2 Ascarididae gen. sp. Brevimulticaecum sp. Camallanidae gen. sp. Contracaecum sp. Eustrongylides sp. Eustrongylides ignotus Procamallanus (S.) inopinatus Procamallanus sp. Brevimulticaecum sp. Cystidocoloides fischeri Hysterothylacium sp. Procamallanus (S.) inopinatus Spiroxys sp. Procamallanus (S.) inopinatus Nematoda gen. sp. Acuariidae gen. sp. Anisakidae gen. sp. Ascarididae gen. sp. Ascaridoidea fam. gen. sp. Contracaecum sp. Contracaecum sp. Type A Contracaecum sp. Type B Contracaecum sp. Type 2 Cystidocoloides fischeri Eustrongylides ignotus

Paracheirodon axelrodi (Schultz) Piaractus brachypomus (Cuvier)

Piaractus mesopotamicus (Holmberg)

Psellogrammus kennedyi (Eigenmann) Pygocentrus cariba (Humboldt) Pygocentrus nattereri Kner

Pygocentrus piraya (Cuvier)

Pygocentrus sp. Roeboides paranensis Pignalberi Salminus brasiliensis (Cuvier)

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Salminus brasiliensis (Cuvier)

Eustrongylides sp. Freitascapillaria maxillosa Goezia spinulosa Hysterothylacium sp. Neocucullanus neocucullanus Paracapillaria (P.) piscicola Procamallanus (S.) inopinatus Procamallanus (S.) sp. (= " P. laticonchus") Amplicaecum sp. Contracaecum sp. Contracaecum sp. Type A Contracaecum sp. Type 2 Cystidocoloides fischeri Filocapsulariinae gen. sp. Klossinemella iheringi Necocucullanus neocucullanus Paracapillaria (P.) piscicola Philometroides maplestoni Procamallanus (S.) hilarii Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. Procamallanus sp. Spinitectus rodolphiheringi Rondonia rondoni Travnema araujoi Contracaecum sp. Cystidocoloides fischeri Hysterothylacium sp. Procamallanus (S.) inopinatus Spiroxys sp. Procamallanus (S.) inopinatus Capillariidae gen. sp. Cucullanus sp. Eustrongylides ignotus Philometridae gen. sp. Procamallanus (S.) inopinatus Procamallanus (S.) inopinatus Brevimulticaecum sp. Capillaria sp. Capillariidae gen. sp. Contracaecum sp. Cucullanus sp. Cucullanus sp. 3 Eustrongylides ignotus Goezia sp. Philometridae gen. sp. Procamallanus sp.

Salminus hilarii Valenciennes

Salminus sp. Serrasalmus altuvei Ramrez Serrasalmus brandtii Ltken

Serrasalmus gouldingi Fink & Machado-Allison Serrasalmus maculatus Kner

Serrasalmus manueli (Fernndez-Ypez & Ramrez) Serrasalmus marginatus Valenciennes

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Serrasalmus marginatus Valenciennes Serrasalmus spilopleura Kner

Procamallanus (S.) inopinatus Capillariinae gen. sp. Cucullanus sp. Eustrongylides ignotus Nematoda gen. sp. Philometridae gen. sp. Procamallanus sp. Procamallanus (S.) inopinatus Procamallanus (S.) rarus Contracaecum sp. Nematoda gen. sp. Paraseuratum albidum Rhabdochona acuminata Rhabdochona obtusus Rhabdochona sp. Hysterothylacium sp. Procamallanus (S.) saofranciscensis Procamallanus (S.) iheringi Rhabdochona acuminata Eustrongylides sp. Klossinemella iheringi Procamallanus (S.) iheringi Rhabdochona acuminata Nematoda gen. sp. Hysterothylacium sp. Procamallanus (S.) saofranciscensis Nematoda gen. sp. Procamallanus (S.) inopinatus Spinitectus rodolphiheringi Procamallanus (S.) barroslimai Neocucullanus neocucullanus Procamallanus (S.) iheringi

Serrasalmus sp. Tetragonopterus argenteus Cuvier

Tetragonopterus chalceus Spix & Agassiz Tetragonopterus sp. Tetragonopterinae gen. sp.

Triportheus angulatus (Spix & Agassiz) Triportheus guentheri (Garman) Triportheus paranensis (Gunther)

Triportheus sp. Characidae gen.sp.

Family Curimatidae Curimata sp. Curimatella lepidura (Eigenmann & Eigenmann) Cyphocharax gilbert (Quoy & Gaimard) Contracaecum sp. Philonema sp. B Cosmoxynemoides aguirrei Camallanidae gen. sp. Cosmoxynema vianai Cosmoxynemoides aguirrei Guyanema sp. Nematoda gen. sp. Philonema sp. B Raphidascaris sp. Travnema araujoi Travnema travnema Travnema sp.

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Cyphocharax nagelii (Steindachner) Cyphocharax plumbeus (Eigenmann & Eigenmann) Potamorhina squamoralevis (Braga & Azpelicueta) Pseudocurimata sp.

Nematoda gen. sp. Travnema travnema Nematoda gen. sp. Cosmoxynema vianai Cosmoxynemoides aguirrei Philonema sp. B Travnema sp. Cosmoxynema vianai Travnema travnema Spinoxyuris sp. Cosmoxynema vianai Cosmoxynemoides aguirrei Procamallanus (S.) hilarii Travnema araujoi Travnema travnema Travnema sp. Cosmoxynema vianai Travnema travnema

Steindachnerina brevipinna (Eigenmann & Eigenmann) Steindachnerina elegans (Steindachner)

Steindachnerina insculpta (FernndezYpez)

Family Cynodontidae Rhaphiodon vulpinus Spix & Agassiz Ascarididae gen. sp. Contracaecum sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Eustrongylides ignotus Eustrongylides sp. Goezia sp. Guyanema raphiodoni Hysterothylacium sp. Nematoda gen. sp.

Family Erythrinidae Erythrinus erythrinus (Bloch & Schneider) Hoplerythrinus unitaeniatus (Spix & Agassiz) Paraseuratum albidum Procamallanus (S.) paraensis Contracaecum sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Eustrongylides sp. Nematoda gen. sp. Procamallanus (S.) hilarii Amplicaecum sp. Ascarididae gen sp. Ascaridoidea fam. gen. sp. Brevimulticaecum sp. Capillostrongyloides sentinosa

Hoplias lacerdae Miranda Ribeiro Hoplias malabaricus (Bloch)

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Hoplias malabaricus (Bloch)

Contracaecum sp. Contracaecum sp. Type B Contracaecum sp. Type 1 Contracaecum sp. Type 2 Cystidicoloides izecksohni Eustrongylides ignotus Eustrongylides sp. Filariidae gen. sp. Filocapsulariinae gen. sp. Goezia spinulosa Klossinemella iheringi Nematoda gen. sp. Paracapillaria (P.) piscicola Paraseuratum soaresi Philonema sp. C Porrocaecum sp. Procamallanus (S.) hilarii Procamallanus (S.) iheringi Procamallanus (S.) inopinatus Procamallanus (S.) sp. Procamallanus (P.) peraccuratus Procamallanus sp. Eustrongylides ignotus Procamallanus (S.) iheringi Procamallanus (S.) paraensis

Hoplias sp. Jeju Family Hemiodontidae Bivibranchia fowleri (Steindachner) Bivibranchia velox (Eigenmann & Myers) Family Prochilodontidae Prochilodus argenteus Spix & Agassiz Prochilodus lineatus (Valenciennes)

Procamallanus (S.) sp. Procamallanus (S.) sp.

Spinitectus asperus Anisakidae gen. sp. Contracaecum sp. Contracaecum sp. Type 2 Procamallanus (S.) inopinatus Procamallanus (S.) sp. Procamallanus sp. Raphidascaris sp. Spinitectus asperus

Order Cypriniformes Family Rhamphichthyidae Rhamphichthys rostratus (Linnaeus) Rhamphichthys hahni (Meinken) Cucullanus (T.) rhamphichthydis Nematoda gen. sp.

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Order Cyprinodontiformes Family Poeciliidae Poecilia latipinna (Lesueur) Poecilia reticulata Peters Poecilia sphenops Valenciennes Poecilia vivipara Bloch & Schneider Xiphophorus helleri Heckel Xiphophorus maculatus (Gnther) Poeciliidae gen. sp. Family Rivulidae Austrolebias bellottii (Steindachner) Order Gadiformes Family Merluccidae Merluccius hubbsi Marini Anisakis sp. Hysterothylacium sp. Raphidascaris sp. Hedruris iheringi Camallanus cotti Camallanus cotti Camallanus maculatus Camallanus sp. Eustrongylides sp. Camallanus cotti Camallanus maculatus Eustrongylides ignotus

Family Phycidae Urophycis brasiliensis (Kaup) Ascarophis marina Contracaecum sp. Cucullanus sp. Hysterothylacium sp. Procamallanus (S.) macaensis Raphidascaris sp. Anisakis sp. Contracaecum sp. Cucullanus cirratus Hysterothylacium sp. Procamallanus (S.) macaensis

Urophycis mystacea Miranda Ribeiro

Urophycis sp. Order Gymnotiformes Family Gymnotidae Gymnotus carapo Linnaeus

Capillariinae gen. sp. Contracaecum sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Eustrongylides sp. Hysterothylacium sp. Nematoda gen. sp. Procamallanus (P.) peraccuratus Spiroxys sp.

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Gymnotus inaequilabiatus (Valenciennes) Gymnotus sp.

Brevimulticaecum sp. Contracaecum sp. 1 Contracaecum sp. 2 Eustrongylides sp. 1 Eustrongylides sp. 2 Hysterothylacium sp. Spiroxys sp. Ascaridoidea fam. gen. sp.

Electrophorus electricus (Linnaeus) Family Sternopygidae Sternopygus macrurus (Bloch & Schneider) Order Lampriformes Family Trachipteridae Trachipterus jacksonensis (Ramsay) Order Lophiiformes Family Lophiidae Lophius gastrophysus Miranda Ribeiro

Spinitectus rodolphiheringi

Lappetascaris lutjani

Hysterothylacium sp. Nematoda gen. sp. Raphidascaris sp.

Order Ophidiiformes Family Ophidiidae Genypterus brasiliensis Regan Anisakidae gen. sp. Anisakis physeteris Anisakis simplex Anisakis sp. Contracaecum sp. Cucullanus genypteri Hysterothylacium sp. Nematoda gen. sp. Procamallanus sp. Pseudoterranova decipiens Pseudoterranova sp. Raphidascaris sp. Terranova sp.

Order Osteoglossiformes Family Arapaimidae Arapaima gigas (Schinz) Camallanus (C.) tridentatus Capillostrongyloides arapaimae Eustrongylides sp. Gnathostoma gracile Goezia spinulosa Nilonema senticosum

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Arapaima gigas (Schinz)

Rumai rumai Terranova serrata

Family Osteoglossidae Osteoglossum bicirrhosum (Cuvier) Camallanus (C.) acaudatus Camallanus (C.) tridentatus

Order Perciformes Acar-a Car-cachimbo Caranx hippos (Linnaeus) Goezia spinulosa Procamallanus sp. Procamallanus (S.) sp. Contracaecum sp. Cucullanus pulcherrimus Philometra sp. Raphidascaris sp. Terranova sp. Anisakis sp. Contracaecum sp. Hysterothylacium sp. Terranova sp. Cucullanus pulcherrimus Contracaecum sp. Contracaecum sp. Contracaecum sp. Porrocaecum secundum Terranova trichiuri Contracaecum sp. Hysterothylacium sp. Raphidascaris sp. Anisakis sp. Contracaecum sp. Hysterothylacium sp. Raphidascaris sp. Terranova sp. Porrocaecum sp. Raphidascaris sp. Ascarophis brasiliensis Contracaecum sp. Terranova sp. Hysterothylacium sp. Raphidascaris sp. Raphidascaris sp.

Caranx latus Agassiz

Caranx lugubris Poey Oligoplites palometa (Cuvier) Oligoplites saliens (Bloch) Oligoplites saurus (Bloch & Schneider)

Parona signata (Jenyns)

Selene setapinnis (Mitchill)

Selene vomer (Linnaeus) Trachinotus carolinus (Linnaeus) Trachurus lathami Nichols Uraspis secunda (Poey) Scombroides occidentalis (Bloch & Schneider) Family Centrarchidae Micropterus salmoides (Lacpde)

Goezia spinulosa

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Family Centropomidae Centropomus parallelus Poey Centropomus undecimalis (Bloch) Angusticaecum sp. Contracaecum sp. Cucullanus sp. Rhabdochona sp.

Family Cichlidae Apistogramma cacatuoides Hoedeman Astronotus ocellatus (Agassiz) Camallanus cotti Camallanus sp. Contracaecum sp. Contracaecum sp. Type 2 Goezia spinulosa Nematoda gen. sp. Procamallanus (S.) inopinatus Procamallanus (P.) peraccuratus Procamallanus sp. Contracaecum sp. Contracaecum sp. Type 1 Eustrongylides sp. Hysterothylacium sp. Procamallanus (S.) inopinatus Spiroxys sp. Contracaecum sp. Eustrongylides sp. Contracaecum sp. Eustrongylides sp. Procamallanus (P.) peraccuratus Contracaecum sp. Type 1 Eustrongylides sp. Procamallanus (P.) peraccuratus Ichthyouris ovofilamentosa Ichthyouris sp. Nematoda gen. sp. Procamallanus (S.) inopinatus Contracaecum sp. Type 1 Contracaecum sp. Hysterothylacium sp. Procamallanus (P.) peraccuratus Nematoda gen. sp. Contracaecum sp. Cosmoxynemoides aguirrei Nematoda gen. sp. Procamallanus (P.) peraccuratus Raphidascaris (S.) sp. Rhabdochona acuminata Raphidascaris (S.) sp.

Australoheros facetus (Jenyns) Cichla kelberi Kullander & Ferreira

Cichla monoculus Spix & Agassiz Cichla ocellaris Bloch & Schneider

Cichla piquiti Kullander & Ferreira Cichlasoma facetum (Jenyns) Cichlasoma sp. Crenicichla britskii Kullander Crenicichla haroldoi (Luengo & Britski) Crenicichla lepidota Heckel

Crenicichla niederleinii (Holmberg) Geophagus brasiliensis (Quoy & Gaimard)

Geophagus proximus (Castelnau)

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Geophagus steindachneri Eigenmann & Hildebrand Oreochromis niloticus niloticus (Linnaeus) Pterophyllum scalare (Schultze) Satanoperca jurupari (Heckel) Satanoperca pappaterra (Heckel) Symphysodon aequifasciatus Pellgrin Family Coryphaenidae Coryphaena hippurus Linnaeus Family Ephippidae Chaetodipterus faber (Broussonet)

Cosmoxynemoides aguirrei Nematoda gen. sp. Camallanus cotti Capillarinae gen. sp. Raphidascaris (Sprentascaris) lanfrediae Pseudoproleptus sp Nematoda gen. sp. Ichthyouris bursata

Nematoda gen. sp.

Porrocaecum sp. Procamallanus (S.) macaensis

Family Gempylidae Thyrsitops lepidopoides (Cuvier) Nematoda gen. sp. Procamallanus (S.) macaensis Raphidascaris sp.

Family Gerreidae Diapterus rhombeus (Cuvier) Contracaecum sp. Cucullanus sp.

Family Haemulidae Haemulon plumierii (Lacpde) Family Labridae Halichoeres radiatus (Linnaeus) Family Lutjanidae Lutjanus synagris (Linnaeus) Raphidascaris (I.) vicentei Philometra sp. Dichelyne travassosi Philometra lateolabracis

Family Mugilidae Mugil curema Valenciennes Mugil liza Valenciennes Mugil platanus Gnther Cucullanus sp. Cucullanus (C.) grandistomis Hysterothylacium sp. Cucullanus pulcherrimus

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Family Mullidae Mullus argentinae Hubbs & Marini Anisakis sp. Contracaecum sp. Procamallanus (S.) cruzi Raphidascaris sp. Capillaria sp. Nematoda gen. sp.

Pseudupeneus maculatus (Bloch)

Family Osphronemidae Betta splendens Regan Family Percophidae Percophis brasiliensis Quoy & Gaimard Anisakis sp. Contracaecum sp. Hysterothylacium sp. Procamallanus sp. Raphidascaris sp. Terranova sp. Camallanus cotti

Family Pinguipedidae Pinguipes brasilianus Cuvier Cucullanus carioca Hysterothylacium sp. Paracapillaria (P.) argentinensis Raphidascaris sp. Anisakidae gen. sp. Anisakis sp. Capillaria sp. Cucullanus carioca Dichelyne sp. Hysterothylacium sp. Nematoda gen. sp. Paracapillaria (P.) argentinensis Procamallanus sp. Raphidascaris sp. Terranova sp. Capillaria sp. Cucullanus carioca Dichelyne sp. Hysterothylacium sp. Paracapillaria (P.) argentinensis Procamallanus sp. Raphidascaris sp. Terranova sp.

Pseudopercis numida Miranda-Ribeiro

Pseudopercis semifasciata (Cuvier)

Family Polynemidae Polydactylus virginicus (Linnaeus) Procamallanus (S.) cruzi

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Family Pomatomidae Pomatomus saltatrix (Linnaeus) Anisakis sp. Contracaecum sp. Nematoda gen. sp. Philometra sp. Phocanema sp. Raphidascaris sp. Terranova sp.

Family Priacanthidae Priacanthus arenatus Cuvier Contracaecum sp. Nematoda gen. sp. Oncophora melanocephala Raphidascaris sp. Terranova sp.

Family Rachycentridae Rachycentron canadum (Linnaeus) Family Sciaenidae Cynoscion guatucupa (Cuvier) Cucullanus sp. Hysterothylacium sp. Terranova sp. Nematoda gen. sp. Contracaecum sp. Cucullanus sp. Hysterothylacium sp. Terranova trichiuri Terranova sp. Capillaria sp. Contracaecum sp. Hysterothylacium sp. Nematoda gen. sp. Procamallanus (S.) macaensis Procamallanus (S.) pereirai Procamallanus sp. Anisakidae gen. sp. Anisakis sp. Contracaecum sp. Cucullanus cassinensis Cucullanus pulcherrimus Cucullanus sp. Dichelyne (C.) amaruincai Dichelyne (C.) elongatus Dichelyne (D.) micropogonii Dichelyne (C.) rodriguesi Dichelyne (C.) sciaenidicola Iheringascaris inquies

Cynoscion leiarchus (Cuvier) Macrodon ancylodon (Bloch & Schneider)

Menticirrhus americanus (Linnaeus)

Micropogonias furnieri (Desmarest)

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Micropogonias furnieri (Desmarest)

Hysterothylacium sp. Nematoda gen. sp. Procamallanus (S.) pereirai Pseudocapillaria (P.) magalhaesi Raphidascaris sp. Terranova sp. Cucullanus cirratus Procamallanus (S.) macaensis Pseudocapillaria (P.) magalhaesi Anisakis sp. Procamallanus (S.) pereirai Raphidascaris sp. Raphidascaris (I.) vicentei Contracaecum sp. Type 2 Nematoda gen. sp. Procamallanus (S.) sp. Rhabdochona uruyeni Spinitectus rodolphiheringi Contracaecum sp. Cucullanus pulcherrimus Dichelyne (C.) elongatus Dichelyne (C.) sp. Philometra sp. Procamallanus (S.) macaensis Procamallanus (S.) pereirai Cucullanus rougetae Procamallanus (S.) macaensis Contracaecum sp. Contracaecum sp. Type 1 Eustrongylides sp. Hysterothylacium sp. Nematoda gen. sp. Thynnascaris sp. Terranova sp. Contracaecum sp. Type 2 Procamallanus (S.) pereirai Dichelyne (C.) sciaenidicola Hysterothylacium sp. Raphidascaris sp. Terranova sp. Anisakis physeteris Anisakis typica Oncophora melanocephala Anisakis sp. Contracaecum sp. Raphidascaris sp.

Micropogonias undulates (Linnaeus)

Nebris microps Cuvier

Pachyurus bonariensis Steindachner Pachyurus squamipennis Agassiz

Paralonchurus brasiliensis (Steindachner)

Plagioscion auratus (Castelnau) Plagioscion squamosissimus (Heckel)

Plagioscion ternetzi Boulenger Stellifer brasiliensis (Schultz) Umbrina canosai Berg

Family Scombridae Auxis thazard thazard (Lacpde)

Euthynnus alletteratus (Rafinesque)

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Katsuwonus pelamis (Linnaeus)

Anisakidae gen. sp. Anisakis sp. Anisakis sp. 1 Anisakis sp. 2 Philometra katsuwoni Contracaecum sp. Oncophora melanocephala Procamallanus sp. Raphidascaris sp. Terranova sp. Raphidascaris sp. Anisakis sp. Contracaecum sp. Hysterothylacium sp. Phocanema sp. Raphidascaris sp. Anisakis sp. Hysterothylacium sp. Raphidascaris sp. Contracaecum sp. Anasikis sp. Contracaecum sp. Hysterothylacium fortalezae Porrocaecum paivai Hysterothylacium fortalezae Oncophora melanocephala Oncophora melanocephala Anisakis typica Anisakis pegreffii Oncophora melanocephala

Sarda sarda (Bloch)

Scomber colias Gmelin Scomber japonicus Houttuyn

Scomber scombrus Linnaeus

Scomberomorus brasiliensis Collette, Russo & ZavalaCamin Scomberomorus cavalla (Cuvier)

Scomberomorus maculatus (Mitchill) Thunnus albacores (Bonnaterre) Thunnus atlanticus (Lesson) Thunnus thynnus (Linnaeus)

Family Serranidae Cephalopholis fulva (Linnaeus) Epinephelus guttatus (Linnaeus) Epinephelus sp. Mycteroperca sp. Family Sparidae Archosargus rhomboidalis (Linnaeus) Diplodus argenteus argenteus (Valenciennes) Pagrus pagrus (Linnaeus) Hysterothylacium sp. Nematoda gen. sp. Anisakidae gen. sp. Anisakis sp. Contracaecum sp. Contracaecum sp. Raphidascaris sp. Hysterothylacium eurycheilum Nematoda gen. sp. Nematoda gen. sp.

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Pagrus pagrus (Linnaeus)

Hysterothylacium sp. Nematoda gen. sp. Phocanema sp. Pseudoterranova sp. Raphidascaris sp. Terranova sp.

Family Sphyraenidae Sphyraena guachancho Cuvier Anisakis sp. Raphidascaris sp.

Family Stromateidae Peprilus paru (Linnaeus) Contracaecum sp. Hysterothylacium sp. Raphidascaris sp.

Family Trichiuridae Trichiurus lepturus Linnaeus Anisakis sp. Contracaecum sp. Hysterothylacium sp. Procamallanus sp. Pseudoterranova sp. Raphidascaris sp. Terranova sp.

Order Pleuronectiformes Family Achiridae Catathyridium jenynsii (Gnther) Family Cynoglossidae Symphurus tessellatus (Quoy & Gaimard) Family Paralichthyidae Citharichthys macrops Dresel Paralichthys isosceles Jordan Procamallanus (S.) halitrophus Anisakis simplex Anisakis sp. Contracaecum sp. Hysterothylacium sp. Raphidascaris sp. Terranova sp. Procamallanus (S.) halitrophus Procamallanus (S.) pereirai Nematoda gen. sp.

Syacium papillosum (Linnaeus) Family Dactylopteridae Dactylopterus volitans (Linnaeus)

Contracaecum sp.

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Dactylopterus volitans (Linnaeus)

Hysterothylacium sp. Procamallanus (S.) macaensis Raphidascaris sp.

Order Scorpaeniformes Family Triglidae Prionotus punctatus (Bloch) Anisakis sp. Cucullanus sp. Hysterothylacium sp. Procamallanus sp. Raphidascaris sp. Terranova sp.

Order Siluriformes Bagre Siluriformes fam. gen. sp. Procamallanus (S.) rarus Ascarididae gen. sp. Procamallanus sp. Procamallanus (S.) solani

Family Ariidae Bagre bagre (Linnaeus) Cathorops agassizii (Eigenmann & Eigenmann) Anisakis sp. Terranova trichiuri Falcaustra mirandafroesi Philometra fariaslimai Porrocaecum jardimfreire Contracaecum sp. Falcaustra mirandafroesi Hysterothylacium sp. Philometra fariaslimai Porrocaecum jardimfreire Falcaustra mirandafroesi Philometra fariaslimai Porrocaecum jardimfreire Cucullanus sp.

Genidens barbus (Lacpde)

Genidens genidens (Cuvier)

Ariidae gen. sp. Family Auchenipteridae Ageneiosus inermis (Linnaeus)

Contracaecum sp. Procamallanus (S.) inopinatus Nematoda gen. sp. Cucullanus (C.) pinnai pinnai Goezia sp. Cucullanus ageneiosus Procamallanus (S.) belenensis Procamallanus (S.) rarus Pseudoproleptus sp.

Ageneiosus militaris Valenciennes Ageneiosus ucayalensis Castelnau

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Auchenipterus nigripinnis (Boulenger)

Anisakidae gen. sp. Eustrongylides ignotus Eustrongylides sp. Cucullanus (C.) brevispiculus Johnstonmawsonia sp. Procamallanus (S.) sp. Rhabdochona acuminata Rhabdochona acuminata Cucullanus sp. Dichelyne (C.) sp. Goezia sp. Contracaeum sp. Cucullanus heliomartinsi Cucullanus sp. Hysterothylacium sp. Paracapillaria (P.) piscicola Procamallanus (P.) peraccuratus

Auchenipterus nuchalis (Spix & Agassiz) Auchenipterus osteomystax (Miranda Ribeiro)

Tatia neivai (Ihering) Trachelyopterus galeatus (Linnaeus)

Trachelyopterus striatulus (Steindachner)

Family Callichthyidae Callichthys callichthys (Linnaeus) Corydoras aeneus (Gill) Corydoras paleatus (Jenyns) Hoplosternum littorale (Hancock) Lepthoplosternum pectorale (Boulenger) Family Doradidae Acanthodoras spinosissimus (Eigenmann & Eigenmann) Franciscodoras marmoratus (Ltken) Rondonia rondoni Porrocaecum sp. Spinitectus rodolphiheringi Spinitectus sp. Rondonia rondonia Spinoxyuris oxydoras Cucullanus (C.) grandistomis Brevimulticaecum sp. Porrocaecum sp. Cucullanus (C.) pinnai pterodorasi Cucullanus sp. Cucullanus sp. 2 Hysterothylacium sp. Klossinemella iheringi Neoparaseuratum travassosi Paracamallanus amazonensis Procamallanus (S.) inopinatus Eustrongylides sp. Procamallanus (S.) pintoi Procamallanus (S.) pintoi Capillariinae gen. sp. Goezia sp. Eustrongylides sp.

Oxydoras kneri Bleeker Oxydoras niger (Valenciennes) Platydoras costatus (Linnaeus) Pterodoras granulosus (Valenciennes)

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Pterodoras granulosus (Valenciennes)

Raphidascaroides brasiliensis Rondonia rondoni Procamallanus (S.) rarus Procamallanus (S.) sp. Nematoda gen. sp. Cosmoxynemoides sp. Ichthyouris laterifilamenta Neoparaseuratum travassosi Procamallanus (S.) inopinatus Procamallanus sp. Rondonia rondoni

Rhinodoras dorbignyi (Kner)

Trachydoras paraguayensis (Eigenmann & Ward)

Family Hemiodontidae Hemiodus sp. Procamallanus (S.) sp. Procamallanus sp.

Family Heptapteridae Pimelodella gracilis (Valenciennes) Pimelodella lateristriga (Lichtenstein) Cucullanus (C.) pinnai pinnai Brasilnema pimelodellae Contracaecum sp. Cucullanus (C.) pimelodellae Procamallanus (S.) pimelodus Procamallanus (S.) rarus Rhabdochona acuminata Spinitectus rodolphiheringi Spinitectus yorkei Capillariinae gen. sp. Contracaecum sp. Type 2 Contracaecum sp. Cucullanus sp. Cucullanus (C.) pinnai pinnai Procamallanus (S.) hilarii

Rhamdia quelen (Quoy & Gaimard)

Family Loricariidae Ancistrus cirrhosus (Valenciennes) Hypostomus affinis (Steindachner) Hypostomus albopunctatus (Regan) Raphidascaris (S.) hypostomi Raphidascaris sp. Paracapillaria (Paracapillaria) piscicola Procamallanus (P.) annipetterae Raphidascaris (S.) hypostomi Raphidascaris sp. Procamallanus (P.) annipetterae Raphidascaris (S.) hypostomi Raphidascaris sp. Raphidascaris (S.) hypostomi

Hypostomus cochliodon Kner Hypostomus commersoni Valenciennes

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Hypostomus derbyi (Haseman)

Capillaria sp. Raphidascaris (S.) hypostomi Raphidascaris sp. Raphidascaris (S.) hypostomi Nematoda gen. sp. Ascarididae gen. sp. Cucullanus (C.) pinnai pinnai Contracaecum sp. Raphidascaris (S.) mahnerti Nematoda gen. sp. Raphidascaris (S.) mahnerti Raphidascaris (S.) mahnerti Hysterothylacium sp. Raphidascaris (S.) mahnerti Eustrongylides ignotus Ichthyouris brasiliensis Nematoda gen. sp. Raphidascaris (S.) mahnerti Parasynodontisia petterae

Hypostomus sp. Hypostomus regani (Ihering) Loricaria sp. Loricariichthys castaneus (Castelnau) Loricariichthys brunneus (Hancock) Loricariichthys platymetopon Isbrcker & Nijssen Loricariichthys labialis (Boulenger) Loricariichthys sp. Loricariidae gen. sp. Megalancistrus aculeatus (Peters) Proloricaria prolixa (Isbrcker & Nijssen) Pseudohemiodon laticeps (Regan) Rhinelepis aspera Spix & Agassiz Family Pimelodidae Bergiaria westermanni (Ltken) Bergiaria sp. Brachyplatystoma sp. Calophysus macropterus (Lichtenstein)

Procamallanus (S.) freitasi Contracaecum sp. Ascarididae gen. sp. Cucullanus sp. Alinema amazonicum Eustrongylides sp. Procamallanus (S.) inopinatus Procamallanus sp. Anisakidae gen. sp. Procamallanus (S.) sp. Rhabdochona sp. Brevimulticaecum sp. Contracaecum sp. 1 Contracaecum sp. 2 Contracaecum sp. Type 1 Contracaecum sp. Type 2 Contracaecum sp. Cucullanus (C.) zungaro Eustrongylides ignotus Goezia sp. Nematoda gen. sp. Hysterothylacium sp. Paracamallanus amazonensis Paracamallanus sp.

Conorynchus conirostris (Valenciennes)

Hemisorubim platyrhynchos (Valenciennes)

Hypophthalmus edentatus Spix & Agassiz

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Hypophthalmus edentatus Spix & Agassiz Iheringichthys labrosus (Ltken)

Nematoda gen. sp. Contracaecum sp. Procamallanus (S.) pimelodus Nematoda gen. sp. Alinema amazonicum Cucullanus (C.) pinnai pinnai Cucullanus (C.) zungaro Rondonia rondoni Cucullanus (C.) pinnai pinnai Nematoda gen. sp. Cucullanus (C.) pinnai pinnai Procamallanus (S.) rarus Cucullanus (C.) pinnai pinnai Cucullanus sp. Procamallanus (S.) inopinatus Procamallanus (S.) pimelodus Procamallanus sp. Cucullanus (C.) pinnai pinnai Klossinemella iheringi Nematoda gen. sp. Philometra baylisi Procamallanus (S.) pimelodus Procamallanus (S.) rarus Procamallanus sp. Rondonia rondoni Camallanidae gen. sp. Contracaecum sp. Cucullanus (C.) pinnai pinnai Cucullanus debacoi Cucullanus fabregasi Cucullanus patoi Cucullanus riograndensis Cucullanus sp. Cucullanus sp. B Dichelyne (D.) pimelodi Dichelyne sp. Eustrongylides sp. Goezia sp. Goezia spinulosa Klossinemella sp. Philometra sp. Procamallanus (S.) freitasi Procamallanus (S.) pimelodus Procamallanus (S.) rarus Procamallanus (S.) sp. Procamallanus (S.) sp. A Rhabdochona uruyeni Spinitectus rodolphiheringi Cucullanus (C.) pinnai pinnai

Luciopimelodus pati (Valenciennes)

Megalonema platanum (Gnther) Pimelodus albicans (Valenciennes) Pimelodus blochii Valenciennes

Synodontis clarias (Linnaeus)

Pimelodus fur (Ltken) Pimelodus maculatus Lacpde

Pimelodus ornatus Kner

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Pimelodus ortmanni Haseman

Contracaecum sp. Procamallanus (P.) peraccuratus Procamallanus (S.) pimelodus Contracaecum sp. Cucullanus (C.) pinnai pinnai Cucullanus sp. Dichelyne (D.) pimelodi Dichelyne sp. Procamallanus (S.) freitasi Procamallanus (S.) pimelodus Procamallanus (S.) sp. Alinema amazonicum Contracaecum sp. Eustrongylides sp. Eustrongylides ignotus Nematoda gen. sp. Anisakidae gen. sp. Brevimulticaecum sp. Capillariidae gen. sp. 2 Contracaecum sp. Contracaecum sp. 1 Contracaecum sp. 2 Contracaecum sp. Type 1 Contracaecum sp. Type 2 Cucullanus (C.) pinnai pinnai Cucullanus (C.) pseudoplatystomae Eustrongylides ignotus Eustrongylides tubifex Eustrongylides sp. Goezia brasiliensis Goezia spinulosa Philometridae gen. sp. Procamallanus (S.) sp. Nematoda gen. sp. [in the cestode Choanoscolex abscissus - Hyperparasitism] Anisakidae gen. sp. Contracaecum sp. Contracaecum sp. Type 1 Contracaecum sp. Type 2 Cucullanus (C.) pseudoplatystomae Cucullanus (C.) pinnai pinnai Cucullanus sp. Eustrongylides ignotus Eustrongylides sp. Amplicaecum sp. Ascaridoidea fam. gen. sp. Brevimulticaecum sp. Cucullanus (C.) pinnai pinnai Cucullanus sp. Nematoda gen. sp.

Pimelodus sp.

Pinirampus pirinampu (Spix & Agassiz)

Pseudoplatystoma corruscans (Spix & Agassiz)

Pseudoplatystoma fasciatum (Linnaeus)

Pseudoplatystoma sp.

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Pseudoplatystoma sp. Sorubim lima (Bloch & Schneider) Steindachneridion parahybae (Steindachner)

Eustrongylides sp. Brevimulticaecum sp. Cucullanus sp. Amplicaecum sp. Cucullanus (C.) pinnai pinnai Eustrongylides sp. Contracaecum sp. Cucullanus (C.) oswaldocruzi Cucullanus (C.) pauliceae Cucullanus (C.) pinnai pinnai Cucullanus schubarti Cucullanus sp. Cucullanus sp. 1 Cucullanus (C.) zungaro Procamallanus (S.) iheringi Rondonia rondoni Seuratoidea gen. sp.

Zungaro zungaro (Humboldt)

Family Pseudopimelodidae Pseudopimelodus mangurus (Valenciennes) Lophiosilurus alexandri Steindachner Family Trichomycteridae Pygidium brasiliensis Reinhardt Trichomycterus piurae (Eigenmann) Order Synbranchiformes Family Synbranchidae Synbranchus marmoratus Bloch Eustrongylides ignotus Eustrongylides sp. Nematoda gen. sp. Ascarididae gen. sp. Procamallanus (S.) pexatus Procamallanus (S.) hilarii Cucullanus (C.) pinnai pinnai Nematoda gen. sp.

Order Tetraodontiformes Family Balistidae Balistes capriscus Gmelin Contracaecum sp. Dichelyne sp. Raphidascaris sp. Terranova sp. Anisakis sp. Contracaecum sp. Dichelyne travassosi Dichelyne sp. Hysterothylacium sp.

Balistes vetula Linnaeus

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Family Monacanthidae Aluterus monoceros (Linnaeus) Anisakis sp. Contracaecum sp.

Family Tetraodontidae Sphoeroides testudineus (Linnaeus) Capillaria (N.) carioca Capillostrogyloides sentinosa Cucullanus dodsworthi

Class Chondrichthyes Order Carcharhiniformes Family Carcharhinidae Carcharhinus brachyurus (Gnther) Carcharhinus signatus (Poey) Galeocerdo cuvieri (Pron & Lesueur) Family Scyliorhinidae Scyliorhinus haeckelii (Miranda Ribeiro) Contracaecum sp. Raphidascaris sp. Contracaecum sp. Anisakis sp. Contracaecum sp. Nematoda gen. sp.

Family Sphyrnidae Sphyrna zygaena (Linnaeus) Contracaecum sp. Parascarophis sphyrnae

Family Triakidae Galeorhinus galeus (Linnaeus) Anisakis sp. Pseudoterranova sp. Contracaecum sp. Pseudoterranova sp. Raphidascaris sp. Anisakis sp. Contracaecum sp. Pseudoterranova sp. Contracaecum sp. Pseudoterranova sp. Terranova rochalimai

Mustelus canis (Mitchill)

Mustelus schmitti Springer Cao Panan Order Hexanchiformes Family Hexanchidae Heptranchias perlo (Bonnaterre)

Anisakis sp. Contracaecum sp.

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Hexanchus griseus (Bonnaterre)

Anisakis sp. Contracaecum sp.

Order Rajiformes Family Potamotrygonidae Paratrygon sp. Potamotrygon hystrix (Mller & Henle) Potamotrygon falkneri Castex & Maciel Eustrongylides ignotus Eustrongylides sp. Echinocephalus daileyi Brevimulticaecum sp. Brevimulticaecum sp. of Moravec et al. (1997) Cucullanus (C.) sp. Cucullanus sp. Echinocephalus sp. Spinitectus sp. Ascarididae gen. sp. Brevimulticaecum regoi Brevimulticaecum sp. Echinocephalus daileyi Procamallanus (P.) peraccuratus Procamallanus (S.) inopinatus Nematoda gen. sp.

Potamotrygon motoro (Mller & Henle)

Family Rajidae Atlantoraja castelnaui (Miranda Ribeiro) Dipturus trachydermus (Krefft & Stehmann) Procamallanus (S.) pereirai Raphidascaris sp. Anisakis sp. Contracaecum sp. Raphidascaris sp. Pseudanisakis sulamericana Pseudanisakis sulamericana

Psammobatis extent (Garman) Rioraja agassizii (Mller & Henle) Family Rhinobatidae Zapteryx brevirostris (Mller & Henle) Order Squaliformes Family Squalidae Squalus megalops (Macleay)

Proleptus acutus

Anisakis sp. Pseudoterranova sp.

Order Squatiniformes Family Squatinidae Squatina sp. Squatina squatina (Linnaeus) Anisakis sp. Contracaecum sp. Pulchrascaris caballeroi Terranova trichiuri

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Discussion
In recent years, parasites have been recognized as an important component of global biodiversity. Given the integral roles played by parasites in natural ecosystems, identifying hotspots of high parasite diversity, as well as areas of relatively low parasite diversity, is crucial for a complete understanding of the functioning of the biosphere (Luque & Poulin 2007). According Luque & Poulin (2007), nematode species represent 13% and 27% of the total number of host-parasite associations in Latin American and the Caribbean marine and freshwater fishes, respectively. In Brazil, a Megadiversity Country with the highest fish biodiversity of the planet, only 17.3% of the known fish species have parasitological records. Consequently, we are in agreement with the statement of Moravec (1998) that our current knowledge of nematodes parasitizing Neotropical fishes is too rudimentary to permit answers even to basic questions concerning their biodiversity, biology and ecology, never mind other issues relating to veterinary medicine and public health. The nematode fauna, therefore, remains an underestimated component of Neotropical fish parasite biodiversity. However, there are some general and incipient patterns in the distribution of nematode species in Brazilian fishes. For example, the number of nematode species parasitizing freshwater fishes is higher than occurs in marine and brackish fish species, and some fish groups, such as the Perciformes and Siluriformes, have a greater number of host-parasite associations. However, these differences may not reflect true biological patterns, as they may only be an echo of study effort and local research priorities. As only a small portion of the potential hosts have been examined for parasitic nematodes, further studies are needed to evaluate the possible implications of the faunal composition of this group. With regard to the distribution of nematode taxa, the greatest number of host-parasite associations involves ascaridoids, with approximately 51% of the records in this study. The significant presence of anisakid nematodes is of importance, because of possible consequences in terms of public health and fish marketing. Anisakids utilize fish as definitive, intermediate or paratenic hosts. The third larval stage of some of those occurring as adults in higher vertebrates can cause a disease referred to as anisakiasis, a typical food-borne parasitic zoonosis (Limbery & Cheah 2007), in man when raw or poorly cooked fish are consumed. Although human anisakiasis in Brazil remains unreported, the presence of anisakid larvae in several Brazilian commercial fish species at a high prevalence and abundance, plus an increase in the habit of eating raw fish, are indicative of a potential health problem in this country (Tavares & Luque 2006).

Acknowledgements
The authors are very grateful to the Fundao Carlos Chagas Filho de Amparo Pesquisa do Estado do Rio de Janeiro (FAPERJ) and the Instituto Oswaldo Cruz for financial support. Jos L. Luque was supported by a research fellowship from Conselho Nacional de Desenvolvimento Cientfico e Tecnolgico do Brasil (CNPq); Fabiano M. Vieira was supported by a Postdoctoral fellowship from CNPq (Conselho Nacional de Pesquisa e Desenvolvimento Tecnolgico, Brasil); and Julio C. Aguiar was supported by an MSc Fellowship from the Coordenao de Aperfeioamento do Pessoal de Ensino Superior (CAPES), Brazil.

References
Abdallah, V.D., Azevedo, R.K. & Luque, J.L. (2004) Metazorios parasitos dos lambaris Astyanax bimaculatus (Linnaeus, 1758), Astyanax parahybae Eigenmann, 1908 e Oligosarcus hepsetus (Cuvier, 1829) (Osteichthyes, Characidae), do Rio Guandu, Estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 13, 5763. Abdallah, V.D., Azevedo, R.K. & Luque, J.L. (2005) Ecologia da comunidade de metazorios parasitos do sair Cyphocharax gilbert (Quoy e Gaimard, 1824) (Characiformes, Curimatidae) do Rio Guandu, Estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 14, 154159. Abdallah, V.D., Azevedo, R.K. & Luque, J.L. (2006) Ecologia da comunidade de metazorios parasitos do tamboat Hoplosternum littorale (Hancock, 1828) (Siluriformes, Callichthyidae) do rio Guandu, Estado do Rio de Janeiro, Brasil. Acta Scientiarum, 28, 413419. Abdallah, V.D., Luque, J.L., Alves, D.R. & Paraguass, A.R. (2002) Aspectos quantitativos das infrapopulaes de metazorios parasitos da cavalinha, Scomber japonicus (Osteichthyes, Scombridae), do litoral do Estado do Rio de Janeiro. Revista da Universidade Rural, Srie Cincias da Vida, 22, 103107.

76 Zootaxa 3082 2011 Magnolia Press

LUQUE ET AL.

Adriano, E.A., Rgo, R.F., Santos, S.M.C. & Ceccarelli, P.S. (2005) Distribuio e prevalncia de parasitos protozoa e metazoa infectando piraputanga (Brycon hilarii) no pantanal mato-grossense, Brasil. Boletim Tcnico do CEPTA, 18, 2938. Albuquerque, M.C., Santos, M.D., Monteiro, C.M., Martins, A.N., Ederli, N.B. & Brasil-Sato, M.C. (2008) Helmintos endoparasitos de Pimelodus maculatus Lacpde, 1803 (Actinopterygii, Pimelodidae) de duas localidades (lagoa e calha do rio) do rio Guandu, Estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 17, 113119. Almeida, F.M., Morais, N.C.M. & Pereira Jr., J. (2008) Pseudocapillaria (Pseudocapillaria) magalhaesi em Micropogonias furnieri do litoral do Rio Grande do Sul, Brasil. Atlntica, 30, 6974. Silva, A.M.O., Tavares-Dias, M., Jernimo, G.T. & Martins, M.L. (2011) Parasite diversity in Oxydoras niger (Osteichthyes: Doradidae) from the basin of Solimes River, Amazonas state, Brazil, and the relationship between monogenoidean and condition factor. Brazilian Journal of Biology, 71, 791796 Alves, D.R. & Luque, J.L. (1999) Aspectos quantitativos das infrapopulaes de metazorios parasitos de indivduos jovens da corvina, Micropogonias furnieri (Osteichthyes, Sciaenidae), do litoral do Estado do Rio de Janeiro, Brasil. Contribuies Avulsas sobre a Histria Natural do Brasil, Srie Zoologia, 10, 14. Alves, D.R. & Luque, J.L. (2000) Metazorios parasitos de Micropogonias furnieri (Osteichthyes, Sciaenidae) do litoral do Estado do Rio de Janeiro, Brasil. Parasitologia al Da, 24, 1925. Alves, D.R. & Luque, J.L. (2001a) Aspectos quantitativos das infrapopulaes de metazorios parasitos da corvina, Micropogonias furnieri (Osteichthyes, Sciaenidae) do litoral do Estado do Rio de Janeiro. Parasitologia al Dia, 25, 3035. Alves, D.R. & Luque, J.L. (2001b) Community ecology of the metazoan parasites of the white croaker Micropogonias furnieri (Osteichthyes, Sciaenidae) from the coastal zone of the State of Rio de Janeiro, Brazil. Memrias do Instituto Oswaldo Cruz, 96, 45 153. Alves, D.R. & Luque, J.L. (2006) Ecologia das comunidades de metazorios parasitos de cinco espcies de escombrdeos (Perciformes, Scombridae) do litoral do estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 15, 167181. Alves, D.R., Luque, J.L. & Abdallah, V.D. (2003) Metazoan parasites of chubmackerel, Scomber japonicus Houttuyn (Osteichthyes, Scombridae), from the coastal zone of the State of Rio de Janeiro, Brazil. Revista Brasileira de Parasitologia Veterinria, 12, 164 170. Alves, D.R., Luque, J.L. & Paraguass, A.R. (2001a) Aspectos ecolgicos de las infrapoblaciones de Cucullanus genypteri (Nematoda, Cucullanidae) parsito del congro Genypterus brasiliensis del litoral del Estado de Rio de Janeiro, Brasil. Parasitologia al Dia, 25, 129131. Alves, D.R., Luque, J.L. & Paraguass, A.R. (2002a) Community ecology of the metazoan parasites of pink cuskeel, Genypterus brasiliensis (Osteichthyes, Ophidiidae), from the coastal zone of the State of Rio de Janeiro, Brazil. Memrias do Instituto Oswaldo Cruz, 97, 683689. Alves, D.R., Luque, J.L. & Paraguass, A.R. (2002b) Metazorios parasitos do congrorosa Genypterus brasiliensis Regan, 1903 (Osteichthyes, Ophidiidae) do litoral do Estado do Rio de Janeiro, Brasil. Revista Brasileira de Zoocincias, 4, 133142. Alves, D.R., Luque, J.L. Paraguass, A.R., Jorge, D.S & Vias, R.A. (2002c) Ecologia da comunidade de metazorios parasitos da abrtea Urophycis mystaceus Ribeiro, 1903 (Osteichthyes, Phycidae) do litoral do Estado do Rio de Janeiro. Revista Brasileira de Zoocincias, 4,1930. Alves, D.R., Luque, J.L., Paraguass, A.R. & Marques, F.A. (2000) Ocorrncia de Camallanus cotti (Nematoda, Camallanidae) parasitando o guppy Poecilia reticulata (Osteichthyes, Poeciliidae) no Brasil. Revista da Universidade Rural, Srie Cincias da Vida, 22, 7779. Alves, D.R., Paraguass, A.R. & Luque, J.L. (2004) Metazorios parasitos daabrtea, Urophycis brasiliensis (Kaup, 1858) (Osteichthyes, Phycidae) do litoral do Estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 13, 4955. Alves, D.R., Paraguass, A.R. & Luque, J.L. (2005) Community ecology of the metazoan parasites of the grey triggerfish, Balistes capriscus Gmelin, 1789 and queen triggerfish B.vetula Linnaeus, 1758 (Osteichthyes, Balistidae) from the State of Rio de Janeiro, Brazil. Revista Brasileira de Parasitologia Veterinria, 14, 7177. Anderson, R.C., Chabaud, A.G. & Willmott, S. (2009). Keys to the nematode parasites of vertebrates. CAB International, Wallingford, UK, 463pp. Andrade, S.M.S. & Malta, J.C.O. (2006) Parasite fauna monitoring of matrinxa Brycon amazonicus (Spix & Agassiz, 1829) raised in an intensive husbandry system in a stream channel in the state of Amazonas, Brazil. Brazilian Journal of Biology, 66, 1123-1132. Andrade, S.M.S., Malta, J.C.O. & Ferraz, E. (2001) Fauna parasitolgica de alevinos de matrinch Brycon cephalus (Gnther, 1869) coletados nos rios Negros e Solimes, na Amaznia Central. Acta Amazonica, 31, 263273. Araujo, C.S.O., Barros, M.C., Gomes, A.L.S. Varella, A.M.B., Viana, G.M., Silva, N.P., Fraga, E.V. & Andrade, S.M.S. (2009a) Parasitas de populaes naturais e artificiais de tucunar (Cichla spp.). Revista Brasileira de Parasitologia Veterinria, 18, 3438. Araujo, C.S O., Gomes, A.L., Tavares-Dias, M., Andrade, S.M.S., Belem-Costa, A., Borges, J.T., Queiroz, M.N. & Barbosa, M. (2009b) Parasitic infections in pirarucu fry, Arapaima gigas Schinz, 1822 (Arapaimatidae) kept in a semi-intensive fish farm in Central Amazon, Brazil. Veterinarski Arhiv, 79, 499507. Arci, A.D.N., Cohen, S.C. & Fernades, B.M.M. (2006) Helmintos em peixe de interesse economic do litoral do Rio de Janeiro. Revista Universidade Rural, Srie Ciencias da Vida, 26, 172-172. Azevedo, R.K., Abdallah, V.D. & Luque, J.L. (2006) Ecologia da comunidade de metazorios parasitos do acar Geophagus brasiliensis (Quoy e Gaimard, 1824) (Perciformes, Cichlidae) do rio Guandu, Estado do Rio de Janeiro, Brasil. Acta Scientiarum, 28, 403 411. Azevedo, R.K., Abdallah, V.D. & Luque, J.L. (2007a) Ecologia da comunidade de metazorios parasitos do apaiar Astronotus ocellatus (Cope, 1872) (Perciformes, Cichlidae) do rio Guandu, estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 16, 1520. Azevedo, R.K., Abdallah, V.D. & Luque, J.L. (2007b) Aspectos quantitativos da comunidade de metazorios parasitos do gordinho Peprilus paru (Linnaeus,1758) (Perciformes, Stromateidae), do litoral do estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 16, 1014. Azevedo, R.K., Abdallah, V.D. & Luque, J.L. (2010) Acanthocephala, Annelida, Arthropoda, Myxozoa, Nematoda and Platyhelminthes parasites of fishes from the Guandu river, Rio de Janeiro, Brazil. Check List, 6, 659667.

NEMATODA IN BRAZILIAN FISHES

Zootaxa 3082 2011 Magnolia Press

77

Azevedo, G.B., Madi, R.R. & Ueta, M.T. (2007c) Metazoans parasites of Astyanax altiparanae (Pisces, Characidae) at Rio das Pedras Farm, Campinas, SP, Brazil. Bioikos, 21, 8996. Barreto, A.L. (1918) Notas helmintolgicas III. Cucullanus pulcherrimus n. sp. de nematdeo. Brasil Medico, 18, 137138. Barreto, A.L. (1922) Reviso da familia Cucullanidae Barreto, 1916. Memrias do Instituto Oswaldo Cruz, 14, 6887. Barros, G.C. (1994) Larvas de anisakdeos de peixes economicamente importantes da costa do estado do Rio de Janeiro. Revista Brasileira de Medicina Veterinria, 16, 205208. Barros, G.C. & Amato, J.F.R. (1993) Larvas de anisakideos de peixeespada, Trichiurus lepturus L., da costa do Estado do Rio de Janeiro, Brasil. Revista Brasileira de Biologia, 53, 241245. Barros, L.A., Mateus, L.A.F., Braum, D.T. & Bonaldo, J. (2010) Aspectos ecolgicos de endoparasitos de piranha vermelha (Pygocentrus nattereri, Kner, 1860) proveniente do rio Cuiab. Arquivo Brasileiro de Medicina Veterinaria e Zootecnia, 62, 228231. Barros, L.A., Moraes-Filho, J. & Oliveira, R.L. (2006) Nematodes with zoonotical potential in fishes with economical importance from Cuiab river. Revista Brasileira de Cincia Veterinria, 13, 5557. Barros, L.A., Moraes-Filho, J. & Oliveira, R.L. (2007) Larvas de nematides de importncia zoontica encontradas em traras (Hoplias malabaricus Bloch, 1794) no municpio de Santo Antonio do Leverger, MT. Arquivo Brasileiro de Medicina Veterinaria e Zootecnia, 59, 533535. Barros, L.A., Oliveira, R.L., Moraes-Filho, J., Justino, C.H.S. & Mateus, L.A.F. (2009a) Analisys of the parasitism by Contracaecum sp. and Eustrongylides sp. in cacharas, Pseudoplatystoma fasciatum (Linnaeus, 1766) (Pisces, Pimelodidae) from Cuiab river, Mato Grosso, Brazil. Revista Brasileira de Cincia Veterinria, 16, 5861. Barros, L.A., Yamanaka, A.R., Silva, L.E., Vanzeler, M.L.A., Braum, D.T. & Bonaldo, J. (2009b) In vitro larvicidal activity of geraniol and citronella against Contracaecum sp. (Nematoda, Anisakidae). Brazilian Journal of Medical and Biological Research, 42, 918 920. Baylis, H.A. (1927) Some parasitic worms from Arapaima gigas (teleostean fish) with a description of Philometra senticosa n. sp. (Filarioidea). Parasitology, 19, 3547. Baylis, H.A. (1936) The nematode genus Rondonia Travassos, 1920. Annals and Magazine of Natural History, 10, 606610. Berg, C. (1895) Enumeracin sistemtica y sinonmica de los peces de las costas Argentina y Uruguaya. Anales Del Museo Nacional de Buenos Aires, 4, 1120. Bicudo, A.A., Tavares, L.E.R. & Luque, J.L. (2005a) Larvas de Anisakidae (Nematoda, Ascaridoidea) parasitas da cabrinha Prionotus punctatus (Bloch, 1793) (Osteichthyes, Triglidae) do litoral do estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 14, 109118. Bicudo, A.A., Tavares, L.E.R. & Luque, J.L. (2005b). Metazorios parasitos da cabrinha Prionotus punctatus (Bloch, 1793) (Osteichthyes, Triglidae) do litoral do estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 14, 2733. Brasil-Sato, M.C. (2003) Parasitos de peixes da bacia do rio So Francisco. In: Godinho, H.P, Godinho, A.L. (Eds), guas, peixes e pescadores do So Francisco das Minas Gerais. Belo Horizonte: PUC Minas, pp. 149165. Brasil-Sato, M.C. & Santos, M.D. (2003) Helmintos de Myleus micans (Ltken, 1875) (Characiformes, Serrasalminae) do Rio So Francisco, Brasil. Revista Brasileira de Parasitologia Veterinria, 12, 131134. Brasil-Sato, M.C. & Santos, M.D. (2005) Metazoan parasites of Conorhynchus conirostris (Valenciennes, 1840), an endemic siluriform fish of the So Francisco Basin, Brazil. Revista Brasileira de Parasitologia Veterinria, 14, 160166. Bruce, N.L., Adlard, R.D. & Cannon, L.R.G. (1994) Synoptic checklist of ascaridoid parasites (Nematoda) from fish hosts. Invertebrate Taxonomy, 8, 583674. Bhrnheim, U. (1976) Levantamento ecolgico dos helmintos parasitos de peixes do rio Mogi Guau na cachoeira de EMAS, Estado de So Paulo. Magister Thesis, Departamento de Zoologia, Instituto de Biocincias, Universidade de So Paulo, So Paulo, 224pp. Campos, C.M., Fonseca, V.E., Takemoto, R.M. & Moraes, F.R. (2008) Fauna parasitria de cachara Pseudoplatystoma fasciatum (Siluriforme, Pimelodidae) do rio Aquidauana, Pantanal Sul Mato-grossense, Brasil. Acta Scientiarum, Biological Sciences, 30, 9196. Campos, C.M., Fonseca, V.E., Takemoto, R.M. & Moraes, F.R. (2009b) Ecology of the parasitic endohelminth community of Pseudoplatystoma fasciatum (Linnaeus, 1776) (Siluriformes: Pimelodidae) from the Aquidauana River, Pantanal, State of Mato Grosso do Sul, Brazil. Brazilian Journal of Biology, 69, 9399. Campos, C.M., Moraes, F.R. & Pereira, G.T. (2007) Mtodo estimativo para amostragem quantitativa de Rondonia rondoni (Nematoda, Atractidae) parasito de peixes. Revista Brasileira de Parasitologia Veterinria, 16, 174176. Campos, C.M., Moraes, J.R.E. & Moraes, F.R. (2009c) Histopatologia do intestino de Pseudoplatystoma fasciatum (Osteicthyes, Pimelodidae) parasitados com cestides proteeocephaldeos e nematides. Boletim do Instiuto de Pesca, 35, 153158. Campos, C.M., Takemoto, R.M., Fonseca, V.E. & Moraes, F. R. (2009a) Ecology of the parasitic endohelminth community of Piaractus mesopotamicus (Holmberg, 1887) (Characiformes) from Aquidauana and Miranda Rivers, Pantanal, state of Mato Grosso do Sul, Brazil. Brazilian Journal of Biology, 69, 8792. Crdenas, M.Q. & Lanfredi, R. M. (2005) Further description of Procamallanus (S.) halitrophus comb. n. (Nematoda, Camallanidae) from flounder off the Brazilian coast by light and scanning electron microscopy. Journal of Parasitology, 91, 606613. Crdenas, M.Q., Moravec, F. & Kohn, A. (2009) First record of Philometra katsuwoni (Nematoda, Philometridae), a parasite of skipjack tuna Katsuwonus pelamis (Perciformes, Scombridae), off South American Atlantic Coast. Biota Neotropica, 9, 263266. Crdenas, M.Q., Souza, W. & Lanfredi, R.M. (2005) Ultrastructure of Procamallanus (S.) halitrophus (Nematoda, Camallanidae) parasite of flounder. Parasitology Research, 97, 478485. Carvalho, A.R., Tavares, L.E.R. & Luque, J.L. (2010) Variao sazonal dos metazorios parasitos de Geophagus brasiliensis (Perciformes, Cichlidae) no rio Guandu, Estado do Rio de Janeiro, Brasil. Acta Scientiarum, Biological Sciences, 32, 159167. Carvalho, S., Guidelli, G.M., Takemoto, R.M. & Pavanelli, G.C. (2003) Ecological aspects of endoparasite fauna of Acestrorhynchus lacustris (Ltken, 1875) (Characiformes, Acestrorhynchidae) on the Upper Paran River floodplain, Brazil. Acta Scientiarum, 25, 479483. Cavalcanti, E.T.S., Takemoto, R.M., Alves, L.C., & Chellappa, S. (2010) First record of endoparasite Philometra sp. (Nematoda: Philometridae) in lane snapper Lutjanus synagris from the coast of Rio Grande do Norte, Brazil. Marine Biodiversity Records, 3, 93. Ceschini, T.L., Takemoto, R.M., Yamada, F.H., Moreira, L.H.A. & Pavanelli, G.C. (2010a) Endoparasites of Steindachinerina brevi-

78 Zootaxa 3082 2011 Magnolia Press

LUQUE ET AL.

pinna (Eigenmann and Eigenmann, 1889), collected in the tributaries Corvo and Guairac of Paranapanema River, Paran State, Brazil. Acta Scientiarum, Biological Sciences, 32, 125130. Ceschini, T.L., Takemoto, R.M., Yamada, F.H., Moreira, L.H.A. & Pavanelli, G.C. (2010b) Metazoan parasite community of Steindachnerina brevipinna (Curimatidae) from Southern Brazil. Helminthologia, 47, 164-169. Chaves, N.D., & Luque, J.L. (1999) Ecology of metazoans parasites of Menticirrhus americanus (Osteichthyes, Sciaenidae), coast area from Rio de Janeiro State, Brazil. Revista Brasileira de Parasitologia Veterinria, 8, 137144. Conroy, G. & Conroy, D.A. (1984) Diseases and parasites detected in grey mullets (Mugilidae) from coastal waters of So Paulo state, Brazil. I. Adult silver mullet (Mugil curema Val., 1836). Estratto dalla Rivista Italiana di Piscicoltura e Ittiopatologia, 1, 1328. Cordeiro, A.S. & Luque, J.L. (2004) Community ecology of the metazoan parasites of Atlantic moon fish, Selene setapinnis (Osteichthyes, Carangidae) from the coastal zone of the State of Rio de Janeiro, Brazil. Brazilian Journal of Biology, 64, 399406. Cordeiro, A.S., & Luque, J.L. (2005a) Aspectos quantitativos dos metazorios parasitos do sargo-de-dente, Archosargus rhomboidalis (Linnaeus, 1758) (Osteichthyes, Sparidae), do litoral do Estado do Rio de Janeiro, Brasil. Revista Brasileira de Zoocincias, 7, 7 14. Cordeiro, A.S. & Luque, J.L. (2005b) Metazorios parasitos do cio Dactylopterus volitans (Linnaeus, 1758) (Osteichthyes, Dactylopteridae) do litoral do Estado do Rio de Janeiro, Brasil. Acta Scientiarum, 27, 119123. Costa, C.H.A. & Camargo, M. (2009) Procamallanus (Spirocamallanus) sp. (Camallanidae), an endoparasite of the digestive tract of Bivibranchia velox (Eigenmann & Myers, 1927) and B. fowleri (Steindachner, 1908), from the middle section of Rio Xingu, Para, Brazil. Uakari, 5, 97103. Costa, H.M.A, Guimares, M.P., Cabral, D.D. & Mundim, M.J.S. (1995) Scanning electron microscopic observations on Goezia spinulosa (Diesing, 1839) (Nematoda, Anisakidae) from Arapaima gigas (Cuvier, 1817). Memrias do Instituto Oswaldo Cruz, 90, 703705. Costa, H.M.A., Moreira, N.I.B. & Oliveira, C.L. (1991) Travassosnema gen. n. (Dracunculoidea, Guyanemidae) parasite of Acestrorhynchus lacustris Reinhardt, 1874 (Characidae) from Trs Marias reservoir, MG, Brazil. Memrias do Instituto Oswaldo Cruz, 86, 437439. Costa, S.C.G. (1960) Sbre um novo gnero da famlia Cobboldinidae Skrjabin, 1948 (Nematoda). Atas da Sociedade de Biologia do Rio de Janeiro, 4, 6263. Costa, S.C.G. (1961) Sbre a sistemtica da famlia Cobboldinidae Skrjabin, 1948 (Nematoda). Atas da Sociedade de Biologia do Rio de Janeiro, 5, 3032. Costa, S.C.G. (1962) Aspectos biolgicos de Rondonia rondoni Travassos, 1920 (Nematoda, Atractidae). Arquivos do Museu Nacional, 52, 7578. Costa, S.C.G. (1963) Rondonia rondoni Travassos, 1920 (Nematoda, Atractidae). Memrias do Instituto Oswaldo Cruz, 61, 7588. Costa, S.C.G., Motta, C.S. & Gomes, D. C. (1968) Reviso do gnero Klossinemella Costa, 1961 (Nematoda, Cobboldinidae), com descrio de uma nova subfamlia e de uma nova espcie. Memrias do Instituto Oswaldo Cruz, 66, 169179. Crisp, D.J. & Klein, V.L.M. (1973) Contribution to the knowledge of Philometra lateolabracis Yamaguti, 1935 (Nematoda, Filarioidea). Memrias do Instituto Oswaldo Cruz, 71, 481484. Deardorff, T.L., Brooks, D.R. & Thorson, T.B. (1981) A new species of Echinocephalus (Nematoda, Gnathostomidae) from Neotropical stingrays with comments on E. diazi. Journal of Parasitology, 67, 433439. Dias, F.J.E., So Clemente, S.C. & Knoff, M. (2010) Nematides anisaqudeos e cestoides Trypanorhyncha de importncia em sade pblica em Aluterus monoceros (Linnaeus, 1758) no Estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 19, 9497. Dias, F.J.E., So Clemente, S.C. Pinto, R.M. & Knoff, M. (2010) Anisakidae nematodes and Trypanorhyncha cestodes of hygienic importance infecting the king mackerel Scomberomosus cavalla (Osteichthyes: Scombridae) in Brazil. Veterinary Parasitology, 175, 351-355. Dias, P.G., Furuya, W.M., Pavanelli, G.C., Machado, M.H. & Takemoto, R.M. (2004) Carga parasitria de Rondonia rondoni Travassos, 1920, (Nematoda, Atractidae) e fator de condio do armado, Pterodoras granulosus Valenciennes, 1833 (Pisces, Doradidae). Acta Scientiarum, Biological Sciences, 26, 151156. Diesing, K.M. (1839) Neue Gattungen von Binnenwrmen nebst einem Nachtrge zur Monographie der Amphistomen. Annalen der Wiener Museums der Naturgeschichte, 2, 219242. Diesing, K.M. (1851) Systema helminthum. Vol. II. Wilhelmum Braumller, Vindobonae, 588 pp. Drasche, R. (1884a) Revision der in der Nematoden-Sammlung des k. k. zoologischen Hofcabinetes befindlichen Original-Exemplare Diesings und Molins. Verhandlungen der Kaiserlich- Kniglichen Zoologisch-Botanischen Gesselschaft in Wien, 33, 107118. Drasche, R. (1884b) Revision der in der Nematoden-Sammlung des k. k. zoologischen Hofcabinetes befindlichen Original-Exemplare Diesings und Molins. Spiruridea. Verhandlungen der Kaiserlich-Kniglichen Zoologisch-Botanischen Gesselschaft in Wien, 33, 193218. Eiras, J.C. & Rgo, A.A. (1988) Histopatologia da parasitose de peixes do rio Cuiab (Mato Grosso) por larvas de Eustrongylides sp. (Nematoda, Dioctophymidae). Revista Brasileira de Biologia, 48, 273280. Eiras, J.C. & Rgo, A.A. (1989) Histopatologia em peixes resultante de infeces parasitarias. Publicaes do Instituto de Zoologia Dr. Augusto Nobre, 208, 112. Eiras, J.C., Takemoto, R.M. & Pavanelli, G.C. (2010) Diversidade dos parasitas de peixes de gua doce do Brasil. Cliche Tec Editora, Maring, Brasil, 333 pp. Eschmeyer, W.N. (2010) Catalog of fishes. Electronic version. Available from: http://research.calacademy.org/ichthyology/catalog/ fishcatmain.asp (May 2010). Fbio, S.P. (1982) Sobre alguns nematoda parasitos de Hoplias malabaricus. Arquivos da Universidade Federal Rural do Rio de Janeiro, 5, 179186. Feijo, L.M.F., Rodrigues, H.O. & Rodrigues, S.S. (1979) Contribuio ao estudo da fauna helmintologica de sardinhas (Sardinella sp.) do litoral do Estado do Rio de Janeiro. Atas da Sociedade de Biologia do Rio de Janeiro, 20, 2328. Felizardo, N.N., Knoff, M., Pinto, R.M. & Gomes, D.C. (2009a) Larval anisakid nematodes of the flounder, Paralichthys isosceles Jordan, 1890 (Pisces, Teleostei) from Brazil. Neotropical Helminthology, 3, 5764.

NEMATODA IN BRAZILIAN FISHES

Zootaxa 3082 2011 Magnolia Press

79

Felizardo, N.N., Menezes, R.C., Tortelly, R., Knoff, M., Pinto, R.M. & Gomes, D.C. (2009b) Larvae of Hysterothylacium sp. (Nematoda, Anisakidae) in the sole fish Paralichthys isosceles Jordan, 1890 (Pisces, Teleostei) from the littoral of the state of Rio de Janeiro, Brazil. Veterinary Parasitology, 166, 175177. Feltran, R.B., Jnior, O.M., Pinese, J.F. & Takemoto, R.M. (2004) Prevalncia, abundncia, intensidade e amplitude de infeco de nematides intestinais em Leporinus friderici (Bloch, 1794) e L. obtusidens (Valenciennes, 1836) (Pisces, Anostomidae), na represa de Nova Ponte (Perdizes, MG). Revista Brasileira de Zoocincias, 6, 169179. Fernandes, B.M.M., Pinto, R.M. & Cohen, S.C. (2002) Report on two species of Digenea from marine fishes in Brazil. Brazilian Journal of Biology, 62, 459462. Fernandes, G.Q., Pereira, M.A.V.C., Mattos Jr., D.G. & Souza, G. (2006) Levantamento de parasitos em infrapopulao de Brycon insignis Steindachner, 1876 (Pisces, Characidae), criada na regio norte fluminense, Rio de Janeiro, Brasil. Cincia Animal Brasileira, 7, 309313. Fernandes, M.T., Artigas, P.T. & Campos, M.S. (1982) Spinitectus asperus Travassos, Artigas & Pereira, 1928 (Nematoda, Spiruridae). Redescrio da fmea e descrio do macho. Revista de Cincias Biomedicas, 3,7985. Fernandes, M.T., Campos, M.S. & Artigas, P.T. (1983) Travnema araujoi n. sp. (Nematoda, Pharyngodonidae) parasito de Curimatus gilberti Quoy & Gaimard (Pisces, Characinidae). Revista da Faculdade de Medicina Veterinaria e Zootecnia da Universidade de So Paulo, 20, 3541. Ferraris Jr., C.J. (2007) Checklist of catfishes, recent and fossil (Osteichthyes, Siluriformes), and catalogue of siluriform primary types. Zootaxa, 1418, 1628. Ferraz, E. & Thatcher, V.E. (1988) Bacudacnitis grandistomis gen. et. sp. n. (Nematoda, Cucullanidae) parasita intestinal de um bagre, Pseudodoras niger (Valenciennes) da Amazonia brasileira. Amazoniana, 10, 249253. Ferraz, E. & Thatcher, V.E. (1990) Camallanus acaudatus sp. n. (Nematoda, Camallanidae) e uma descrio do macho de Camallanus tridentatus (Drasche, 1884), parasitas de peixes da Amaznia Brasileira. Amazoniana, 11, 135145. Ferraz, E. & Thatcher, V.E. (1992) Paracamallanus amazonensis sp. n. (Nematoda, Camallanidae) um parasita de bagre, Hypophthalmus edentatus (Pisces, Hypophthalmidae) da Amazonia Brasileira. Amazoniana, 12, 16. Fischer, C., Malta, J.C.O. & Varella, A.M.B. (2003) A fauna de parasitas do tambaqui, Colossoma macropomum (Cuvier, 1818) (Characiformes, Characidae) do medio rio Solimes, Estado do Amazonas (AM) e do baixo rio Amazonas, Estado do Par (PA), e seu potencial como indicadores biologicos. Acta Amazonica, 33, 651662. Fortes, E. (1981) Descrio de quatro espcies novas de helmintos de bagres (Pisces, Bagridae) do esturio do Guaba, Porto Alegre, RS, Brasil. Arquivos da Faculdade de Veterinaria, 9, 6978. Fortes, E. & Hoffmann, R.P. (1995) Levantamento da fauna parasitria de peixes do lago Guaba, Porto Alegre, Rio Grande do Sul, Brasil. Revista Brasileira de Medicina Veterinaria, 17, 107111. Fortes, E., Hoffmann, R.P. & Querol, M.V.M. (1999) Presena de Raphidascaris (Sprentascaris) mahnerti (Petter et Cassone, 1984) Nematoda, Anisakidae em Loricariichthys platymetopon (cascudo viola) da bacia do Rio Uruguai medio, Uruguaiana, Rio Grande do Sul, Brasil. Revista Brasiliera de Medicina Veterinaria, 21, 1314. Fortes, E., Hoffmann, R.P. & Sarmento,T.M. (1992) Descrio de Cucullanus patoi sp. n. (Nematoda, Cucullanidae) de pintado Pimelodus maculatus Lacpde, 1803 (Pisces), do Lago do Guaiba, RS, Brasil. Cincia Rural, 22, 325328. Fortes, E., Hoffmann, R.P. & Sarmento, T.M. (1993a) Nova especies de nematdeo Cucullanus, C. riograndensis sp. n. de Pimelodus maculatus Lacpde, 1803 do Lago do Guaiba, Rio Grande do Sul, Brasil. Revista Brasileira de Medicina Veterinaria, 15, 7982. Fortes, E., Hoffman, R.P. & Sarmento, T.M. (1993b) Cucullanus fabregasi sp. n. nematoda parasita do intestino do peixe Pimelodus maculatus (Lacpde, 1803) do Rio Guaiba, Porto Alegre, Rio Grande do Sul, Brasil. Arquivos da Faculdade de Veterinaria, 21, 7176. Freitas, J.F.T. & Lent, H. (1935) Capilariinae de animaes de sangue frio (Nematoda, Trichuroidea). Memrias do Instituto Oswaldo Cruz, 30, 241284. Freitas, J.F.T. & Lent, H. (1946) Infestao de apaiars Astronotus ocellatus (Agassiz) pelo nematdeo Goezia spinulosa (Diesing, 1839). Revista Brasileira de Biologia, 6, 215222. Froese, R. & Pauly, D. (Eds) (2010) FishBase. World Wide Web electronic publication.Available from: www.fishbase.org, version (May 2010). Fujimoto, R.Y., Santana, C.A., Carvalho, W.L.C., Diniz, D.G. Barros, Z.M.N., Varella, J.E.A. & Guimares, M.D.F. (2009) Hematologia e parasitas metazorios de camurim (Centropomus undecimalis Bloch, 1792) na regio bragantina, Bragana-Par. Boletim do Instituto de Pesca de So Paulo, 35, 441450. Gallego-Berenger, J. (1947) Revisin de La famlia Atractidae Travassos, 1920 com descripcin de nuevas espcies. Revista Ibrica de Parasitogia, 7, 390. Gibbons, L.M. (2010) Keys to nematode parasites of vertebrates. Supplementary Volume. CAB International, Wallingford, 416p. Gibson, D.I., Bray, R.A. & Harris, E.A. (2005) Host Parasite Database of the Natural History Museum, London. Available from: http:/ /www.nhm.ac.uk/researchcuration/research/projects/hostparasites/database/index.jsp (May 2010). Giese, E.G., Furtado, A.P., Lanfredi, R.M. & Santos, J.N. (2010) A new cucullanid species (Nematoda) from Ageneiosus ucayalensis Castelnau, 1855 (Pisces, Auchenipteridae) from Par, Brazil. Journal of Parasitology, 96, 389394. Giese, E.G., Santos J.N. & Lanfredi R.M. (2009) A new species of Camallanidae from Ageneiosus ucayalensis (Pisces, Siluriformes) from Para State, Brazil. Journal of Parasitology, 95, 407412. Gomes, D.C. & Kohn, A. (1970) Sobre a subfamlia Ancyracanthinae Yorke et Maplestone, 1926 (Nematoda, Spiruroidea). Atas da Sociedade de Biologia do Rio de Janeiro, 13, 8388. Graa, R.J. & Machado, M.H. (2007) Ocorrncia e aspectos ecolgicos de metazorios parasitos de peixes do Lago do Parque do Ing, Maring, Estado do Paran. Acta Scientiarum, Biological Sciences, 29, 321326. Guidelli, G.M., Isaac, A., Takemoto, R.M. & Pavanelli, G.C. (2003) Endoparasite infracommunities of Hemisorubim platyrhynchos (Valenciennes, 1840) (Pisces, Pimelodidae) of the Baa River, upper Paran River floodplain, Brazil: Specific composition and ecological aspects. Brazilian Journal of Biology, 63, 261268. Guidelli, G.M., Tavechio, W.G., Takemoto, R.M. & Pavanelli, G.C. (2006) Fauna parasitria de Leporinus lacustris e Leporinus friderici (Characiformes, Anostomidae) da plancie de inundao do alto rio Paran, Brasil. Acta Scientiarum, Biological Sciences, 28,

80 Zootaxa 3082 2011 Magnolia Press

LUQUE ET AL.

281290. Guimares, J.F. (1975) Contribuio ao estudo da fauna helmintolgica do estado da Bahia. Boletim do Instituto Biolgico da Bahia, 14, 4452. Guimares, J.F. & Cristofaro, R. (1974) Contribuio ao estudo da fauna helmintolgica de peixes do Estado da Bahia. Atas da Sociedade de Biologia do Rio de Janeiro, 17, 8185. Guimares, J.F., Cristofaro, R., Rodrigues, H.O. (1976) Alguns nematdeos de peixes de Salvador, Bahia. Atas da Sociedade de Biologia do Rio de Janeiro, 18, 2125. Hodda, M. (2007) Phylum Nematoda. Zootaxa, 1668, 265293. Inglis, W.G. & Ogden, C.C. (1964) Miscellanea Nematodologica. IV. The male of Alinema alli Rasheed, 1963. Annals and Magazine of Natural History, 81, 523525. Iiguez, A.M., Santos. C.P. & Vicente, A.C.P. (2009) Genetic characterization of Anisakis typica and Anisakis physeteris from marine mammals and fish from the Atlantic Ocean off Brazil. Veterinary Parasitology, 165, 350356. Isaac, A., Guidelli, G.M., Frana, J.G. & Pavanelli, G.C. (2004) Composio e estrutura das infracomunidades endoparasitrias de Gymnotus spp. (Pisces, Gymnotidae) do rio Baa, Mato Grosso do Sul, Brasil. Acta Scientiarum, Biological Sciences, 26, 453462. Ito, K.F., Moreira, S.T., Takemoto, R.M. & Pavanelli, G.C. (2005) Ecological aspects of the Procamallanus (S.) pintoi parasite of Corydoras paleatus (Jenyns, 1842) (Siluriformes, Callichthydae) in reservoirs of the State of Paran, Brazil. Acta Scientiarum, Biological Sciences, 27, 239242. Klein, V.L.M. (1973) Helmintos parasitos das espcies Scomberomus cavalla (Cuvier) e Scomberomus maculatus (Mitchill) do litoral Cearense. Contracaecum fortalezae sp. n. (Nematoda, Ascaroidea). Memrias do Instituto Oswaldo Cruz, 71, 199202. Kloss, G.R. (1966) Helmintos parasitos de espcies simptricas de Astyanax (Pisces, Characidae). Papis Avulsos do Departamento de Zoologia de So Paulo, 18, 189219. Knoff, M., Luque, J.L. & Amato, J.F.R. (1997) Community ecology of the metazoan parasites of grey mullets, Mugil platanus (Osteichthyes, Mugilidae) from the littoral of the State of Rio de Janeiro, Brazil. Revista Brasileira de Biologia, 57, 441454. Knoff, M., So Clemente, S.C., Fonseca, M.C.G., Andrada, C.G., Padovani, R.E.S. & Gomes, D.C. (2007) Anisakidae parasitos de congro-rosa, Genypterus brasiliensis Regan, 1903 comercializados no estado do Rio de Janeiro, Brasil de interesse na sade pblica. Parasitologa Latinoamericana, 62, 127133. Knoff, M., So Clemente, S.C., Lima, F.C. & Ferreira, M.S. (2003a) Helmintos importantes na industrializao e comercializao de peixes telesteos utilizados para consumo. Revista Higiene Alimentar, 17, 6565. Knoff, M., So Clemente, S.C., Lima, F.C., Padovani, R.E.S., Castro, R.R.T. & Gomes, D.C. (2003b) Helmintos de importncia sanitria presentes no congro-rosa Genypterus brasiliensis Regan, 1903, comercializados no Estado do Rio de Janeiro. Revista Higiene Alimentar, 17, 8990. Knoff, M., So Clemente, S.C., Pinto, R.M. & Gomes, D.C. (2001) Nematodes of elasmobranch fishes from southern coast of Brazil. Memrias do Instituto Oswaldo Cruz, 96, 17. Kohn, A. & Fernandes, B.M.M. (1987) Estudo comparativo dos helmintos parasitos de peixes do Rio Mogi Guassu, coletados nas excurses realizadas entre 1927 e 1985. Memrias do Instituto Oswaldo Cruz, 82, 483500. Kohn, A. & Fernandes, B.M.M. (1988a) Helminth parasites of fishes from hydroelectric power station of Eletrosul (Brazil). I. Procamallanus petterae n. sp. and S. pintoi n. sp. (Nematoda, Camallanidae) from the reservoir of Salto Osrio. Memrias do Instituto Oswaldo Cruz, 83, 293298. Kohn, A. & Fernandes, B.M.M. (1988b) Procamallanus annipetterae nom. n. for Procamallanus petterae Kohn & Fernandes, 1988 preocupied by Procamallanus (Procamallanus) petterae Moravec & Sey, 1988. Memrias do Instituto Oswaldo Cruz, 83, 535. Kohn, A., Fernandes, B.M.M., Baptista-Farias, M.F.D., Cohen, S.C., Fernandez, D.R. & Canzi, C. (2003) Helmintos em peixes do Reservatrio de Itaipu e reas de influncia. Revista Brasileira de Medicina Veterinria, 25, 148153. Kohn, A., Fernandes, B.M.M., Baptista-Farias, M.F.D., Cohen, S.C., Santos, A.L., Pamplona-Basilio, M.C., Vieira, J.A.F. & Feitosa, V.A. (2004) Prevalncia de helmintos parasitos dos peixes do Aude Pereira de Miranda e viveiros do DNOCS (Pentecoste, Cear, Brasil). Revista Brasileira de Cincia Veterinria, 11, 5557. Kohn, A., Fernandes, B.M.M., Macedo, B. & Abramson, B. (1985) Helminths parasites of freshwater fishes from Pirassununga, SP, Brazil. Memrias do Instituto Oswaldo Cruz, 80, 327336. Kohn, A., Fernades, B.M.M., Pipolo, H.V. & Godoy, M.P. (1988) Helmintos parasitos de peixes das usinas hidreltricas Eletrosul (Brasil). II. Reservatrios de Salto Osrio e de Salto Santiago, bacia do rio Iguau. Memrias do Instituto Oswaldo Cruz, 83, 299303. Kohn, A., Fernades, B.M.M., Pipolo, H.V. & Godoy, M.P. (1989) List of helminth parasites of fishes from the Passo Fundo reservoir, Uruaguay River basin, Brazil. Memrias do Instituto Oswaldo Cruz, 84, 427428. Kohn, A., Gomes, D.C. & Motta, C.S. (1968) Nota prvia sobre um novo gnero de Ancyracanthinae Yorke & Maplestone, 1926 (Nematoda). Atas da Sociedade de Biologia do Rio de Janeiro, 12, 27. Kohn, A. & Macedo, B. (1984) First record of Aspiculuris tetraptera (Nitzsch, 1821) (Nematoda, Oxyuroidea) and Dollfusentis chandleri (Golvan, 1969) (Acanthocephala, Illiosentidae) in Haemulon sciurus (Shaw, 1803) (Pisces, Pomadasyidae). Annales de Parasitologie Humaine et Compare, 59, 477482. Lacerda, A.C.F., Santin, M., Takemoto, R.M., Pavanelli, G.C., Bialetzki, A. & Tavernari, F.C. (2009) Helminths parasitizing larva fish from Pantanal, Brazil. Journal of Helminthology, 83, 5155. Lacerda, A.C.F., Takemoto, R.M. & Pavanelli, G.C. (2008) Digenea, Nematoda, Cestoda, and Acanthocephala, parasites in Potamotrygonidae (Chondrichthyes) from the upper Paran River floodplain, states of Paran and Mato Grosso do Sul, Brazil. Check List, 4, 115122. Lacerda, A.C.F., Takemoto, R.M. & Pavanelli, G.C. (2009) Ecology of endoparasites of the fluvial stingray Potamotrygon falkneri (Chondrichthyes, Potamotrygonidae) from the upper Paran River floodplain, Brazil. Brazilian Journal of Biology, 69, 297303. Lamothe-Argumedo, R., Garca-Prieto, L., Osorio-Sarabia, D. & Prez-Ponce de Len, G. (1996) Catlogo de la Coleccin Nacional de Helmintos depositada en el Instituto de Biologia. Universidad Nacional Autonoma de Mxico, Publicacin Especial Instituto de BiologiaCONABIO, Mxico D. F., 211 pp. Leo, E.L.M., Leite, R.G., Chaves, P.T.C. & Ferraz, E. (1991) Aspectos de reproduo, alimentao e parasitofauna de uma espcie de piranha, Serrasalmus altuvei Ramirez, 1965 (Pisces, Serrasalmidae) do Baixo Rio Negro. Revista Brasileira de Biologia, 51, 545

NEMATODA IN BRAZILIAN FISHES

Zootaxa 3082 2011 Magnolia Press

81

553. Lent, H. & Freitas, J.F.T. (1937) Alguns helminthos da colleco de Pedro Severiano de Magalhes. Memrias do Instituto Oswaldo Cruz, 31, 305309. Lima, M.M., Teixeira, W.C., Ramos, R.A.N., Lima, A.M.A., Alves, L.C. & Faustino, M.A.G. (2008) Ocorrncia de ovos de Capillaria sp. em fil de peixe Saramunete (Pseudupeneus maculatus) comercializado na Regio Metropolitana de Recife-PE/Brasil. Medicina Veterinria, Recife, 2, 3537. Lizama, M.A.P., Takemoto, R.M. & Pavanelli, G.C. (2005) Influence of host sex and age on infracommunities of metazoan parasites of Prochilodus lineatus (Valenciennes, 1836) (Prochilodontidae) of the Upper Paran River floodplain, Brazil. Parasite, 12, 299 304. Lordello, L.G.E. & Monteiro, F.L. (1959) Larvas de nematdeos do gnero Eustrongylides parasitando pintado do rio Piracicaba (Diotophymidea). Revista Agrcola, 34, 3740. Luque, J.L. & Alves, D.R. (2001) Ecologia das comunidades de metazorios parasitos do xaru, Caranx hippos (Linaneus) e do xerelete, Caranx latus Agassiz (Osteichthyes, Carangidae) do litoral do Estado do Rio de Janeiro, Brasil. Revista Brasileira de Zoologia, 18, 399410. Luque, J.L., Alves, D.R. & Ribeiro, R.S. (2003) Community ecology of the metazoan parasites of banded croaker, Paralonchurus brasiliensis (Osteichthyes, Sciaenidae), from thecoastal zone of the State of Rio de Janeiro, Brazil. Acta Scientiarum, 25, 273 278. Luque, J.L., Alves, D.R. & Sabas, C.S.S. (2000a) Metazorios parasitos do xaru Caranx hippos (Linnaeus, 1766) e do xerelete Caranx latus Agassiz, 1831 (Osteichthyes, Carangidae) do litoral do Estado do Rio de Janeiro, Brasil. Contribuies Avulsas Sobre a Histria Natural do Brasil, 25, 117. Luque, J.L. & Chaves, N.D. (1999) Ecologia da comunidade de metazorios parasitos de Pomatomus saltator (Teleostei, Pomatomidae) do litoral do Estado do Rio de Janeiro. Revista Brasileira de Zoologia, 16, 711723. Luque, J.L., Cordeiro, A.S. & Oliva, M. E. (2010) Metazoan parasites as biological tags for stock discrimination of whitemouth croaker Micropogonias furnieri. Journal of Fish Biology, 76, 591600. Luque, J.L., Felizardo, N.N. & Tavares L.E.R. (2008) Community ecology of the metazoan parasites of namorado sandperches, Pseudopercis numida and P. semifasciata (Perciformes, Pinguipedidae), from the coastal zone of the state of Rio de Janeiro, Brazil. Brazilian Journal of Biology, 68, 269278. Luque, J.L., Porrozzi, F. & Alves, D.R (2002) Community ecology of the metazoan parasites of Argentine goatfish Mullus argentineus (Osteichthyes, Mullidae) from the coastal zone of the State of Rio de Janeiro, Brazil. Revista Brasileira de Parasitologia Veterinria, 11, 3338. Luque, J.L. & Poulin, R. (2004) Use of fish as intermediate hosts by helminth parasites: a comparative analysis. Acta Parasitologica, 49, 353361. Luque, J.L. & Poulin, R. (2007) Metazoan parasite species richness in Neotropical fishes: hotspots and the geography of biodiversity. Parasitology, 134, 865878. Luque, J.L., Vias, R.A., Paraguass, A.R. & Alves, D.R. (2000b) Metazorios parasitos das sardinhas Sardinella brasiliensis e Harengula clupeola (Osteichthyes, Clupeidae) do litoraldo Estado do Rio de Janeiro, Brasil. Revista da Universidade Rural, Srie Cincias da Vida, 22, 7176. Machado, M.H., Pavanelli, G.C. & Takemoto, R.M. (1994) Influence of the hosts sex and size in endoparasitic infrapopulations of Pseudoplatystoma corruscans and Schizodon borelli (Osteichthyes) of the high River Paran. Revista Brasileira de Parasitologia Veterinria, 3, 143148. Machado, M.H., Pavanelli, G.C. & Takemoto, R.M. (1995) Influence of the type of environment and the hydrological level variation in the endoparasitic infrapopulations of Pseudoplatystoma corruscans and Schizodon borelli (Osteichthyes) of the high River Paran. Revista Brasileira de Zoologia, 12, 961976. Machado, M.H., Pavanelli, G.C. & Takemoto, R.M. (1996) Structure and diversity of endoparasitic infracomunities and the trophic level of Pseudoplatystoma corruscans and Schizodon borelli (Osteichthyes) of the high River Paran. Memrias do Instituto Oswaldo Cruz, 91, 441448. Machado, P.M., Almeida, S.C., Pavanelli, G.C. & Takemoto, R.M. (2000) Ecological aspects of endohelminths parasitizing Cichla monoculus Spix, 1831 (Perciformes, Cichlidae) in the Paran River near Porto Rico, State of Paran, Brazil. Comparative Parasitology, 67, 210217. Madi, R.R. & Silva, M.S.R. (2005) Contracaecum Railliet & Henry, 1912 (Nematoda, Anisakidae): o parasitismo relacionado biologia de trs espcies de peixes piscvoros no reservatrio do Jaguari, SP. Revista Brasileira de Zoocincias, 7, 1524. Marques, M.C., So-Clemente, S.C., Barros, G.C. & Lucena, F.P. (1995) Effect of cold (chilling and freezing) on survival of anisakid nematode larvae in Trichiurus lepturus (L.). Higiene Alimentar, 9, 2328. Martins, M.L., Fenerick Jr., J., Nomura, D.T. & Braga, F.M.S. (2003) Prevalence, intensity and seasonality of Ichthyouris voltagrandensis (Nematoda: Pharyngodonidae) in Myleus tiete (Osteichthyes: Characidae) from Volta Grande reservoir, Minas Gerais state, Brazil. Boletim do Instituto de Pesca, 29, 3946. Martins, M.L., Fujimoto, R.Y., Moraes, F.R., Andrade, P.M., Nascimento, A.A. & Malheiros, E.B. (2000) Description and prevalence of Thynnascaris sp. larvae Dollfus, 1933 (Nematoda, Anisakidae) in Plagioscion squamosissimus Heckel, 1840 from Volta Grande Reservoir, state of Minas Gerais, Brazil. Revista Brasileira de Biologia, 60, 519526. Martins, M.L., Garcia F., Piazza, R.S. & Ghiraldelli, L. (2007) Camallanus maculatus n. sp. (Nematoda, Camallanidae) in an ornamental fish Xiphophorus maculatus (Osteichthyes, Poeciliidae) cultivated in So Paulo State, Brazil. Arquivo Brasileiro de Medicina Veterinaria e Zootecnia, 59, 12241230. Martins, M.L. & Onaka, E.M. (2004) Larvae of Porrocaecum sp. (Nematoda, Ascarididae) in the swim bladder of cultured Piaractus mesopotamicus (Osteichthyes, Characidae) in Brazil. Boletim do Instituto de Pesca de So Paulo, 30, 5761. Martins, M.L., Onaka, E.M. & Fenerick Jr., J. (2005) Larval Contracaecum sp. (Nematoda, Anisakidae) in Hoplias malabaricus and Hoplerythrinus unitaeniatus (Osteichthyes, Erythrinidae) of economic importance in occidental marshlands of Maranho, Brazil. Veterinary Parasitology, 127, 5159. Martins, M.L., Santos, R.S., Marengoni, N.G., Takahashi, H.K. & Onaka, E.M. (2009) Seasonality of Eustrongylides sp. (Nematoda,

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LUQUE ET AL.

Dioctophymatidae) larvae in fishes from Paran River, south-western Brazil. Boletim do Instituto de Pesca, 35, 2937. Martins, M.L., Tavares-Dias, M., Fujimoto, R.Y., Onaka, E.M. & Nomura, D.T. (2004) Haematological alterations of Leporinus macrocephalus (Osteichthyes, Anostomidae) naturally infected by Goezia leporini (Nematoda, Anisakidae) in fish pond. Arquivos Brasileiros de Medicina Veterinria e Zootecnia, 56, 640646. Martins, M.L. & Urbinati, E.C. (1993) Rondonia rondoni Travassos, 1919 (Nematoda, Atractidae) parasite of Piaractus mesopotamicus Holmberg, 1887 (Osteichthyes, Characidae), in Brazil. Ars Veterniria, 9, 7581. Martins, M.L. & Yoshitoshi, E.R. (2003) A new nematode species Goezia leporini n. sp. (Anisakidae) from cultured freshwater fish Leporinus macrocephalus (Anostomidae) in Brazil. Brazilian Journal of Biology, 63, 497506. Martins, M.L., Yoshitoshi, E.R. & Umekita, H. (2001) Ichthyouris voltagrandensis n. sp. (Nematoda, Pharyngodonidae) from Myleus tiete Eigenmann & Norris, 1900 (Osteichthyes, Characidae) in the Volta Grande Reservoir, MG, Brazil. Revista Brasileira de Biologia, 61, 305310. Mattiucci, S. & Nascetti, G. (2006) Molecular systematic, phylogeny and ecology of anisakid nematodes of the genus Anisakis Dujardin, 1845: an update. Parasite, 13, 99113. Mattiucci, S., Paggi, L., Nascetti, G., Santos, C.P., Costa, G., Di-Beneditto, A.P., Ramos, R., Argyrou, M., Cianchi, R. & Bullini, L. (2002) Genetic markers in the study of Anisakis typica (Diesing, 1860): Larval identification and genetic relationships with other species of Anisakis Dujardin, 1845 (Nematoda, Anisakidae). Systematic Parasitology, 51, 159170. Melo, M.F., Giese, E.G., Santos, J.N. & Portes Santos, C. (2011a) First record of Pseudoproleptus sp. (Nematoda: Cystidicolidae) in fish host. Acta Tropica, 117, 212215. Melo, M.F., Santos, J.N., Giese, E.G, Santos, E.G.N. & Portes Santos, C. (2011b). Raphidascaris (Sprentascaris) lanfrediae sp. nov. (Nematoda: Anisakidae) from the fish Satanoperca juruparii (Osteichthyes: Cichlidae). Memrias Instituto Oswaldo Cruz, 71, 623627. Mendona, J.M. (1963) Sbre dois Capilarineos parasitos de peixes (Nematoda, Trichuroidea). Memrias do Instituto Oswaldo Cruz, 61, 321327. Menezes, R.C. (2006) Camallanus cotti Fujita, 1927 (Nematoda, Camallanoidea) in ornamental aquarium fishes: pathology and morphology. Memrias do Instituto Oswaldo Cruz, 101, 683687. Menezes, R.C., Santos, S.M.C., Ceccarelli, P.S., Tavares, L.E.R., Tortelly, R. & Luque, J.L. (2011) Tissue alterations in the pirarucu, Arapaima gigas, infected by Goezia spinulosa (Nematoda). Revista Brasileira de Parasitologia Veterinria, 20, 207-209. Meschiatti, A.J. & Arcifa, M.S. (2009) A review on the fishfauna of Mogi-Guau River basin: a century of studies. Acta Limnologica Brasiliensia, 21, 135159. Mesquita, R.L.B., Azevedo, R.K., Abdallah, V.D. & Luque, J.L. (2011) Ectoparasites as numerical dominant species in parasite community of Trachelyopterus striatulus (Siluriformes: Auchenipteridae) from Guandu River, southeastern Brazil. Brazilian Journal of Biology 71, 623627. Molin, R. (1860) Una monografia del genere Spiroptera. Sitzungsberichte der Akademie der Wissenschaften in Wien, 38, 9111005. Monteiro, C.M., Santos, M.D., Zuchi, N.A. & Brasil-Sato, M.C. (2009) Ecological parameters of the endohelminths in relation to size and sex of Prochilodus argenteus (Actinopterygii: Prochilodontidae) from the Upper So Francisco River, Minas Gerais, Brazil. Zoologia, 26, 753757. Moravec, F. (1972) General characterization of the nematode genus Rhabdochona with a revision of the South American species. Acta Societatis Zoologicae Bohemoslovenicae, 36, 2946. Moravec, F. (1994) Parasitic nematodes of freshwater fishes of Europe. Kluwer Academic Publishers, Dordrecht, The Netherlands, 473 pp. Moravec, F. (1998) Nematodes of freshwater fishes of the Neotropical Region. Academia, Prague, Czech Republic, 464 pp. Moravec, F. (2001) Trichinelloid nematodes parasitic in cold-blooded vertebrates. Academia, Prague, Czech Republic, 429 pp. Moravec, F. (2006) Dracunculoid and anguillicoloid nematodes parasitic in vertebrates. Academia, Prague, Czech Republic, 634 pp. Moravec F., Kohn A. & Fernandes B.M.M. (1990) First record of Raphidascaris (Sprentascaris) hypostomi (Petter et Cassone, 1984) comb. n. and R. (S.) mahnerti (Petter et Cassone, 1984) comb. n. (Nematoda, Anisakidae) from Brazil with remarks on the taxonomic status of the genus Sprentascaris Petter et Cassone, 1984. Folia Parasitologica, 37, 131140. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1992a) Nematode parasites of fishes of the Paran River, Brazil. Part 1. Trichuroidea, Oxyuroidea and Cosmocercoidea. Folia Parasitologica, 39, 327353. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1992b) Three new species of oxyuroid nematodes, including two new genera, from freshwater catfishes in Brazil. Systematic Parasitology, 21, 189201. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1992c) Neoparaseuratum travassosi n.g., n.sp. (Nematoda, Quimperiidae), a new parasite from thorny catfish Pterodoras granulosus in Brazil. Memrias do Instituto Oswaldo Cruz, 87, 145150. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1993a) Nematode parasites of fishes of the Paran River, Brazil. Part 2. Seuratoidea, Ascaridoidea, Habronematoidea and Acuarioidea. Folia Parasitologica, 40, 115134. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1993b) Nematode parasites of fishes of the Paran River, Brazil. Part 3. Camallanoidea and Dracunculoidea. Folia Parasitologica, 40, 211229. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1993c) Travassosnema travassosi paranaensis subsp. n. and first description of the female of Guyanema raphiodoni Moravec, Kohn and Fernandes, 1993 (Nematoda, Guyanemidae), dracunculoid parasites of characid fishes in Brazil. Annales de Parasitologie Humaine et Compare, 68, 229233. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1994a) Structure of the cephalic end of two little-known oxyuroid genera, Travnema Pereira, 1938 and Cosmoxynemoides Travassos, 1949, parasites of fishes, as revealed by SEM. Journal of Helminthology, 68, 319 322. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1994b) Two new species of the genus Goezia, G. brasiliensis sp. n. and G. brevicaeca sp. n. (Nematoda, Anisakidae), from freshwater fishes in Brazil. Folia Parasitologica, 41, 271278. Moravec, F., Kohn, A. & Fernandes, B.M.M. (1997) New observations on seuratoid nematodes parasitic in fishes of the Paran River, Brazil. Folia Parasitologica, 44, 209223. Moravec, F., Kohn, A. & Santos, L.A. (1999) New data on Oncophora melanocephala (Nematoda, Camallanidae), a little known parasite of scombrid fishes. Parasite, 6, 7984.

NEMATODA IN BRAZILIAN FISHES

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83

Moravec, F. & Laoprasert, T. (2008) Redescription of Ichthyouris bursata Moravec & Prouza, 1995 (Nematoda: Pharyngodonidae), a parasite of wild and aquarium-reared discus Symphysodon spp. (Osteichthyes). Systematic Parasitology, 71, 137143. Moravec, F., Santos, M.D. & Brasil-Sato, M.C. (2008) Redescription of Cystidicoloides fischeri based on specimens from Piranhas in Brazil, and erection of a new genus (Nematoda, Cystidicolidae). Journal of Parasitology, 94, 889897. Moravec, F., Scholz, T., Dykov, I., Kuchta, R., Fiala, I. & Kohn, A. (2006) Redescription of Alinema amazonicum (Travassos, 1960) n. comb., a philometrid nematode with unusual morphology. Journal of Parasitology, 92, 138144. Moravec, F. & Sey, O. (1988) Nematodes of freshwater fishes. Part. I. Camallanoidea and Habronematoidea. Vstnk Ceskoslovensk Spolenosti Zooligisk, 52, 128148. Moravec, F. & Thatcher, V.E. (1997a) Procamallanus (Denticamallanus subgen. n.) dentatus sp. n. (Nematoda, Camallanidae) from the characid fish, Bryconops alburnoides, in the Brazilian Amazon. Parasite, 4, 239243. Moravec, F. & Thatcher, V.E. (1997b) New data on the morphology and systematic status of Klossinemella iheringi (Nematoda, Atractidae) from an Amazonian serrasalmid fish. Folia Parasitologica, 44, 4854. Moravec F. & Thatcher V.E. (1997c) Raphidascaroides brasiliensis n. sp. (Nematoda, Anisakidae), an intestinal parasite of the thorny catfish Pterodoras granulosus from Amazonia, Brazil. Systematic Parasitology, 38, 6571. Moravec, F. & Thatcher, V.E. (1999) Myleusnema brasiliense sp. n. (Nematoda, Kathlaniidae), a new intestinal parasite of the serrasalmid fish Myleus sp. in Brazil. Folia Parasitologica, 46, 216220. Moravec, F. & Thatcher, V.E. (2001) New oxyurid nematodes of the genera Ichthyouris and Spinoxyuris from South American freshwater fishes. Folia Parasitologica, 48, 311320. Moreira, L.H.A., Takemoto, R.M., Yamada, F.H., Ceschini, T.L. & Pavanelli, G.C. (2009) Ecological aspects of metazoan endoparasites of Metynnis lippincottianus (Cope, 1870) (Characidae) from Upper Paran River floodplain, Brazil. Helminthologia, 46, 214219. Moreira, L.H.A., Yamada, F.H., Ceschini, T.L., Takemoto, R.M. & Pavanelli, G.C. (2010) The influence of parasitism on the relative condition factor (Kn) of Metynnis lippincottianus (Characidae) from two aquatic environments: the upper Parana river floodplain and Corvo and Guairaca rivers, Brazil. Acta Scientiarum Biological Sciences, 32, 8386. Moreira, N.I.B., Oliveira, C.L. & Costa, H.M.A. (1991) New helminth parasite of fish: S. freitasi sp. n. (Nematoda, Camallanidae). Memrias do Instituto Oswaldo Cruz, 86, 429431. Moreira, N.I.B., Oliveira, C.L. & Costa, H.M.A. (1994) Spirocamallanus inopinatus (Travassos, Artigas & Pereira, 1928) e Spirocamallanus saofranciscensis sp. n. (Nematoda, Camallanidae) em peixes da represa de Trs Marias. Arquivo Brasileiro de Medicina Veterinaria e Zootecnia, 46, 485500. Moreira, N.I.B., Rocha, G.N. & Costa, H.M.A. (2000) A new nematode species (Seuratoidea, Cucullanidae) parasitizing Parauchenipterus striatulus (Steindachner, 1876) (Pisces, Auchenipteridae) in Brazil. Memrias do Instituto Oswaldo Cruz, 95, 3941. Moreira, S.T., Ito, K.F., Takemoto, R.M. & Pavanelli, G.C. (2005) Ecological aspects of the Iheringichthys labrosus (Ltken, 1874) (Siluriformes, Pimelodidae) in reservoirs of Paran river basin and upper Paran river floodplain, Brazil. Acta Scientiarum, Biological Sciences, 27, 317322. Motta, C.S. & Gomes, D.C. (1968) Sobre uma nova espcie do gnero Porrocaecum Railliet & Henry, 1912 (Nematoda, Anisakidae). Atas da Sociedade de Biologia do Rio de Janeiro, 11, 163164. Muniz-Pereira, L.C., Vieira, F.M. & Luque, J.L. (2009). Checklist of helminth parasites of threatened vertebrate species from Brazil. Zootaxa, 2123, 145. Nelson, J.S. (2006) Fishes of the world. John Wiley and Sons, New York, 601 pp. Padovani, R.E.S., Knoff, M., So-Clemente, S.C., Mesquita, E.F.M., Jesus, E.F.O. & Gomes, D.C. (2005) The effect of in vitro gamma radiation on Anisakis sp. larvae collected from the pink cusk-eel, Genypterus brasiliensis Regan, 1903. Revista Brasileira de Cincia Veterinria, 12, 137141. Paraguass, A.R, Alves, D.R. & Luque, J.L. (2005) Metazorios parasitos do acar Geophagus brasiliensis (Quoy & Gaimard, 1824) (Osteichthyes, Cichlidae) do reservatrio de Lajes, estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 14, 3539. Paraguass, A.R. & Luque, J.L. (2006) Ecologia das comunidades de metazorios parasitos de seis espcies de peixes do reservatrio de lajes, estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 16, 121128. Paraguass, A.R., Luque, J.L. & Alves, D.R. (2000) Aspectos quantitativos do parasitismo por larvas de anisakdeos (Nematoda, Ascaridoidea, Anisakidae) no pargo, Pagrus pagrus (Linaneus, 1758) (Osteichthyes, Sparidae) do litoral do Estado do Rio de Janeiro, Brasil. Contribuies Avulsas Sobre a Historia Natural do Brasil, 24, 18. Paraguass, A.R., Luque, J.L. & Alves, D.R. (2002) Community ecology of metazoan parasites of red porgy Pagrus pagrus (Osteichthyes, Sparidae) from the coastal zone of the state of Rio de Janeiro, Brazil. Acta Scientiarum, 24, 461467. Parra, J.E.P., Brandao, D.A. & Ceccarelli, P.S. (1997) Identificao e prevalncia de nematides do pacu Piaractus mesopotamicus (Holmberg, 1887), da Estao de Piscicultura do CEPTA, Pirassununga, SP, Brasil. Cincia Rural, 27, 291295. Pavanelli, G.C., Machado, M.H. & Takemoto, R.M. (1997) Fauna helmintolgica de peixes do rio Paran, regio de Porto Rico, Paran. In: Vazzoler, A.E.M., Agostinho, A.A. & Hahn, N.S. (Eds.), A plancie de inundao do alto rio Paran: Aspectos fsicos, biolgicos e socioeconmicos. Editora da Universidade Estadual de Maring, Maring, pp. 307329. Pavanelli, G.C., Machado, M.H., Takemoto, R.M., Guidelli, G.M. & Lizama, M.A.P. (2004) Helminth fauna of fishes: diversity and ecological aspects. In: Thomaz, S.M., Agostinho, A.A. & Hahn, N.S. (Eds.), The upper Paran river and its floodplain: Physical aspects, ecology and conservation. Backhuis Publishers, Leiden, pp. 309329. Pereira, C. (1935) Ascaridida e Spirurata parasitos de peixes do nordeste brasileiro. Archivos do Instituto Biolgico So Paulo, 6, 53 62. Pereira, C. (1938) Travnema travnema n. gen. n. sp. (Nematoda, Oxyuridae) parasito de Curimata elegans (Pisces, Characinidae) no nordeste brasileiro. In: Livro Jubilar do Professor Lauro Travassos. Tipografia do Instituto Oswaldo Cruz, Rio de Janeiro, Brasil, pp. 318385. Pereira, C. & Vaz, Z. (1933) Nematides parasitos de Cynolebias bellotti (Pisces, Cyprinodonta). Revista de Biologia e Hygiene, 4, 59 61.

84 Zootaxa 3082 2011 Magnolia Press

LUQUE ET AL.

Pereira, C., Vianna, D. & Azevedo, P. (1936) Biologia do nematide Procamallanus cearensis n. sp. Archivos do Instituto Biolgico, 7, 209226. Pereira-Jr., J. & Costa, M.A.S. (1996) Cucullanidae (Nematoda, Seratoidea) em Micropogonias furnieri (Desmarest, 1823) (Sciaenidae) do Rio Grande do Sul, com a descrio de Cucculanus cassinensis sp. n. e Dichelyne (Dichelyne) micropogonii sp. n. Comunicaes do Museu de Ciencias e Tecnologia da PUCRS. Serie zoologia, 9, 1130. Pereira-Jr., J., Almeida, F.M., Morais, N.C.M. & Vianna, R.T. (2004) Hysterothylacium sp. larvae (Nematoda, Anisakidae) in Micropogonias furnieri (Sciaenidae) from Rio Grande do Sul Coast, Brazil. Revista Atlntica, Rio Grande, 26, 5560. Pereira-Jr, J., Costa, M.A.S. & Vianna, R.T. (2002) ndices parasitolgicos de Cucullanidae (Nematoda, Seratoidea) em Micropogonias furnieri (Desmarest, 1823) no litoral do Rio Grande do Sul, Brasil. Atlntica, 24, 97101. Pereira Cardoso, T., Lima Salgado, R., Firmino Andrade, P., So Clemente, S.C. & Lima, F.C. (2006) Nematides da famlia Anisakidae e cestides da ordem Trypanorhyncha em peixes telesteos comercializados no estado do Rio de Janeiro. Revista Brasileira de Cincia Veterinria, 13, 98101. Petter, A.J. & Thatcher, V.E. (1988) Observations sur la capsule buccale de Spirocamallanus inopinatus (Nematoda), parasite de poisons brsiliens. Bulletin du Musum National d'Histoire Naturelle, 4 685692. Pinna, M.C.C. & Wosiacki, W. (2003) Family Trichomycteridae. In: Reis, R.E., Kullander, S.O. & Ferraris Jr., C.J. (Eds.), Check list of freshwater fishes of South and Central America. EDIPUCRS, Porto Alegre, pp. 270290. Pinto, R.M., Fbio, S.P. & Noronha, D. (1970) Sbre uma nova espcie do gnero Cucullanellus Trnquist, 1931 (Nematoda, Camallanoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 14, 7577. Pinto, R.M., Fbio, S.P., Noronha, D. & Rolas, F.J.T. (1974) Procamallanus brasileirosParte I (Nematoda, Camallanoidea). Memrias do Instituto Oswaldo Cruz, 72, 205211. Pinto, R.M., Fbio, S.P., Noronha, D. & Rolas, F.J.T. (1975) Novas contribuies ao conhecimento do gnero Procamallanus (Nematoda, Camallanoidea). Memrias do Instituto Oswaldo Cruz, 73, 183191. Pinto, R.M., Fbio, S.P., Noronha, D. & Rolas, F.J.T. (1976) Novas consideraes morfolgicas e sistemticas sobre os Procamallanus brasileiros (Nematoda, Camallanoidea). Memrias do Instituto Oswaldo Cruz, 74, 7784. Pinto, R.M. & Fernandes, B.M.M. (1972) Sobre Procamallanus probus n. sp. (Nematoda, Camallanoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 15, 133136. Pinto, R.M. & Noronha, D. (1972) Redescrio de Procamallanus inopinatus Travassos, Artigas & Pereira, 1928, (Nematoda, Camallanoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 15, 105108. Pinto, R.M. & Noronha, D. (1976) Procamallanus brasileiros (Nematoda, Camallanoidea): consideraes finais, com chave para determinao das espcies. Memrias do Instituto Oswaldo Cruz, 74, 323339. Pinto, R.M., Noronha, D., Knoff, M. & Gomes, D.C. (2010) Rhabdochona (R.) uruyeni (Nematoda, Rhabdochonidae) in Brazil: present status of South American Rhabdochona Railliet with a worldwide survey of the genus to 2010. Neotropical Helminthology, 4, 5069. Pinto, R.M., Vicente, J.J. & Noronha, D. (1984) First report of Ascarophis van Beneden, 1871: A. brasiliensis n. sp. (Nematoda, Ascarophidinae) and Procamallanus (S.) pereirai Annereaux, 1946 (Nematoda, Procamallaninae) in South America. Memrias do Instituto Oswaldo Cruz, 79, 491494. Pinto, R.M., Vicente, J.J. & Noronha, D. (1992) On some family related parasites (Nematoda, Cucullanidae) from the marine fish Paralonchurus brasiliensis (Steindacher, 1875) (Pisces, Ostraciidae). Memrias do Instituto Oswaldo Cruz, 87, 207212. Pinto, R.M., Vicente, J.J., Noronha, D. & Fbio, S.P. (1988) Redescription of Oncophora melanocephala (Rudolphi, 1819) BaudinLaurencin, 1971 (Nematoda, Camallanidae). Memrias do Instituto Oswaldo Cruz, 83, 233237. Rasheed, S. (1963) A revision of the genus Philometra Costa, 1945. Journal of Helminthology, 37, 89130. Rasheed, S. (1965) On a remarkable new nematode, Lappetascaris lutjani gen. et sp. n. (Anisakidae, Ascaridoidea), from marine fishes of Karachi and an account of Thynnascaris inquires (Linton, 1901) n. comb. and Goezia intermedia n. sp. Journal of Helminthology, 39, 313343. Rgo, A.A. (1979) Contribuio ao conhecimento dos helmintos de raias fluviais Paratrygonidae. Revista Brasileira de Biologia, 39, 879890. Rgo, A.A., Carvajal, J. & Schaeffer, G. (1985) Patogenia del higado de peces (Pagrus pagrus) provocada por larvas de nematodos Anisakidae. Parasitologia al Dia, 9, 7579. Rgo, A.A. & Eiras, J.C. (1988) Ecologia da parasitose de peixes e aves do rio Cuiab (Mato Grosso, Brasil) por Eustrongylides ignotus (Nematoda: Dioctophymidae). Actas do Colquio Luso-Espanhol sobre Ecologia das Bacias Hidrogrficas e Recursos Zoolgicos, pp. 335341. Rgo, A.A. & Gibson, D.I. (1989) Hyperparasitism by helminths: new records of cestodes and nematodes in proteocephalid cestodes from South American siluriform fishes. Memrias do Instituto Oswaldo Cruz, 84, 371376. Rgo, A.A. & Santos, C.P. (1983) Helmintofauna de cavalas Scomber japonicus Houtt, do Rio de Janeiro. Memrias do Instituto Oswaldo Cruz, 78, 443448. Rgo, A.A. & Vicente, J.J. (1988a) Eustrongylides ignotus Jgerskild, 1909 (Nematoda, Dioctophymatoidea), parasito de peixes, anfbios, rpteis e aves. Distribuio e taxonomia. Cincia e Cultura, 40, 478483. Rgo, A.A. & Vicente, J.J. (1988b) Excurso cientfica a Zona do Pantanal, Estado de Mato Grosso, para coletas de helmintos. Cincia e Cultura, 40, 6568. Rgo, A.A., Vicente, J.J., Santos, C.P. & Wekid, R.M. (1983) Parasitas de anchovas, Pomatomus saltatrix (L.) do Rio de Janeiro. Cincia e Cultura, 35, 13291336. Ribeiro, R.S., Luque, J.L. & Alves, D.R. (2002) Aspectos quantitativos dos parasitos da Maria-luiza, Paralonchurus brasiliensis (Osteichthyes, Sciaenidae), do litoral do Estado do Rio de Janeiro. Revista da Universidade Rural, Srie Cincias da Vida, 22, 151154. Rocha, C.A.M. (2011) Parasitic helminths of the freshwater Neotropical fich Hoplias malabaricus (Characiformes, Erythrinidae) from South America Basins. Reviews in Fisheries Science, 19, 150156. Rodrigues, E.L., Ranzani-Paiva, M.J.T. & Santos, A.A. (2002) Alteraes histolgicas em figado de dourado Salminus maxillosus Valenciennes, 1840 (Osteichthyes, Characidae) causadas por Neocucullanus neocucullanus Travassos, Artigas & Pereira 1828 (Nematoda). Acta Scientiarum, Biological Sciences, 24, 455459.

NEMATODA IN BRAZILIAN FISHES

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85

Rodrigues, H.O., Pinto, R.M. & Fernandes, B.M.M. (1971) Contribuio ao conhecimento de Heterotyphlum eurycheilum Olsen, 1952 (Nematoda, Ascaridoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 14, 177178. Rodrigues, H.O., Pinto, R.M. & Noronha, D. (1991) Key to the species of Brazilian Procamallanus with general considerations (Nematoda, Camallanoidea). Memrias do Instituto Oswaldo Cruz, 86, 107113. Rodrigues, H.O., Rodrigues, S.S. & Faria, Z. (1990) Contribution to the knowledge of the helminthological fauna of vertebrates of Marica, Rio de Janeiro State, Brazil. Memrias do Instituto Oswaldo Cruz, 85, 115116. Rodrigues, S.S. (1974) Raphidascaris atlanticus, nova espcie parasita de sardinha (Nematoda, Ascaridoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 17, 5961. Rudolphi, C.A. (1819) Entozoorum synopsis cui accedunt mantissa duplex et indices locupletissimi. Berolini, 811 pp. Saad, C.D.R & Luque, J.L. (2009) Larvas de Anisakidae na musculatura do pargo, Pagrus pagrus, no Estado do Rio de Janeiro, Brasil. Revista Brasileira de Parasitologia Veterinria, 18, 7173. Sabas, C.S.S. & Luque, J.L. (2003) Metazoan parasites of weakfish, Cynoscion guatucupa and Macrodon ancylodon (Osteichthyes, Sciaenidae), from the coastal zone of the State of Rio de Janeiro, Brazil. Revista Brasileira de Parasitologia Veterinria, 12, 171 178. Santos, C.P., Crdenas, M.Q. & Lent, H. (1999) Studies on Procamallanus (S.) pereirai Annereaux, 1946 (Nematoda, Camallanidae), with new host records and new morphological data on the larval stages. Memrias do Instituto Oswaldo Cruz, 94, 635640. Santos, C.P. & Gibson, D.I. (2007) Nilonema gymnarchi Khalil, 1960 and N. senticosum (Baylis, 1922) (Nematoda: Dracunculoidea): Gondwana relicts? Systematic Parasitology, 67, 225234. Santos, C.P., Lent, H. & Gibson, D.I. (2004) A new species of Pseudanisakis Layman & Borovkova, 1926 (Nematoda, Ascaridida) from Rioraja agassizii and Psammobatis extenta (Rajidae) in Brazilian southwestern Atlantic waters. Systematic Parasitology, 57, 229235. Santos, C.P. & Moravec, F. (2009a) Camallanus tridentatus (Drasche) (Nematoda, Camallanidae): new taxonomically important morphological data. Memrias do Instituto Oswaldo Cruz, 104, 9399. Santos, C.P. & Moravec, F. (2009b) Goezia spinulosa (Nematoda, Raphidascarididae), a pathogenic parasite of the arapaima Arapaima gigas (Osteichthyes). Folia Parasitologica, 56, 5563. Santos, C.P. & Moravec, F. (2009c) Tissue-dwelling philometrid nematodes of the fish Arapaima gigas in Brazil. Journal of Helminthology, 83, 295301. Santos, C.P., Moravec, F. & Venturieri, R. (2008) Capillostrongyloides arapaimae sp. n. (Nematoda: Capillariidae), a new intestinal parasite of the arapaima Arapaima gigas from the Brazilian Amazon. Memrias do Instituto Oswaldo Cruz, 103, 392-395. Santos, E. (1970) Nova espcie do gnero Raphidascaris Railliet & Henry, 1915 (Nematoda, Ascaridoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 14, 3940. Santos, E., Vicente, J.J. & Jardim, C.R. (1979) Helmintos de peixes de rios amaznicos da Coleo Helmintolgica do Instituto Oswaldo Cruz. II. Nematoda. Atas da Sociedade de Biologia do Rio de Janeiro, 20, 1119. Santos, M. & Brasil-Sato, M.C. (2004) Parasitos metazorios de Franciscodoras marmoratus (Reinhardt, 1874), serrudo (Siluriformes, Doradidae) do Rio So Francisco, Brasil. Revista Brasileira de Parasitologia Veterinria, 13, 1822. Santos, M.D., Albuquerque, M.C., Monteiro, C.M., Martins, A.N., Ederli, N.B. & Brasil-Sato, M.C. (2009) First report of larval Spiroxys sp. (Nemata, Gnathostomatidae) in three species of carnivorous fish from Trs Marias Reservoir, So Francisco River, Brazil. Pan American Journal of Aquatic Sciences, 4, 306311. Santos, M.D., Lemos-Pita, S.R.L.C. & Brasil-Sato, M.C. (2007) Fauna de parasitos metazorios de Pimelodus maculatus La Cpde, 1803 (Siluriformes, Pimelodidae) do rio Guandu, Estado do Rio de Janeiro, Brasil. Acta Scientiarum, 29, 101-107. Santos, S.M.C., Ceccarelli, P.S. & Luque, J.L. (2008) Helmintos parasitos do pirarucu, Arapaima gigas (Schinz, 1822) (Osteoglossiformes, Arapaimidae), no rio Araguaia, estado de Mato Grosso, Brasil. Revista Brasileira de Parasitologia Veterinria, 17, 171 173. Santos, S.M.C., Ceccarelli, P. S. & Rgo, R.F. (2003) Helmintos em peixes do Pantanal sul-mato-grossense: primeira expedio do Programa Pantanal. Boletim Tcnico do CEPTA, 16, 1526. So-Clemente,S.C., Lima, F.C. & Uchoa, C.M.A. (1995) Parasitos de Balistes vetula (L.) e sua importncia na inspeo do pescado. Revista Brasileira de Cincia Veterinria, 2, 3941. So-Clemente, S.C., Uchoa, C.M.A. & Serra Freire, N.M. (1994) Larvas de anisakideos em Pagrus pagrus (L.) e seu controle atravs de baixas temperaturas. Revista Brasileira de Ciencia Veterinaria, 1, 2124. Saraiva, A., Rosim, D.F. & Silva-Souza, A. T. (2006a) Nematode parasites of characoid fishes from Brazil. Bulletin of the European Association of Fish Pathologists, 26, 271274. Saraiva, A., Silva, F.A. & Silva-Souza, A.T. (2006b) Nematode parasites of the characid fish Brycon hilarii from the River Juba, Mato Grosso, Brazil. Helminthologia, 43, 158160. Saraiva, A., Silva, F.A. & Silva-Souza, A.T. (2006c) Neocucullanus neocucullanus Travassos, Artigas et Pereira, 1928 (Nematoda, Cucullanidae) from the Characidae fish, Brycon hilarii Valenciennes, 1850, from Brazil. Memrias do Instituto Oswaldo Cruz, 101, 669672. Sarmento, T.M., Fortes, E. & Hoffmann, R.P. (1995) Cucullanus debacoi sp.n. nematoda, parasito do intestino do Pimelodus maculatus Lacpde, 1803 (Pisces, Pimelodidae) do rio Guaiba, Porto alegre, Rio Grande do Sul. A Hora Veterinria, 15, 3840. Shibuya, A., Rosa, R.S. & Gadig, O.B.F. (2005) Stomach contents of Galeocerdo cuvieri and Carcharhinus plumbeus (Elasmobranchii, Carcharhinidae) caught off Paraba State, Brazil. Arquivos de Cincias do Mar, 38, 105107. Silva, L.O., Luque, J.L. & Alves, D.R. (2000a) Metazorios parasitos do peixe-espada, Trichiurus lepturus (Osteichthyes, Trichiuridae) do litoral do estado do Rio de Janeiro, Brasil. Parasitologia al Dia, 24, 97201. Silva, L.O., Luque, J.L., Alves, D.R. & Paraguass, A.R. (2000b) Ecologia da comunidade de metazorios parasitos do peixe espada Trichiurus lepturus Linnaeus (Osteichthyes, Trichiuridae) do litoral do estado do Rio de Janeiro, Brasil. Revista Brasileira de Zoocincias, 2, 115133. Silva, R.J., Baldassin, P., Gonzlez, G.K., Sakiyama, D.T.P., Lavor, U.L., Werneck, M.R. & Gallo, H. (2008) Proleptus acutus Dujardin, 1845 (Nematoda, Physalopteridae) parasite of rays Zapteryx brevirostris Mller & Henle, 1841 (Rhinobatiformes, Rhinobatidae) in Brazil. Arquivo Brasileiro de Medicina Veterinria e Zootecnia, 60, 15731575.

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Silva-Souza, A.T. & Saraiva, A. (2002) Ecological data of Travassosnema travassosi travassosi (Dracunculoidea, Guyanemidae) from the humour of the eyes of Acestrorhynchus lacustris from Tibagi River, Parana, Brazil. Memrias do Instituto Oswaldo Cruz, 97, 5152. Sprent, J.F.A. (1990) Some ascaridoid nematodes of fishes: Heterocheilinae. Systematic Parasitology, 16, 149161. Takemoto, R.M., Amato, J.F.R. & Luque, J.L. (1996) Comparative analysis of the metazoan parasite communities of leatherjackets, Oligoplites palometa, O. saurus and O. saliens (Osteichthyes, Carangidae) from Sepetiba Bay, Rio de Janeiro, Brazil. Revista Brasileira de Biologia, 56, 639650. Takemoto, R.M. & Lizama, M.A.P. (2010) Helminth fauna of fishes from the Upper Parana River Floodplain, Brazil. Neotropical Helminthology, 4, 5-8. Takemoto, R.M., Lizama, M.A.P., Guidelli, G.M. & Pavanelli, G.C. (2004) Parasitos de peixes de guas continentais. In: RanzaniPaiva, M.J.T., Takemoto, R.M. & Lizama, M.A.P. (Eds.), Sanidade de organismos aquticos. Varela, So Paulo, pp. 179198. Takemoto, R.M., Pavanelli, G.C., Lizama, M.A.P., Lacerda, A.C.F.,Yamada, F.H., Moreira, L.H.A., Ceschini, T.L. & Bellay, S. (2009) Diversity of parasites of fish from the Upper Paran River floodplain, Brazil. Brazilian Journal of Biology, 69, 691705. Tavares, L.E.R, Bicudo, A.A. & Luque, J.L. (2004a) Metazoan parasites of the needlefish Tylosurus acus (Lacpde, 1803) (Osteichthyes, Belonidae) from the coastal zone of the state of Rio de Janeiro, Brazil. Revista Brasileira de Parasitologia Veterinria, 13, 3640. Tavares, L.E.R. & Luque, J.L. (2004a) Community ecology of the metazoan parasites of white catfish Netuma barba (Osteichthyes, Ariidae) from the coastal zone of the State of Rio de Janeiro,Brazil. Brazilian Journal of Biology, 64, 18. Tavares, L.E.R. & Luque, J.L. (2004b) Community ecology of the metazoan parasites ofcommon snook Centropomus undecimalis (Osteichthyes, Centropomidae) from the coastal zone of the state of Rio de Janeiro, Brazil. Brazilian Journal of Biology, 64, 523 529. Tavares, L.E.R. & Luque, J.L. (2006) Sistemtica, biologia e importncia em sade coletiva de larvas de Anisakidae (Nematoda, Ascaridoidea) parasitas de peixes sseos marinhos do Estado do Rio de Janeiro, Brasil. In: Silva-Souza, A.T. (Ed.), Sanidade de Organismos Aquticos no Brasil. Abrapoa, Maring, pp. 297328. Tavares, L.E.R. & Luque, J.L. (2008) Similarity between metazoan parasite communities of two sympatric brackish fish species from Brazil. Journal of Parasitology, 94, 985989. Tavares, L.E.R., Luque, J.L. & Bicudo, A.A. (2004b) Metazoan parasites of the Brazilian menhaden, Brevoortia aurea (Spix & Agassiz, 1829) (Osteichthyes, Clupeidae) from the coastal zone of the State of Rio de Janeiro, Brazil. Brazilian Journal of Biology, 64, 553554. Tavares, L.E.R., Luque, J.L. & Bicudo, A.A. (2005) Community ecology of the metazoan parasites of anchovy Anchoa tricolor (Osteichthyes, Engraulidae) from the coastal zone of the State of Rio de Janeiro, Brazil. Brazilian Journal of Biology, 65, 533540. Tavares, L.E.R., Luque, J.L. & Botelho-Neto, S.L. (2001) Ecologia da comunidade de metazorios parasitos do olhodeco, Priacanthus arenatus (Cuvier, 1829) (Osteichthyes, Priacanthidae) do litoral do Estado do Rio de Janeiro, Brasil. Revista Brasileira de Zoocincias, 3, 4559. Tavares, L.E.R., Saad, C.D.R., Cepeda, P.B. & Luque, J.L. (2007) Larvae of Terranova sp. (Nematoda, Anisakidae) parasitic in Plagioscion squamosissimus (Perciformes, Sciaenidae) from Araguaia river, state of Tocantins, Brazil. Revista Brasileira de Parasitologia Veterinria, 16, 110115. Tavares-Dias, M., Brito, M.L.S. & Lemos, J.R.G. (2009) Protozorios e metazorios parasitos do cardinal Paracheirodon axelrodi Schultz, 1956 (Characidae), peixe ornamental proveniente de exportador de Manaus, Estado do Amazonas, Brasil. Acta Scientiarum, Biological Sciences, 31, 2328. Tavares-Dias, M., Lemos, J.R.G. & Martins, M.L. (2010) Parasitic fauna of eight species of ornamental freshwater fish species from the middle Negro River in the Brazilian Amazon Region. Revista Brasileira de Parasitologia Veterinria, 19, 103-107. Tavernari, F.C., Takemoto, R.M., Guidelli, G.M., Perez Lizama, M.A., Figueiredo Lacerda, A.C. & Pavanelli, G.C. (2009) Parasites of Auchenipterus osteomystax (Osteichthyes, Auchenipteridae) from two different environments, Rosana's reservoir and upper Parana river floodplain, Brazil. Acta Scientiarum Biological Sciences, 31, 49-54. Thatcher, V.E. (1981) Patologia de peixes da Amaznia brasileira, 1. Aspectos gerais. Acta Amazonica, 11, 125140. Thatcher, V.E., (1991) Amazon fish parasites. Amazoniana, 11, 263572. Thatcher, V.E. (2006) Amazon fish parasites. Pensoft Publishers, Sofia and Moscow. 508 pp. Thatcher, V.E. & Oliveira, E.F. (1991) Paracamallanus amazonensis sp. n. (Nematoda, Camallanidae) um parasita de bagre, Hypophthalmus edentatus (Pisces, Hypophthalmidae) da Amazonia brasileira. Amazoniana, 12, 16. Thatcher, V.E. & Padilha, T.N. (1977) Spinitectus jamundensis sp. n. (Nematoda, Spiruroidea) from a Colombian freshwater fish (Prochilodus reticulatus). Revista Brasileira de Biologia, 37, 799801. Timi, J.T., Lanfranchi, A.L., & Luque, J.L. (2010) Similarity in parasite communities of the teleost fish Pinguipes brasilianus in the southwestern Atlantic: Infracommunities as a tool to detect geographical patterns. International Journal for Parasitology, 40, 243 254. Timi, J.T., Lanfranchi, A.L., Tavares, L.E.R. & Luque, J.L. (2009) A new species of Dichelyne (Nematoda, Cucullanidae) parasitizing sciaenid fishes from off the South American Atlantic coast. Acta Parasitologica, 54, 4552. Timi, J.T., Luque, J.L. & Sardella, S. (2005) Parasites of Cynoscion guatucupa along South American Atlantic coasts: evidence for stock discrimination. Journal of Fish Biology, 67, 16031618. Travassos, L. (1919) Informaes sobre o material helmintolgico colecionado na ilha Trinidade em 1916. Archivos do Museu Nacional do Rio de Janeiro, 22, 159167. Travassos, L. (1920) Esbo de uma chave geral dos nematdeos parasitos. Revista Veterinria e Zootecnia, 10, 5970. Travassos, L. (1923) Informaes sobre a fauna helmintolgica de Mato Grosso. Folha Mdica. 5, 2930. Travassos, L. (1925) Contribuio ao conhecimento da evoluo dos Dioctophymoidea. Brazil Medico, 2, 286287. Travassos, L. (1927) Uma nova Capillaria parasita de peixes de gua doce: Capillaria sentinosa n. sp. Boletin Biologico, 10, 215217. Travassos, L. (1940a) Relatrio da terceira excurso zona da Estrada de Ferro Noroeste do Brasil, realizada em fevereiro e maro de 1940. II. Pesquisas helimintolgicas. Memrias do Instituto Oswaldo Cruz, 35, 607634. Travassos, L. (1940b) Relatrio da quarta excurso do Instituto Oswaldo Cruz zona da Estrada de Ferro Noroeste do Brasil, realisada

NEMATODA IN BRAZILIAN FISHES

Zootaxa 3082 2011 Magnolia Press

87

em agosto e setembro de 1940. Memrias do Instituto Oswaldo Cruz, 35, 698722. Travassos, L. (1945) Relatrio da excurso do Instituto Oswaldo Cruz ao rio Parana (Porto Cabral), em maro e abril de 1944. Memrias do Instituto Oswaldo Cruz, 42, 151165. Travassos, L. (1947) Contribuio ao conhecimento da fauna helmintolgica dos peixes dgua doce do Brasil. III. Duas novas espcies do gnero Cucullanus Meller, 1877. Memrias do Instituto Oswaldo Cruz, 45, 551559. Travassos, L. (1948) Contribuio ao conhecimento da fauna helmintolgica dos peixes dgua doce do Brasil IV. Dois novos gneros de Cosmocercidae (Nematoda) e uma nota de nomenclatura helmintolgica. Memrias do Instituto Oswaldo Cruz, 46, 633641. Travassos, L. (1960) Sobre nematdeos cavitrios de peixes do Rio Amazonas. Atas da Sociedade de Biologia do Rio de Janeiro, 4, 1520. Travassos, L., Artigas, P. & Pereira, C. (1928) Fauna helmintolgica dos peixes de gua doce do Brasil. Archivos do Instituto Biolgico, 1, 568. Travassos, L. & Freitas, J.F.T. (1940) Relatrio da excurso cientfica realizada na zona da Estrada de Ferro Noroeste do Brasil em julho de 1939. Memrias do Instituto Oswaldo Cruz, 35, 525556. Travassos, L. & Freitas, J.F.T. (1941) Relatrio da quinta excurso do Instituto Oswaldo Cruz realizada zona da Estrada de Ferro Noroeste do Brasil, em janeiro de 1941. II. Memrias do Instituto Oswaldo Cruz, 36, 272295. Travassos, L. & Freitas, J.F.T. (1942) Relatrio da sexta excurso do Instituto Oswaldo Cruz, realizada zona da Estrada de Ferro Noroeste do Brasil, em novembro de 1941. Memrias do Instituto Oswaldo Cruz, 37 259286. Travassos, L. & Freitas, J.F.T. (1943) Relatrio da stima excurso cientfica do Instituto Oswaldo Cruz realizada zona da estrada de ferro Noroeste do Brasil, em maio de 1942. Memrias do Instituto Oswaldo Cruz, 38, 385412. Travassos, L. & Freitas, J.F.T. (1948) Relatrio da excurso do Instituto Oswaldo Cruz ao norte do Estado do Esprito Santo, junto ao parque de reserva e refgio Soretama, em fevereiro e maro de 1948. Memrias do Instituto Oswaldo Cruz, 46, 605631. Travassos, L. & Freitas, J.F.T. (1964) Pesquisas helmintolgicas realizadas em Maicur, estado do Par. Publicaes Avulsas do Museu Paraense Emilio Goeldi, 2, 316. Travassos, L. & Kohn, A. (1965) Lista dos helmintos parasitos de peixes encontrados na estao experimental de biologia e piscicultura de Emas, Pirassununga, Estado de So Paulo. Papis Avulsos do Departamento de Zoologia de So Paulo, 17, 3552. Travassos, L., Freitas, J.F.T. & Lent, H. (1939) Relatrio da excurso cientfica do Instituto Oswaldo Cruz realizada na zona da Estrada de ferro Noroeste do Brasil, em outubro de 1938. Pesquisas helmintolgicas. Boletin Biolgico, 4, 221249. Travassos, L., Freitas, J.F.T. & Mendona, J.M. (1964) Relatrio da excurso do Instituto Oswaldo Cruz ao Parque de Reserva e Refgio Soretama, no Estado do Espirito Santo, em outubro de 1963. Boletim do Museu de Biologia Mello Leito, 23, 126. Travassos, L., Freitas, J.F.T., Mendona, J.M. & Rodrigues, H.O. (1963) Terceira excurso a Cabo Frio, Estado do Rio de Janeiro. Atas da Sociedade de Biologia do Rio de Janeiro, 7, 67. Vaz, Z. & Pereira, C. (1934) Contribuio ao conhecimento dos nematides de peixes fluviais do Brasil. Archivos do Instituto Biolgico de So Paulo, 5, 87103. Venancio, A.C.P., Aguiar, G.R., Lopes, P.S. & Alves, D.R. (2010) Metazoan parasites of Mandi-amarelo Pimelodus maculatus and of Jundi Rhamdia quelen (Osteichthyes: Siluriformes) of Paraba do Sul River, Volta Redonda, Rio de Janeiro. Revista Brasileira de Parasitologia Veterinria, 19, 157163. Vicente, J.J. & Fernandes, G.L. (1978) Contribuio ao conhecimento dos helmintos de Bagre bagre (Linnaeus, 1766) Fowler, 1841 e de Macrodon ancylodon (Bloch, 1801) Jordan, Evermann & Clark, 1930, no litoral da ilha de So Lus, estado do Maranho, Brasil. Boletim do Laboratrio de Hidrobiologia, 2, 9196. Vicente, J.J. & Jardim, C.R. (1980) Filardeos da Coleo Helmintolgica do Instituto Oswaldo Cruz. I. Peixes, anfbios e rpteis. Atas da Sociedade de Biologia do Rio de Janeiro, 21, 4757. Vicente, J.J., Mincarone, M.M. & Pinto, R.M. (2002) First report of Lappetascaris lutjani Rasheed, 1965 (Nematoda, Ascaridoidea, Anisakidae) parasitizing Trachipterus arawatae (Pisces, Lampridiformes) on the Atlantic Coast of Brazil. Memrias do Instituto Oswaldo Cruz, 97, 9394. Vicente, J.J. & Pinto, R.M. (1981) Nematoda, zooparasitic forms. In: Hurlbert, S.H. & Santos, N. (Eds), Aquatic biota of tropical South America, Part 2. Anarthropoda. San Diego State University, San Diego, pp. 136145. Vicente, J.J. & Pinto, R.M. (1999) Nematides do Brasil. Nematides de peixes. Atualizao: 1985-1998. Revista Brasileira de Zoologia, 16, 561610. Vicente, J.J. & Santos, E. (1972) Sobre um novo nematdeo camalandeo, parasito de peixe, marinho (Nematoda, Camallanoidea). Atas da Sociedade de Biologia do Rio de Janeiro, 15, 145147. Vicente, J.J. & Santos, E. (1973) Alguns helmintos de peixe do litoral norte fluminenseI. Memrias do Instituto Oswaldo Cruz, 71, 95113. Vicente, J.J. & Santos, E. (1974) Alguns helmintos de peixes do litoral norte fluminenseII. Memrias do Instituto Oswaldo Cruz, 72, 173180. Vicente, J.J., Pinto, R.M. & Aguilera, O. (1989) On Dichelyne (Cucullanellus) elongatus (Tornquist, 1931) Petter, 1974: South American correlated species (Nematoda, Cucullanidae) and some other helminths of Micropogonias furnieri (Desmarest, 1823) (Pisces, Sciaenidae). Memrias do Instituto Oswaldo Cruz, 84, 357361. Vicente, J.J., Rodrigues, H.O. & Gomes, D.C. (1985) Nematides do Brasil. 1 parte: Nematides de peixes. Atas da Sociedade de Biologia do Rio de Janeiro, 25, 179. Vicentin, W.,Vieira, K.R.I., Costa, F.E.S., Takemoto, R.M., Tavares, L.E.R. & Paiva, F. (2011) Metazoan endoparasites of Serrasalmus marginatus (Characiformes: Serrasalminae) in the Negro River, Pantanal, Brazil. Revista Brasileira de Parasitologia Veterinria, 20, 61-63. Vieira, K.R.I., Vicentin, W., Paiva, F., Pozoc, C.F., Borges, F.A., Adriano, E.A., Costae, F.E.S. & Tavares, L.E.R. (2010) Brevimulticaecum sp. (Nematoda, Heterocheilidae) larvae parasitic in freshwater fish in the Pantanal wetland, Brazil. Veterinary Parasitology, 172, 350354. Yamaguti, S. (1961) Systema helminthum. Vol. III. The nematodes of vertebrates. Interscience Publishers, New York, 1261 pp.

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