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STRUCTURAL, PHYSIOLOGICAL, SYSTEMATICAL,

AND MEDICAL

X,

BEING A FOURTH EDITION OF

THE OUTLINE OF THE FIRST PRINCIPLES OF BOTANY.

BY JOHN LINDLEY,

PH.D. F.R.S.

VICE-SECRETARY OP THE HORTICULTURAL SOCIETY OP LONDON;


PROFESSOR OP BOTANY IN UNIVERSITY COLLEGE, LONDON,
THE ROYAL INSTITUTION OP GREAT BRITAIN, AND TO THE SOCIETY OP APOTHECARIES.

LONDON:
'PRINTED FOR TAYLOR AND WALTON,
BOOKSELLERS AND PUBLISHERS TO UNIVERSITY COLLEGE,

UPPER GOWER STREET.


1841.

<

LONDON

PRINTED BY SAMUEL BENTLEY,


Bangor House, Shoe Lane.

PREFACE.

THE work now laid before the public is a fourth edition of


the Author's " Outline of the First Principles of Botany,"

much extended

That work was


and, it is hoped, improved.
written for the use of students, and entirely for the purpose
of enabling

them

to fix correctly in their

minds the more

important points which the teacher brings before them in an


When facts are mixed up with extended
academical course.
and
discussions,
rapidly adverted to, either in a lecture-room
or in a written dissertation, the beginner is apt to lose sight of
the exact nature of an argument, and is unable to distinguish

with certainty the points upon which

it is

most material

for

him to fix his attention. That there existed a want of such


a work has been sufficiently proved by the many editions the
original Outline has passed through, in various
guages indeed, while the present new edition
:

advice has been

European

lan-

in the press,
received of the translation of the work into
is

The propositions which it contained were such


of the most indispensable importance for a student to
understand ; and were all, apparently, deducible from the
Hungarian.

as

it is

evidence which had at that time been collected by Botanists.


The wish of the Author was to sketch a slight but accurate
outline, the

details

of which were to be

filled

up by the

reader himself, who, for this purpose, was referred to the


Author's more extended Introductions to Botany.
The original " Outline " contained nothing more than the
fundamental propositions upon which the principles of Organic

and Physiological Botany depend


but, when two editions
had been exhausted, the Author was induced, by the favour
with which the book had been received, and by its recognized
utility, notwithstanding its many defects, to combine with it a
;

2091 \ 27

PKEFACE.

IV

sketch of Systematical Botany, treated in the same manner.


He undertook the far more difficult task of reducing to their
that distinguish the various
simplest expression the characters
classified
which
are
in
by modern systematiplants
groups
the object being to diminish, by a very careful
analysis, the difficulties which present themThe
selves to the student of this branch of the subject.

cal writers

and extensive

made in the form of a series of tables, called


" Alliances of Plants
;" and it has been satisfactory to

attempt was
the

the Author to find that this too has been advantageous to


The work
students, notwithstanding its extreme conciseness.
"
thus altered appeared in 1835, under the title of
Key to
Structural, Physiological, and Systematical Botany."
In the edition now offered to students many important

improvements have been introduced, without deviating from


the original plan of the work. The skill of the wood-engraver
has enabled the Author to

fill

his

pages with

illustrations, ex-

planatory not only of the technical terms employed in Botany,


but also of the Natural Orders of plants. An analysis of the
latter, upon the plan of Lamarck, an account of De Candolle's
celebrated system of arrangement, into which a large number
of wood-cuts are introduced, and some new views relating to
natural classification, are added to the matter to be found in

previous editions besides which, the whole of the Structural


and Physiological part has been corrected with great care,
and made to include all the most important views of modern
:

physiologists, so as to present the reader with a view of the


state of Botanical knowledge in these
departments in the

spring of 1841.
It is

what

it

hoped that these improvements will render the work


was originally intended for, a complete Botanical

wherein all the principal topics which the teachers


Note-book,
of Botany introduce into their lectures are
arranged methodiThe student will naturally look to his instructor or to
cally.

more extensive works


in his

for explanations of those


points

Note-book are merely adverted

University College, London,

April 1841.

to.

which

ELEMENTS OF BOTANY;
STRUCTURAL, PHYSIOLOGICAL, SYSTEMATICAL,

AND MEDICAL.

I.

STRUCTURAL AND PHYSIOLOGICAL


BOTANY.

PLANTS are not separable from animals by any absolute


character; the simplest individuals of either kingdom not
being distinguishable by our senses.
2. Animals are for the most part incapable of multiplying
by mechanical or spontaneous division of their trunk, and are
supported by nutritious matter, carried into their system from
an internal bag or stomach.
3. Plants are for the most part congeries of individuals,
1.

multiplying by spontaneous or artificial division of their trunk


or axis, and are supported by nutritious matter conveyed into
their system by the absorption of their lower extremities
or roots, or

by

their surface.

Generally speaking, the latter are fixed to some substance from which they grow, are destitute of locomotion, and
are enabled to digest their food by the action of light upon
4.

their epidermis.
5. Plants consist

of a hygrometrical membranous transparent tissue, chemically composed of oxygen, hydrogen, and


carbon, to which nitrogen is always superadded.
They are
also found to contain many mineral substances, which they
are supposed to separate from their proper food during the
process of digestion, and to deposit in their tissue.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

Their component parts are held together by an organic


mucus, out of which the tissue itself is generated.
the woody, the
7. Tissue is found in the form of the cellular,
each of which has
and the
the
6.

vascular,

laticiferous,

pitted,

certain modifications, constituting the

Elementary organs.

ELEMENTARY ORGANS.

I.

CELLULAR TISSUE (Tela cellulosa, Lat. ; Tissu


old writers ; Zellencellulaire, Fr. ; Pulp and Parenchyma, of
found in plants ;
gewebe, Germ.) is the only form universally
8.

Of

these,

the other forms are often either partially or entirely wanting.


9. Cellular tissue is composed of vesicles, the sides of which
are not originally perforated by visible pores (22).
10. Each vesicle is a distinct individual, cohering with the
vesicle

with which

it

is

and originating from a


;
which either remains visible on its

in contact

primitive point or cytollast


sides or is absorbed.

1
,

11. Therefore the apparently simple membrane


two contiguous cells is in fact double.

which

di-

vides

12. If the adhesion of the contiguous cells be imperfect,


Such spaces are called interspaces will exist between them.
cellular passages.
13. The sides of cellular tissue are often thickened

by the

on their inner surface, of matter of lignification or


z
which is stratified, and often pierced with passages
sclerogen
deposit,

leading to the circumference.


14. The cells contain fluid;

grains of colouring matter


or
(cJiromule., chromogen,
chlorophyll) ; starch in granules (peand
renchyma) ;
crystals, which, when acicular, are named
raphides.
15. The

vesicles

of cellular

tissue,

when

separate,

are

round or oblong ; when slightly and equally pressed together,


4
with an hexagonal
they acquire a dodecahedral appearance
,

section

stretched lengthwise they

drical, fusiform,

16.

When

together

by

become

cellular tissue

is

composed of

vesicles fitting

called in general terms


prosencJiyma if the vesicles are fusiform.

their plane faces,

it

is

parenchyma ; and
Both these are sometimes branched, and
culate.

prismatical, cylin-

&c.

their divisions inos-

ELEMENTARY ORGANS.
4
or spharendtyma ; conical, conenchySpheroidal cellular tissue is merenchyma ,
ma 5 ; oval, ovenchynui 6 ; fusiform, atractenchyma ; cylindrical, cylindrenchyma 8 ;
w;
9
prismatical, prismenchyma,
sinuous, colpenckyma ; branched, cladenchyma
1
7
which, when compressed, becomes muriform ; stellate, actinenchyma *; entann.
and
branched
tubular, d(edalencliyma
gled,

=-.<#4

17. Parenchyma constitutes all the pulpy parts; the medulla or pith (98), the medullary rays (132), a portion of the
bark (120), and all that intervenes between the veins of leaves

and other appendages of the axis. Consequently it occurs in


every part of a plant, and especially in those which are succulent.
It, however, sometimes acquires, by the deposit of scle3
rogen (13), excessive hardness, as in the stone of fruits and
the bony skin of some seeds.
,

destines are large

of Parenchyma, in which raphides (60 a) are often de-

cells

posited.

18.

Prosenchyma

which

it

when

is

confined to the bark and

wood, in

only occasionally occurs.


19. Besides these a spiral line is often found in the inside of

cell,

and

it

13

or inenchyma, is produced ;
sometimes happens that the membrane connecting such
fibro-cellular tissue

fibres is absorbed,

leaving the fibres only to constitute

the

cell.

20.

The

function of the cellular tissue

in all directions

the

is

membrane of which

to transmit fluids
it

is

composed

is

therefore permeable, although not in general furnished with


B 2

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

thickened by the deposition of


with
are left in the latter communicating
sclerogen, passages
of
being
the sides of the tissue, and giving it the appearance

When

visible pores (9).

is

it

dotted or pitted.

one

cell

generating
21. Cellular tissue is self-productive,
In Chara, Marchantia, &c. young
others upon its surface.
in the spaces
cells are said to be formed at the points of and
cells are
new
between older cells ; in Confervse and in anthers

formed by the internal divisions of an older cell; while,


mode of production is
according to Schleiden, the most general
of vegetation (6).
mucus
in
the
from cytoblasts (10), generated
of the
modification
a
is
22. PITTED TISSUE (Boihrenchyma)
of ordinary cylindrical cells placed
and forming continuous

cellular, either consisting

end to end, opening


tubes

into each other,

or originally tubular

15

Its sides are

marked by

pits,

consequence of the sclerogen (13)


resembling dots, produced
It is
the inside of the cells.
over
being unequally deposited
common in wood, of which it forms what is vulgarly called
in

the porosity.
Its office is to convey fluids with rapidity in
the direction of the woody tissue that surrounds it. Formerly
it

was considered a form of vascular

ducts, or

msiform

tissue,

and called dotted

tissue.

Pitted Tissue is articulated, when composed of short cylinders placed end to


end, or continuous when it was originally tubular.

23. WOODY TISSUE (Pleurenchyma) consists of elongated


tubes tapering to each end, and, like the vesicles of cellular
It may be considered a form of
tissue, imperforate to the eye.

the cellular tissue


it

is

itself,

to

which

practically distinguished

length, extreme fineness,

and

it is

by

frequently referred

but

its

cylindrical form, great


toughness; the latter of which

properties is produced by the thickness of its sides.


24. It is found in the wood, among the parenchyma of the
liber (124),

and

in the veins of the leaves, or other

appendages

of the axis.
25. Its functions are to give
strength to the vegetable fabric,
to serve as a medium for the
passage of fluid from the

and

lower to the upper extremities.


Common Pleurenchyma

14
has its sides destitute of markings ; the
is a
glandular
variety in which the sides of the tubes are furnished with circular disks ; the
latter occur chiefly in coniferous
plants and such as have aromatic secretions.

ELEMENTARY ORGANS.
VASCULAR TISSUE (Trachmchyma)

26.

consists of very thin-

sided cylinders tapering to each end, and having a spiral fibre

generated in their inside.

Of

27.

this

Their

fibre is

rolling

when

kind of tissue spiral vessels 1516 are the type.


of a highly elastic nature, and is capable of un-

stretched.

28. Spiral vessels are found in the medullary sheath, and in


all parts that emanate from it,
especially the veins of the
leaves,

and everything that

is

a modification of them.

usually absent from the wood and bark.


They, however, occur in these and other unusual parts in a
few extremely rare cases ; as in the wood, and bark, and pith
29.

They

are

of Nepenthes.
30. The spiral vessels appear intended for the conveyance
of air, which has been found to contain 7 or 8 per cent, more
oxygen than the atmosphere.
31. Ducts are transparent tubes, the sides
rings, bars, or transverse streaks.

of which are

marked with
32.

They

are slight modifications of the spiral vessel, differ-

ing principally in being incapable of unrolling ; and, in some


cases, in the turns of the spiral fibre being distant or broken,
or even, in appearance, branched.
33. In those cases

where the turns of the

spire actually

touch each other, the ducts, which are then called closed, can
only be distinguished from spiral vessels by their inability to
unrol ; while at rest they appear to be absolutely the same.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

Ducts are closed 50 when the spires touch each other


17

to consist of separate rings


scalariform, when the lines
septate,

when

the interior

when they

annular,

seern
19

reticulated, when the spires cross each other


upon their sides are horizontal and equidistant

is

divided

by

pierced disks, as in Echinocactus.

Ducts occur among the woody tissue of herbaceous


plants ; are abundant in the wood of the higher tribes of cellular plants, such as Ferns and Lycopodiacese ; and their ends
are often in immediate connection with the loose cellular tissue
34.

occupying the extremities of the roots.


35. Their functions have not been accurately determined.
It is probable that they act as. spiral vessels when
young ; but
it

certain that

is

they become

filled

with

as soon as

fluid

their spires are separated.

36. LATICIFEROUS TissuE 18 (Cinenchyma) consists of uninterrupted anastomozing tubes, whose final divisions are so delicate, that

the

eye only discovers them when aided by the


It forms the proper vessels of

most powerful microscopes.


old writers.

37. It principally occurs in the liber of


Exogens (124),
whence the ramifications proceed to the surface of all the
organs, and penetrate the hairs, where they form a most
delicate network.

38. Laticiferous tissue


conveys latex, a peculiar fluid, usually
and coloured red, white, or yellow ; often however

turbid,

colourless.

The use of

39.

this tissue is to

carry the latex to

all

the

newly formed organs, which are supposed to be nourished by


The

large trunks of Cinenchyma are vasa expansa, or opopkora


rasa cm: tracta.

40. There are no other


vessels,

Reservoirs of

oil,

elementary forms of

Lenticular glands, are

the small are

tissue.
all

it.

Air-

either dis-

tended intercellular
passages, or cavities built up with cellular
tissue, or large cells filled with
peculiar secretions.
41. When such cavities are essential to
the existence of a
species,

they are formed by a regular -arrangement of cellular


a definite and
unvarying figure ; Ex. Water-plants.
they are not essential to the existence of a

tissue in

When

species,

they are mere irregular distensions or lacerations of the tissue


Ex. Pith of the Walnut-tree.

42
lied

All these forms of tissue are


enclosed within a skin
the epidermis, which is one
or more external
layers of

ELEMENTARY ORGANS.

parenchyma, the vesicles of which are compressed, and in a


firm state of cohesion.

43.

The spaces seen upon the

epidermis,

when examined by

a microscope, represent these vesicles.


44. It is, therefore, not a peculiar membrane, but a form of
cellular tissue.

45. It

posed to
46. It

is

air,
is

spread over all the parts of plants which are exexcept the stigma (397).
not found upon parts habitually living

under

water.
47. It

which

is

is itself

by an extremely

protected

thin

pellicle,

apparently inorganic and homogeneous, and which

covers every part, except the openings through the stomates


This membrane is the cuticle.
(49).

The epidermis is furnished with stomates.


49. STOMATES are oval spaces lying between the sides of the
cells, opening into intercellular cavities in the subjacent tissue,
48.

and appearing
from above 22

to be bordered

by a limb when they

are viewed

23 25
.

50. This appearance. of a limb is


owing to the juxtaposition
of two or more elastic vesicles,
the
closing up or

opening

aperture which they form, according to circumstances, as is


manifest when the stomate is divided
perpendicularly to the
21 24 26
plane of the epidermis
.

51. Stomates are found


abundantly upon leaves, particularly
on the lower surface of those
organs; occasionally upon all

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

such as are of
parts that are modifications of leaves, especially
the
stem.
and
on
a leafy texture ;

upon the roots, nor on


submersed parts of plants,

52. Stomates have not been found


colourless parasitical plants, nor the

nor on Fungi, Algse, and Lichens

they

are,

moreover, rare,

or altogether absent, in succulent parts and in seeds.


53. It frequently happens, that they are so incompletely

formed, as to be either altogether incapable of action, or to


act in a very imperfect manner ; as in succulent plants.
54. The function of stomates is to regulate evaporation

and

has been thought, that the former funcwhich they are destined ; and,
that the cause of certain parts becoming succulent, is the
respiration.

It

tion, in particular, is that for

absence of stomates in sufficient numbers to carry off the


watery part of the sap. But some succulent plants have

more stomates than ordinary


requires reconsideration.
55. HAIRS are minute

plants,

so

that

this

opinion

expansions of transparent cellular

from the surface of plants. They are of


two kinds, lymphatic and secreting.
56. Lymphatic hairs are formed
by vesicles of cellular tissue placed end to end, and not
varying much in dimensions.
57. Glandular hairs are formed
by vesicles of cellular tissue placed end to end, and
sensibly distended at the apex or
tissue proceeding

base into receptacles of

fluid.

COMPOUND ORGANS.
58. Lymphatic hairs are for the absorption of moisture, for
the protection of the surface on which they are placed, and
for the control of evaporation through the stomates
(49).

They always proceed from

the veins, while the stomates oc-

cupy the interjacent parenchyma.


59. Glandular hairs are receptacles of the fluid peculiar to
certain species of plants, such as the fragrant volatile oil of the
sweet brier, and the acrid colourless fluid of the nettle, and

may

be regarded as organs of excretion.

Hairs are simple 35


ticulated 31 3a
liated 50

setaceous 28

27
;

septate

40 41
;

glochidiate

rumentaceous 48

capitate

armoniliform
;
;
strangulated
39
ciclavate 34 ; scabrous
;
;
43
36
45 40
araneose
stellate
scutate
;
;
;
;

38

knotted 33

branched 42

29

37
;

compound

49
.

60. Hairs are usually planted, more or less perpendicularly,


upon the surface on which they grow. In some cases, however, they are attached by their middle (peltate), as in Mal51
pighiaceous and Brassicaceous plants
.

RAPHIDES are crystals of any kind, usually acicular,


parenchyma.
61. PRICKLES are conical hairs of large size, sharp-pointed,
and having thin tissue very hard.

60

found

a.

in the interior of cells of

II.

COMPOUND ORGANS.

62. From peculiar combinations of the elementary organs


are formed the compound organs.

63.

The compound organs

are the axis (64) and

its

ap-

pendages (189).
64.

The Axis may be compared

to the vertebral

column of

animals.

formed from an embryo or leaf-bud, by the deof


a root in one direction, and of a stem in the
velopement
65. It

is

opposite direction.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

10

An embryo is a young plant, produced by the agency of


and
sexes,
developed within a seed.
67. A leaf-bud is a young plant, produced without the agency
of sexes, enclosed within rudimentary leaves called scales, and
66.

developed on a stem.
68.

An

70.

When

embryo propagates the species.


69. Leaf-buds propagate the individual.
the vital action of an embryo or bud

is

excited,

the tissue developes in three directions, upwards, downwards,

and

horizontally.
71. That part which developes downwards is called the
descending axis or root ; that upwards, the ascending axis or

that horizontally, the medullary system


and the part
;
from which these two axes start is called the crown or collar.

stem

72. This elongation in three directions takes place simultaneously ; hence it follows that all plants must necessarily

have an ascending and descending


and a medullary system.
73.

axis,

or a stem

and

root,

The only apparent exceptions

tribes of plants, in

to this are the lower


which the developement seems to be either

spherical, filamentous, or horizontal.


III.

74.

The

root

is

ROOT.

formed by the descending and dividing

fibres of the stem.

75. Anatomically it differs from the stem in the absence of


normal buds, and of stomates (49), and in Exogens of
pith.
76. Although the root has no distinct pith in
Exogens, yet
it
possesses a distinct medullary system.
77.

The

functions of the root are to fix plants in the


earth,

and to absorb nutriment from it. As it has to force


through substances which offer resistance to its
lengthens exclusively
of its divisions.

by

its

way

passage, it
successive additions to the
points

78. This absorption takes place almost


exclusively by the
extremities called spongelets, or
spongioles, which consist of a
lax coating of cellular tissue
lying upon a concentric layer of
woody tissue, in the midst of which is often placed a bundle
of ducts (31).
Spongioles are not, however, a distinct organ,
but are merely the young extremities of roots.

11

ROOT.

Roots are nodose 52


tubercular 56

53
;

placentiform

coralline

58
;

conical 54

tuberous 59

and

moniliform 55

fasciculate,

when

testiculate, or

in clusters as in

the Asphodel.

79. Occasionally the epidermis separates from the end of


the roots in the form of a cup or cap 57 , as in Pandanus and

Lycopodium.
80. The power of affording nutriment to the stem and other

by the root exclusively in consequence


The root is often a reservoir
the
soil.
from
absorption
of nutritious matter ready formed, and consisting of starch,
as in the Dahlia mucilage, as in the Orchis ; alkaline matter,
as in Rhubarb ; upon which the young stem feeds, even although the root itself is cut off from communication with any

parts, is not possessed

of

its

source of supply.
Moniliform, tuberous, testiculate, placentiin short, all which are unusually thickform, conical roots,
are intended by nature as reservoirs of food.
ened,
They

must not be confounded with tubers (152), rootstocks (152),


or corms (153), all which are forms of stem.
IV.

81.

The stem

is

STEM.

produced by the successive developement

of leaf-buds (164), which lengthen in opposite directions.


82. If an annular incision be made below a branch of an

Exogenous plant

(95),

the

upper

lip

of the

wound

heals

the part above the incision inrapidly, the lower lip


the
in
creases sensibly
diameter,
part below does not.

not

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

12

83. If a ligature be made round the bark, below a branch,


the part above the ligature swells, that below it does not
swell.

84. Therefore

the matter which

plants in

causes

the

increase

of

diameter descends.

Exogenous
85. If a growing branch

is

cut through below a leaf-bud,


between the section

that branch never increases in diameter


first bud below
The diameter of

and the

it.

Exogenous stems increases in each


number of leaf-buds developed.
species in proportion to the
of leaf-buds above a given part,
number
The
87.
greater
the greater the diameter of that part and vice versa.
86.

all

88. In the spring the newly forming wood is to be traced


in the form of organic fibres descending from the leaf-buds ;
is most newly formed lying on the outside, and
from
the most newly developed buds.
proceeding

that which

89. Therefore the descending matter, by successive addiwhich Exogenous plants increase in diameter, pro-

tions of

ceeds from the leaf-buds.


90. Their elongation upwards gives rise to new axes, with
appendages ; their elongation downwards increases the

their

diameter of that part of the axis which pre-existed, and produces roots.
91. Roots, therefore, in all cases, should consist of extensions of

woody

92. Hence,

tissue

and

this is

while the stem

is

evolution of leaf-buds, the root,


evolution, has no leaf-buds.
93.

The

conformable to observation.

formed by the successive


which is the effect of that

leaf-buds thus successively developed are


firmly
by the medullary system of the stem, which

held together

proceeds from the bark inwards, connecting the circumference


with the centre.
94.

The stem

95. It

is

side of the

varies in structure in four principal


ways.
formed by successive additions to the out-

either

wood, when

it

is

called Exogenous^ ; or
by sucis called
Endogenous^ ;

cessive additions to its centre, when it


or by the union of the bases of leaves,

and by addition to the


point of the axis, or by simple elongation or dilatation where
no leaves or buds exist ; this is called
Acrogenous.
96. In what are" called
Dictyogens^, the stem has the

STEM.

structure of Endogens, the root that of the stem of

Ex. Smilax.
97. The stem of EXOGENS

nearly

Exogens

may

Pith, the Medullary Sheath, the

Medullary Rays.
98. The PITH consists of

be distinguished into the


the Bark, and the

Wood,

cellular tissue,

occupying the cen-

tre of the stem.

99. It occasionally contains scattered spiral vessels, which

appear to originate in the medullary sheath (104), or scattered bundles of vascular and woody tissue, as in Ferula.
is produced by the elongation of the axis
upwards.
101. It serves to nourish the young buds until they have

100. It

acquired the power of procuring nourishment for themselves.


For this purpose it is filled with starch, which, in the process
of vegetation, becomes converted into mucilage
passes out of the pith into the nascent organs.

102. It
cases

when

it
it

is

always

solid

when

first

organized

and the

latter

but in some

separates into regular cavities, as in the Walnut,


is called disciform ; or it tears into irregular
spaces,

as in Umbelliferous plants.

of nourishing the young parts being acno further importance, and dies.
of
complished,
63
104. The MEDULLARY SHEATH consists of spiral vessels ".
103. Its

office
it is

105. It immediately surrounds the pith, projections of


which pass through it into the medullary rays (132) 63 *.
106. It is in direct communication with the leaf-buds and
the veins of the leaves.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

14

the de107. It carries upwards the oxygen liberated by


it into
conducts
and water, and
composition of carbonic acid
the leaves.
108.
sists

The WOOD

lies

and conupon the medullary sheath,

of concentric layers.

109. It is formed by the successive deposit of organized


matter descending from the buds, and by the interposition
of the medullary system, here called medullary rays, con63 b b

.
necting the pith and the bark
110. The first concentric layer

lies immediately upon the


medullary sheath and pith, and consists of woody and vasiform tissue 63 *.

Each succeeding

concentric layer consists <tf woody


and vasiform tissue, which either form themselves intoMistinct
strata, in which case the latter is innermost, or are con-

111.

founded together.
112.

When

there

is

any material

difference

between the

compactness of the tissue of the two sides of a concentric


layer, zones are formed in which the woody tissue is outerbut when the vasiform and woody tissues are equally
intermingled, no apparent zones exist.

most

113.

concentric layer, once formed, never alters in di-

mensions.
114.

Each

concentric layer,

which

is

distinctly limited, is

usually the produce of one year's growth.


115. Therefore, the age of an
Exogenous tree

should be

15

STEM.

known by

number of

the

But
numerous diswhich the period

concentric circles of the wood.

this rule is of uncertain application,

owing

to

turbing causes, especially in countries in


of rest is less distinctly marked than in the winter of northern
latitudes.

The

116.

secretions of plants are deposited

in the oldest concentric layers

most abundantly

while those layers which are

most recently formed contain but a

slight deposit.

When

117.

secretions,

it

the tissue of the concentric layers is


ceases to perform any vital functions.

The dead and

118.

fully

filled

with

formed central layers are called

the heart-wood.

The

119.

and incompletely formed external layers

living

are called the alburnum.

120.
like the

wood

Upon

the outside of the

wood,

consists of concentric layers.

lies

the BAKK, which,

66
121. It consists of four distinct parts
1. the Epidermis
";
65
65 66c
2. the EpipJilceum
3. the Mesophlceum
and 4,
;

the

65

Endophlmum or Liber
122. Each of these parts

to its

own

^d

increases

by

successive additions

except the epidermis, which

inside,

is

never re-

newed.
123.

The Epiphloeum and Mesophlosum

cellular tissue only

but their

rections with respect

large and

soft,

and

to each

may

cells are

other.

are both formed of

placed in different diThe former is often

separate spontaneously from the

young

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

16

layers forming beneath

it,

as in Cork, which

is

the epiphlceum

of Quercus Suber.
124. The Endophloeum or Liber consists of cellular tissue
on the alburnum, of laticiferous tissue (36), and of
resting

pleurenchyma (23). The tubes of the latter


ened rapidly by a deposit of sedimentary matter ; in which
case, sections of the tubes present the appearance of concentric
^ 65 66
Hence arises the toughness of the tubes of
circles
are often thick-

pleurenchyma which occur in the liber, and are manufactured


into cordage, as in the Lace-bark tree, the Lime-tree, &c.
125. Occasionally the liber is only formed during the first
year's

growth;

after

which

it

is

enclosed in wood, and

is

eventually found near the pith. This has as yet been observed
only in the Menispermaceous order.

The power of renewing themselves by the production


new matter upon their inner surface, is apparently given to
126.

of

the layers of bark in order to compensate for the gradual


incessant distension of the wood beneath them.

and

127. As the older parts die, from becoming too small to


bear the strain upon them, new parts form, each in its allotted
place, and take the station of that which went before it.
128.

bark

The

secretions of a plant are often deposited in the

any other part.


Hence chemical or medicinal

in preference to

129.

principles are often to be


sought in the bark rather than in the wood.
130. The immediate functions of the bark are to protect the
young wood from injury, and to serve as a filter through which

the descending elaborated juices of a plant


may pass horizontally into the stem, or downwards into the root.
131. It also contains the laticiferous vessels
(36), by which
is
conveyed to all parts of the surface of a plant.
132. The MEDULLARY RAYS or PLATES consist of

the latex

compressed

parallelograms of cellular tissue (muriform cellular tissue), belonging to the medullary system.
133. They connect together the tissue of the
trunk, maintaining a communication
ference.

134.

They

of the wood.

between the centre and the circum-

act as braces to the

woody and vasiform

tissue

They convey secreted matter horizontally from


the bark to the heart-wood, and
they generate adventitious
leaf-buds.

17

STEM.
135.

Cambium

a viscid secretion, which, in the spring,


Exogenous plant from the liber.

is

separates the alhurnum of an

It is free vegetable mucilage, out of which the new elementary


organs (8) are constructed, whether in the form of vessels, or
wr oody tissue, or of the cellular tissue of the medullary system,

whose office is to extend the medullary plates, and maintain the


communication between the bark and central part of a stem.
136. As Exogenous plants increase by annual addition of
new matter to their outside, and as their protecting integu-

capable of distension in any degree, commensurate with the increase of the wood that forms below it, it

ment or bark

is

follows, taking all circumstances into consideration, that there


are no assignable limits to the life of an Exogenous tree.
137. The stem of ENDOGENOUS plants offers no absolute dis-

Medullary Bays, Wood, and Bark.


formed by the intermixture of bundles of vascular
tissue among a mass of cellular tissue, the whole of which is
surrounded by a zone of cellular and woody tissue, inseparable
from the stem itself, and therefore not bark.
tinction of Pith,

138. It

is

139. It increases

by the successive descent of new bundles


down into the central cellular tissue,

of fibro- vascular tissue

curving outwards as they descend.


140. The vascular bundles of the centre gradually force outwards those which were first formed, the cellular mass augments
simultaneously, and in this way the diameter of a stem increases.
141. What appears to be bark in these plants is an external
layer of cellular tissue, into which the lower extremities of the
arcs of fibro-vascular tissue descend obliquely, losing their vascularity as soon as they reach the cortical integument, or falsebark.

142. It

is

an oblique
and the external cortical

in consequence of this continuity in

direction of the fibro-vascular bundles

integument, that the latter can never, in Endogens, be separated from the wood beneath it.
143.

The diameter

of the stem of an Endogenous plant is


its tissue possesses of distending, and

determined by the power

by

its

hardness.

144.

When

the external tissue has once

become indurated,

the stem can increase no further in diameter.


145.

When

the tissue

is

soft

and capable of continual

dis-

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

18

there is no more certain limit to the life of an


Endogenous than of an Exogenous tree.
146. Generally, the terminal bud only of Endogenous plants
tension,

is

developed

but very often a considerable number develope

Ex. Asparagus.
147.

When

a terminal bud only of an Endogenous plant

is
Ex. Palms when several
cylindrical
conical
Bamboo.
becomes
Ex.
;
develope,
148. In Acrogens no other stem is formed than what arises
from the simple union between the bases of the leaves and the
original axis of the bud from which they spring, and which
they carry up along with them.
149. In the order of Ferns the section of a stem indicates
the same structure as that of the numerous petioles (197) out
of which it is constituted.

developes, the stem

it

150. When Acrogens have no proper leaves, they are mere


expansions of cellular matter, sometimes in all directions ; Ex.

Fungi

sometimes in particular directions

Ex. Lichens,

Algse, &c.

151. The stem of a plant assumes numerous


ent appearances in different
plants.

and very

differ-

67
68
72
If above ground it is
;
rootshaped, or knotted
;
ascending
creeping
71
culated 73 ; leafless, succulent, and deformed 69 ; or
leafy .
If it bears the fmcers,
proceeding immediately from the soil or near it,
70

scape

10

arti-

it is

19

STEM.

152. It often burrows beneath the earth, when it is vulgarly


Sometimes the internodes become

called a creeping root'*.

much

thickened,

when what

or the stem lies prostrate


its

under

side,

when

it is

are called tubers 11

78

are formed

upon the earth, emitting roots from


called a rhizoma, or rootstock 67
.

distend underground, without creeping or rooting, but always retaining a round or oval figure, it is called a

153. If

it

75
.

154. All these forms of stem are vulgarly called roots.

155.

No

ments of

root can have either scales, which are the rudiwhich are the rudiments of buds.

leaves, or nodes,

scaly root

156.

is,

therefore,

a contradiction

The ascending axis,

in terms.

or stem, has nodes

and internodes.

157. Nodes are the places where the leaves are expanded
and the buds formed.
158. Internodes are the spaces between the nodes.
Whatever is produced by the evolution of a leaf-bud
(164) is a branch.

159.

160.

is the
imperfect evolution of a leaf-bud, and
a
branch.
is, therefore,
161. All processes of the stem which are not the evolutions
of leaf-buds, are mere dilatations of the cellular
integument of

the bark.

spine

Such are prickles (61).

162. In solid form the stem

lowing are

common terms

is

extremely variable; the

relating to

it

c 2

fol-

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

20

Terete 74

angled

half-terete 75

obtuse-angled

">

octangular
85
triquetrous

80

multangular

trilateral

Buds

are of

164. LEAF-BUDS

acute-

88

87

86

V.

163.

triangular

77

76

76

two-edged ;
piano-compressed
82
81
83
quinquangular ;
;
quadrangular
79
78
;
obscurely
quadriquetrous
triquetrous

compressed
81

78

quadrilateral

quinquelateral

LEAF-BUDS.

two kinds, Leaf-buds and Flower-buds.


(Bourgeon, Fr.)

consist

of rudimentary

leaves surrounding a growing vital point, the tissue of

which

capable of elongation, upwards in the form of stem, and


downwards in the form of root.

is

165. FLOWER-BUDS (Bouton, Fr.) consist of rudi-

mentary leaves surrounding a fixed

vital point,

and

assuming, when

fully developed, the form of floral


or
sexual
apparatus.
envelopes
166. Notwithstanding this difference, a leaf-bud

sometimes indicates a tendency to become a flower-

bud

and flower-buds frequently assume the chaEx. Monstrous Pears.


;
167. In appearance a leaf-bud seems 89 to be a
;

racters of leaf-buds

collection

of scales

above the other.


ary leaves.

The

arranged symmetrically one


scales are the rudiment-

These
centre

upon which they are placed

with a thin stratum


of spiral vessels, and these two
parts answer to
the pith (98) and the
medullary sheath (104) in

is

cellular substance coated

Exogens.
168. By the growth of a leaf-bud a branch is formed
and
;
the scales gradually change into true leaves as
vegetation ad-

vances 92

169. Sometimes they


separate spontaneously from the stem

21

LEAF-BUDS.
deciduous), and are
Ex. Lilium bulbiferum.
(are

then called

bulbills

or bulblets 9 *.

170. Often they are of large size, and are formed under9 91
;
they are then called bulbs or scaly bulbs

ground

171. Although the corm (153) is regarded as a kind of


underground stem, it may also be considered as a sort of leaf-

bud, the centre of which is very large and the scales very thin.
172. In bulbs, young buds or bulbs then called cloves
(nuclei),

Garlic

upon

are often formed in the axils of the scales, as in


and then gradually destroy the old bulb by feeding

In like manner corms produce other corms at the

it.

and are destroyed by their offspring.


some Gladioli 93 94 an old corm produces the

axils of their scales,

173. Thus

in

new one always


remains of

its

at

its

point

the latter

parent, and, being in like

is

then seated on the

manner devoured by

own offspring, becomes the base of the third generation 94 :


this process enables such plants by degrees to raise themselves

its

.out of the earth in which they were born.


174. In like manner the Crocus 75 produces

two or more

corms near the apex, and gradually dies as they develope ;


and the Colchicum bears its mother in the form of a shrivelled
96
spungy lump on one side of its base ", while on the opposite
966
side a new bud
is prepared by which the now vigorous
parent will hereafter perish.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

22

175. Both corms and bulbs are reservoirs of nutriment in


either a starchy or mucilaginous condition, or both.
176. Leaf-buds are of two kinds, the regular and the adventitious.

177. Regular or normal Leaf-buds are only found in the


axils of leaves.

178.

They

exist in a developed or

axils of all leaves,

179. Therefore

undeveloped state in the

and of all modifications of leaves.


they

may

be expected to appear at the

axils of scales of the bud, of stipules (223), of bracts (274), of

sepals (335), of petals (336), of stamens (348), and of carpels


(406) ; in all of which situations they are generally undeveloped,
for these different

180.

They

organs are

all

modifications of leaves.

are frequently not called into action, even in the

axils of leaves.

181. As regular buds are only found in the axils of leaves,


or of their modifications, and as branches are always the deit follows, that, whatever
may be the
arrangement of the leaves, the same will be the disposition of
the branches ; and vice versa.

velopement of buds,

182. This corresponding symmetry is, however, continually


destroyed by the unequal developement of the buds.
183. Leaf-buds which are formed

among

the tissue of plants

LEAF-BUDS.

23

subsequently to the developement of the stem and leaves, and


without reference to the latter, are called latent, adventitious,
or abnormal.

184. Adventitious Leaf-buds

be produced from any

may

part of the horizontal medullary system, or wherever cellular


tissue is present.
It has been distinctly proved, that, while
roots are prolongations of the vertical or woody system, leafbuds universally originate in the horizontal or cellular system.

185.

are formed'in the root,

They

among

the wood, and at

the margin or on the surface of leaves.


186. They are constructed anatomically exactly as regular
buds, having pith in their centre, surrounded by a medullary

sheath of spiral vessels,


cellular integument.

and coated over by woody

tissue

and

187. Hence, as adventitious buds, containing spiral vessels,


can be produced from parts such as the root or the wood, in
which no spiral vessels previously existed, it follows that this

form of tissue

is

by some other

either generated spontaneously, or


a manner unknown to us.

tissue, in

is

produced
most

It is

probable, that spiral vessels are spontaneous modifications of


vesicles of cellular tissue.

188.

Embryo buds

woody nodules found

are

in

the bark of

and apparently rudimentary branches formed without


leaves, within a space in which they are forcibly pressed upon
trees,

by the surrounding

tissue.

VI.

189.

leaf

is

LEAVES.

an expansion of the bark immediately below


is an appendage of the

the origin of a regular leaf-bud, and


axis (64).

Whenever a regular leaf-bud is formed, a leaf, either


or
perfect
rudimentary, is developed also ; and vice versa.
191. Leaves are developed alternately 97 , one above and op190.

posite the other, around their common axis


in consequence of the internodes

leaves

become opposite 98 or

produced

side

192. In

by

side,

Exogenous

but sometimes,

being unequally developed,


103
They are never

verticillate

except by irregular developement.


plants, the primordial or seed-leaves

(cotyledons) are opposite

hence, in .such plants the supposed

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

24

non-developement of the
formation of the embryo.

193. There

is

axis takes place during the original

a constant tendency in opposite or verticillate

become alternate.
194. This law applies equally

leaves to

to the

arrangement of

all

of leaves.
parts that are modifications
195. In leaves the developement takes place at their point
of junction with the stem ; and consequently the tissue at the
point of a leaf is the oldest.
196.

leaf consists of a petiole or stalk, a

lamina or blade,

and a pair of stipules.


197.

The PETIOLE

is

the channel through which the vessels

of the leaf are connected with those of the stem


of one or more bundles of spiral vessels and
enclosed in a cellular integument.

it is

woody

formed
tissue,

198. The spiral vessels of the leaf of Exogenous plants derive their origin from the medullary sheath ; those of En-

dogenous plants from the bundles of fibro-vascular tissue.


199. The cellular integument of the petiole is a continuation
of that of the bark.
200.
it is

and the lamina

is

abortive,

When the petiole becomes dilated and hollowed out at


upper end, the lamina being articulated with and closing

201
its

When

the petiole is leafy


called a phyttodium".
.

25

LEAVES.

up

as in Sarracenia

its orifice,

a pitcher or ascidium

if it

is

104

100

it is called
and Nepenthes
mere
a
is
and
sac, as
unclosed,
,

101

in Utricularia, it is called

ampulla
202. Sometimes the petiole has no lamina, or is lengthened
or taper
beyond the lamina, and retains its usual cylindrical
a
such
and
twists
becomes
but
;
petiole
spirally
long,
figure,
is

called

fewrfriJ (Vrille,

Fr.).
1

The petiole is usually either taper, or channelled ; and it has often a struma" ,
or apex, especially in those leaves which
(coussinet, Fr.) at either its base
107
103
In other cases it is inflated 106 , sheathing , ainplexicaul ,
are sensitive.
winged

203.

108

108
,

The

auriculate, leafless, jointed

petiole is

'

spinescent

10
,

&c.

sometimes articulated transversely as in

the Orange.
204. The

LAMIXA of a leaf is an expansion of the- parenof the petiole, and is traversed by veins which are ramifications or extensions of the bundles of vascular tissue of the

chyma

petiole, or,

when

there

is

no

petiole, of the stem.

205. Sometimes one, sometimes


leaf are furnished with stomates.
206.

The

both

the

surfaces of a

veins either branch in various directions

among

the parenchyma, anastomosing and forming a kind of network, or they run parallel to each other, being connected by
single transverse

uubranched

207. The former

is

of Endogenous plants.

veins.

characteristic of Exogenous, the latter

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

26
208.

The

petiole,

and

leaf is a continuation of the


principal vein of a
the base to the apex of
from
line
runs in a direct

It usually prothe lamina; this vein is called the midrib.


both such
from
or
or
base
from
its
veins
other
duces
sides,
and proceeding from the
veins are called ribs, if
:

very strong,

base to the apex; under other circumstances, they are fre-

quently named nervures.


209. There are two strata of veins, the one belonging to the
upper, and the other to the under surface.

The upper stratum conveys the

210.

juices from the stem


and elabo-

into the lamina, for the purpose of being aerated


rated ; the under returns them into the bark.

The

211.

veins are interposed

cellular

among

substance,

called diackyma, diploe^ or mesophyllum ; which is often stratified differently below the two surfaces of the leaf; the upper

stratum being more compact than the lower, and having its
in such cases, the
cells perpendicular to the plane of the leaf
:

cells

of the lower stratum are

commonly more or

less parallel

with the under surface.

The lamina is variously divided and formed; it is


thin
and membranous, with a distinct upper and under
usually
surface ; but sometimes becomes succulent, when the surfaces
212.

are often not distinguishable.


213. The upper surface is presented to the sky, the lower
to the earth ; this position is
rarely departed from in nature,
and cannot be altered artificially, except by violence.

leaf is simple when its lamina is undivided, or when,


separated into several divisions, those divisions do not
reach the midrib ; Ex. Lime-tree, Oak.
215. The form of the simple leaf is
extremely variable, and

214.

if it

is

the terms employed to denote the variations are numerous in


proportion.

216.
outline

Of such
forms

Some

leaves have the margin so continuous, that the

scarcely interrupted, except by small toothings.


leaves the following are among the more common

is

Orbicular 120

ovate 121

cordate" 9

124

peltate

oblique"

auriculate

122

113
oval 118 ; oblong 117 ; roundish
;
oblong
cordate ovate 114 ; cordate acuminate 119 ; renifonn 123

lanceolate
;

139
.

;
;

121

12:2

21 7. In other leaves the margin is produced here and there


in which cases the following terms are
;

into manifest angles


commonly in use
:

Sagittate or arrow-headed
128
.
lar 131 ; triangular

126
;

cuneate or wedge-shaped 127

218. In other cases the margin

a definite manner along


as the following are

its

is

hastate 130

angu-

repeatedly interrupted in
and then such terms
;

whole course

employed

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

28

Palmate 138

seven-lobed

133

134
;

pinnatifid

sinuated 132

panduriform

leaf is compound when the divisions pass down to


219.
the midrib so as to subdivide the leaf into smaller and distinct
leaves, or leaflets (foliola).

220.
is

When

expressed by
Ternate 144

leaves are compound, their


such terms as the following :

biternate 146 or triternate

terruptedly pinnate

147
;

lyrate

bijugate

140
;

digitate

143

148

conjugate-pinnate

150
;

bipinnate

mode

pedate

of division

142
;

decompound

pinnate

221. In speaking of the margin,

we say

that

145
;

in-

or tripinnate 141

149

it is

29

LEAVES.
Entire

126

serrate 114

tate 125

biserrate 137

129
crisp or curled

dentate 139

137

duplicato-dentate

tri-den-

crenate 120 .

222. The point of the leaves gives rise to other terms, such
as the following
:

Acute 118

obtuse 139
;
truncate 129 .

retuse

113

113
;

emarginate

acuminate 119

mucronate 132

223. STIPULES are attached to each side of the base of the


have, if leafy, veins, the anatomical structure
the same as that of the veins of the leaves.

petiole.

They

of which

is

224. Sometimes only one stipule

is

formed, the other being

constantly abortive, as in Azara.


225. Stipules are sometimes transformed into leaves

sometimes have buds in their axils

and may

they

be, therefore,

considered rudimentary leaves.


226. Whatever arises from the base of a petiole, or of a
leaf, if sessile, occupying the same place, and attached to each
side, is

considered a stipule.

The appearance

some being

so extremely variable,

large

and

of this organ is
leaflike, others

being mere rudiments of scales, that botanists are obliged to


define it by its position, and not by its organization.
227.

The

stipules

must not be confounded with

cellular

marginal appendages of the petiole, as in Apocynacese.


228. Stipules, the margins of which cohere in such a
that they form a
called ochreee

way

membranous tube sheathing the stem, are

Ex. Rhubarb.

229. All leaves are originally continuous with the stem^


as they grow, an interruption of their tissue at their junction

with the stem takes place, by which a more or


is formed sooner or later.

less

complete

articulation

The articulation between a leaf and stem being comthe


tissue of the former becomes gradually incrusted
pleted,
the
foreign matter deposited by the sap in the process of
by
230.

secretion
action,

and

digestion,

when -it

in diameter, as

dies.

and

a dead leaf

is incapable of further
the stem continues to increase

at last

When

will not increase

with

it,

the latter

But in
eventually thrown off; this is the fall of the leaf.
some Endogens the articulation is so slight, and the stem

is

increases so little in diameter, that the leaf

but simply withers and decays.

is

never thrown

off,

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

30

231. All leaves ultimately

fall

off;

evergreen leaves later

than others.

The mode

232.

bud

is

in

which leaves are arranged within their

called vernation, or gemmation.

the
have, under particular circumstances,
Ex.
from their margin (185) ;
power of producing leaf-buds
and proliferous Ferns.
Malaxis
paludosa,
Bryophyllum,
233. Leaves

FOOD AND SECRETIONS.

VII.

by the absorption of food from


consequence of which they grow, and

234. Plants are nourished

the air and earth, in

produce their peculiar secretions.


235. The growth of plants is very rapid ; that of the leaves
is such that they often acquire six or seven times their original
weight per hour.
236. The food of plants always consists of carbonic acid,
and water, and also of various mineral matters,

nitrogen,
chiefly

alkaline,

the

nature

of which

varies

according to

species.

237. Boots have the power of absorbing most substances in


a fluid or gaseous form, even although their extremities are
unbroken.
It appears probable that when plants are incapable of imbibing certain substances, such as strontian, there is no isomorphism between their ordinary

mineral constituents and those they reject. Thus, lime and magnesia, which
are isomorphous ; but between them and
plants will indifferently absorb,
strontian, which they will not absorb, no isomorphism exists.
Daubeny.
is obtained by plants in the form of carbonic
the atmosphere, or generated in soil by the
from
derived
acid,
of
vegetable matter.
decay

238. Carbon

239. Hydrogen is obtained principally by the decomposition


of water, and is assimilated along with carbonic acid, while
the oxygen of the water is liberated.
240. Nitrogen can only be obtained by plants in the form
The nitrogen of the atmosphere cannot be the
of ammonia.
source of supply, because it cannot be made to enter into combination with any element except oxygen, even by the employment of the most powerful chemical means.

241.

Ammonia exists

the stem, and in

all

in every part of plants, in the roots, in


blossoms and fruits in an unripe condition.

SI

FOOD AND SECRETIONS.


It is supplied

by

rain-water,

which

carries

it

down from

the

consequence of the putrefacThis ammonia affords


tion of animal and vegetable matters.
all vegetables, without exception, the nitrogen which enters
into the composition of their constituent substances.
air, in

which

it is

suspended, in

certain portion of the ammonia which falls with


242.
rain evaporates again with the water ; but another portion is
taken up by the roots of plants, and, entering into new combinations, produces albumen, gluten,

and a number of other

compounds, containing nitrogen.


243. But

it is

not so

much

the quantity of

ammonia

that

is

important to plants, as the form in which it is presented to


them.
in a volatile state, it is in great measure lost
before it can be imbibed.
fixed, in the state of salts,

When

When

overcome, and not the smallest portion of the


lost to the plants, for it is all dissolved by water

its volatility is

ammonia

is

and imbibed by the roots.


244. But carbonic acid, water, and ammonia, are not the
Ceronly elements necessary for the support of vegetables.
tain inorganic constituents are also essential.
245. Phosphate of magnesia in combination with ammonia
is

an invariable constituent of the seeds of all kinds of grasses.


acids found in the different families of plants are of vari-

The

ous kinds.

It

cannot be supposed that their presence and

peculiarity are the result of accident.

If these acids are con-

stantly present and necessary to life, it is equally certain that


some alkaline base is also indispensable, in order to enter into
combination with the acids, which are always found in the
state of salts.

246. If a plant does not produce more of its peculiar acids


it requires for its own existence, a
plant must contain an

than

invariable quantity of alkaline bases,

acids

may

form

wherewith the vegetable

salts.

The proportion of alkaline bases in a plant is indicated by the quantity of ashes they yield.
The quantity of
ashes obtained from the same quantity of vegetable matter
varies constantly in different species.
Therefore the propor247.

tion of alkaline bases varies in different species, and consequently different species demand a different amount of alkaline

food in the

soil.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

32

a plant is therefore deperfect developement of


or alkaline matter ; for
alkalies
of
on
the
presence
pendent
when these substances are totally wanting, growth will be
and when they are deficient, it must be impeded in
arrested
248.

The

proportion.

249. But other substances besides alkalies are required to


life of
Phosphoric acid has been found in
plants.

sustain the

plants hitherto examined ; and common salt,


sulphate of potash, nitre, salts of iron and copper, chloride of
potassium, and other matters, may be regarded as necessary

the ashes of

all

constituents of several plants.


is indispensable that every plant should
cultivated in those inorganic constituents
which nature has rendered necessary to it, just as it is neces-

250. Therefore

find in the soil

it

it is

sary for animals that they should find in their food the phosphates of lime and magnesia, which harden their bones.
251. As soon as food is absorbed, it begins to ascend into
the stem, or to diffuse itself through the system, and receives
the name of sap.
252. In the course of the sap upwards, the water and carbonic acid are partially decomposed and their elements are
deposited along with nitrogen in the interior of the tissue,
forming a layer over the interior of every cell and vessel,
which thus become in part solidified.

253. As soon as the sap reaches the leaves or the surface of


the bark, green matter, or
occasionally some other colour, is
formed, provided the part is exposed to light.
254. This appears to arise
chiefly from the decomposition of
carbonic acid, ammonia, and water, when the
carbon, nitrogen, and hydrogen are fixed by the plant, and the oxygen
restored to the atmosphere.
Such action is called the assimilating power of plants.
Plants are, therefore, the great purifiers of the
atmosphere, consuming the products of animal respiration and of all
organic putrefaction, and converting them
again into matter suited to the wants of man.

255. In the absence of


light, plants re-absorb oxygen from
the atmosphere, and re-combine it with the
matter they contain, to be again liberated at the return of
light.
256. They also, at all
times, especially at night, part with
carbonic acid in small
quantities.

FOOD AND SECRETIONS.


It has,

by

is

33

however, been proved experimentally that they purify the air much more
(254) action, than they vitiate it by their respiration.

their assimilating

257. No plants can long exist in which this alternate action


prevented, unless, perhaps, Fungi and brown parasites.
258. The amount of assimilation is determined by the de-

It is light alone
gree of light to which a plant is exposed.
that causes, in conjunction with vital forces, the decomposition
of the matters contained in living plants.

259. Hence, if a plant is compelled to grow in darkness, no


assimilation takes place of the food that the roots receive ;
oxygen accumulates ; its natural proportion to other elements

disarranged ; and a destruction of the tissue takes place.


260. In order to avoid this, plants will always lengthen
themselves in the direction in which the smallest ray of light
is

approaches them, as
ness into light.
etiolated,

and then

From

261.

the case of seed which shoot from dark-

is

If this

is

impossible, they

the continued assimilation

constituents of plants,

of the elementary
result, and serve for the
solid matters of a similar

new products

woody fibre, and all


The leaves produce

formation of

become blanched or

die.

sugar, starch, and acids,


which were preyiously formed by roots, when necessary for the
developement of the stem, buds, leaves, and branches.

composition.

Some phyto-chemists

believe that during the chemical transformations that result


in plants from the separation and re-combination of their elements, two compounds are necessarily formed, one of which remains as a component part,

while the other is separated by the roots, in the form of excrementitious matBut the experiments upon which this supposition is founded are not conand great doubt is entertained whether plants have really
the power of rejecting excrementitious matter by their roots.
It appears more
probable that the necessary separation of effete matter takes place by the
hairs and glands that clothe the surface of plants, or by a fluid secretion from
ter.

sidered conclusive

their whole surface.

262. Sap (251)

is put in motion
by the newly developing
by constantly consuming the sap that is near
them, attract it upwards from the roots as it is required.
Therefore, the movement of the sap is the effect, and not the

leaf-buds, which,

cause, of the

and

growth of plants.

It

depends upon

vital irritabi-

lity,
independent of mechanical causes.
263. This irritability is indicated not only by the motion
of the sap, but by several other
phenomena of vegetation ;
such as,

The

is

elasticity

with which the stamens sometimes spring up when touched, and

the sudden collapse of

many

leaves

when

stimulated

the apparently spon-

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

34

taneous oscillation of the labellum of some Orchidaceous plants ; the expansion


of flowers and leaves under the stimulus of light, and the collapse of them
when light is withdrawn (this phenomenon in leaves is called the sleep of
and
the effects of mineral and vegetable poisons being the same
;

by

plants)

upon plants as upon animals.


rosion

vegetable poisons

Mineral poisons

by the

kill

by inflammation and

cor-

destruction of irritability.

264. After the sap has been distributed through the veins
it becomes
exposed to the influence of air and

of the leaves,

and undergoes peculiar chemical changes.


called the proper juice.

light,
it is

265.

When

In this state

the proper juice has been once formed,

back, and descends towards the

it

flows

roots, passing off horizontally

into the centre of the stem.

266. Hence the great importance of leaves to plants, and


the necessity of exposing them to the full influence of light and
air, for the purpose of securing a due execution of their natu-

Hence also the impropriety of mutilating


by the destruction of their leaves.
267- In Exogenous plants (95), the upward course of the
fluids is through the young wood
their downward passage
through the bark, towards, or into the root and their horizonfunctions.

ral

plants

tal diffusion takes place

268.

Hence the

by the medullary

peculiar principles of such plants are, in


be sought either in the bark or the heart-

trees

and shrubs,

wood

(118), not in the alburnum (119).

to

rays.

But

in plants

whose

stems are annually


nial,

destroyed while the roots are perenthe latter are the chief reservoir of secretions ; and in

annuals, whose root

and stem both

perish, the secretions are

dispersed equally through the stem and root.


269. As they are the result of the

growth of a plant, they


be found more abundantly in annual
plants at the end
than at the commencement of their
growth.
will

270. In Endogenous plants (95) it is


probable that the upward course of the fluids is through the bundles of vascular
and woody tissue, and that the downward and horizontal

passage takes place through the cellular tissue.


271. The precise direction of the
sap in Acrogens (95) is

unknown.
VIII.

272.

The FLOWER-BUD

ed by imbricated,

FLOWER-BUD.

consists of a fixed
point, surround-

rudimentary,

or

metamorphosed

leaves,

FLOWER-BUD.

35

the external or inferior of which are usually alternate, and the


internal or superior always verticillate, or opposite ; the latter
are called floral envelopes

As

273.

and

sexes.

every flower-bud proceeds from the axil of a

either fully developed or


rudimentary,

exactly the same


bud.

leaf,

therefore occupies
position with respect to the leaf as a leafit

274. The leaf from the axil of which a flower-bud

arises, is

called bract or flower-leaf ; and all


rudimentary leaves, of
what size or colour soever, which appear on the peduncle (284)

between the

floral leaf

and the calyx (325), are

called Iracteola

or bractlets.

275. But, in common language, botanists constantly confound these two kinds, which are, nevertheless, essentially
distinct.

276. Although the buds in the axils of bracts are often not
developed, yet they have the same power of developement as
those in the axils of leaves ;
they are generally flower-buds,

very rarely leaf-buds.


277. When a single bract

is rolled together,
highly developed, and coloured, and is placed at the base of that form of
inflorescence called a spadix (304), it is named spathe ; Ex.

Arum.
278. When several bracts are verticillate or densely imbricated around the base of the forms of inflorescence, called the
umbel or capitulum (306), they receive the name of involucre ;

Ex. Carrot, Daisy.


279. When the bracts of an involucre form a single whorl,
and cohere by their margins, it is impossible to distinguish
them from the calyx by any other mark than by their position,
and by their usually surrounding more flowers than one.

The minute or colourless bracts at the base of the


of a capitulum (306) are called pale*.
281. Small imbricated bracts are often called scales.
280.

florets

when placed immediately below the sexes,


apetalous flowers, are only distinguished from the
calyx by being alternate with each other, and not verticillate ; hence the
glumes and paleee of grasses are bracts and
not calyx.
282. Bracts,

as

in

283.

The

axis of the flower-bud in its natural state does

D2

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

36
not

series of

lengthen beyond those upper

metamorphosed

leaves which constitute the sexes.

part of the axis, from the point of its


connection with the stem, as far as the floral envelopes, is
called the peduncle.

284.

The lengthened

285. When several peduncles spring from the axis at short


distances from each other, the axis receives the name of racMs,
and the peduncles themselves are called pedicels.

never more than one flower to each peduncle,


therefore, when we speak of a two-flowered
strictly speaking
we
peduncle,
only mean that two flowers, each having its
286. There

is

peculiar pedicel, terminate the axis,


peduncle common to each pedicel.

which

is

then considered a

287. Every flower, with its peduncle and bractlets, being


developement of a flower-bud, and flower-buds being

the

altogether analogous to leaf-buds, it follows, as a corollary,


that every flower, with its peduncle and bractlets, is a meta-

morphosed branch.
288. And further, the flowers being abortive branches,
whatever are the laws of the arrangement of branches with
respect to each other, the same will be the laws of the arrangement of flowers with respect to each other.
289. Flower-buds,

however, being

abortion than leaf-buds, flowers are

much

less

posed than branches, and appear to possess their


order of developement.
290.
bract,

As
it

subject

more symmetrically

own

to
dis-

peculiar

flower-buds can only develope from the axil of a


while a pedicel without bracts can

follows, that

never accidentally produce other flowers,


any one-flowered
pedicel, on which bracts are present, can, and frequently does,
bear several flowers.
291. In consequence of a flower and its
peduncle being a
in a particular state, the
rudimentary or metamor-

branch

phosed leaves which constitute bracts, floral envelopes, and


sexes, are subject to exactly the same laws of
arrangement as
regularly formed leaves.
292. The manner in which the floral
organs, especially the
calyx and corolla, are arranged before expansion takes place,
is called the Aestivation or
pra-flomtion.

FLOWER-BUD.

37

valvate 151 ; valvate and


following are the principal kinds of aestivation:
involute 156 ;
imbricate 157 ; alternate 159 16 ; convolute 152 ; induplicate 155 ;

The

153

plicative

293.

quincuncial

The modes

in

l54

157 l58
;

lei

supervolutive

vexillary

which the flower-buds are arranged are


which they un-

called forms of inflorescence ; and the order in


fold is called the order of expansion.

IX.

294. Inflorescence

is

INFLORESCENCE.
the ramification of that part of the

plant intended for reproduction by seed.


295. The greater developement of some forms of inflores-

cence than of others, is owing to the greater power one plant


possesses than another of developing buds, latent in the axils
of the bracts.

29 6. In consequence of flower-buds obeying the laws whieh


all forms of inflorescence must, of necessity,

regulate leaf-buds,

be axillary to a leaf of some kind.


297. Those forms which are called opposite the leaves, extraaxillary^ petiolar or epiphyllous, and even the terminal itself,
are

mere modifications of the

298.

axillary.

The kinds of inflorescence which

botanists

more

parti-

cularly distinguish are the following


299. When no elongation of the general axis of a plant takes
place beyond the developement of a flower-bud, the flower be:

comes what

is

called terminal

and

solitary

Ex. Pseony.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

38
300.

When

a single flower-bud unfolds

in the axil of

leaf,

and the general axis continues to lengthen, and the leaf underno sensible diminution of size, the flower which is devegoes

and axillary.
loped is said to be solitary
of
a newly formed elongated branch
301. If all the buds
at the same time produce peand
develope as flower-buds,
duncles, a raceme

is

formed 163

302. If buds, under the same circumstances, develope with1

out forming peduncles, a spike is produced


303. Hence the only difference between a spike and raceme
flowers are sessile, and in the latter
is, that in the former the
.

stalked.

304.
spadix differs from a spike in nothing more than in
the flowers being packed close together upon a succulent axis,

which

is

equal

enveloped in aspathe (277).


amentum is a spike the bracts of which are

An

305.

size,

and

closely imbricated,

and which

is

of

all

articulated

with the stem.


306.

When

tion of its

own

a bud produces flower-buds, with little elonga170 172


or an umbel 16 is
axis, either a capitulum
,
',

produced.
307. The capitulum bears the same relation to the umbel
as the spike to the raceme ; that is to say, these two forms
differ in the flower-buds of the
capitulum being sessile, and of
the umbel having pedicels.

39

INFLORESCENCE.
308.

The

dilated depressed axis of the capitulum

is

called

the receptacle.

raceme, or panicle, the lowest flowers of which


long pedicels, and the uppermost short ones, is a

309.

have

corymb
310.

l65 ier
.

A panicle is a raceme,

the flower-buds of which have,

in elongating, developed other flower-buds

173
.

middle branches of which are longer


than those of the base or apex, is called a thyrsus.
311.

312.

which

A
is

panicle, the

panicle, the elongation of all the ramifications of


arrested, so that it assumes the appearance of an

umbel, is called a cyme


313. In all modes of inflorescence which proceed from the
buds of a single branch, the axis of which is either elongated
or not, the flowers expand first at the base of the inflorescence,

and

last at

the summit.

This kind of expansion

is

called

centripetal.

314. When the uppermost or central flowers open first,


and those at the base or the circumference last, the expansion
is

called centrifugal.

315. The centripetal order of expansion always indicates


that the inflorescence proceeds from the developement of the
buds of a single branch.
316.

When

inflorescence

is

the result of the developement

of several branches, each particular branch follows the centri-

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

40

but the
petal law of expansion,

whole mass of inflorescence

the centrifugal.

317. This arises from the partial centripetal developement


the inflorescence,
commencing among the upper extremities of
instead of

among

the lower.

318. Consequently, this difference of expansion will indicate


whether a particular form of inflorescence proceeds from the
when it is called
developement of the buds of a single branch,

when

simple, or not,

319.

it is

called compound.

Whenever

the order of expansion is centripetal, the


to be understood as simple ; when it is centricompound, although in appearance simple. This

inflorescence

is

fugal, it is
difference is often of great importance.

320.

When

that the
either

the order of expansion is irregular, it indicates


of developement of the flowers is irregular also,

mode

on account of abortion or other causes.

321. Sometimes
abortive,

all

the flowers of the

and the ramifications, or the axis

twisted or spiral direction


formed; Ex. the Vine.

X.

when

this

inflorescence
itself,

are

assume a

happens, a tendril

is

FLORAL ENVELOPES.

322. The Floral Envelopes are the parts which immediately


surround the sexual organs.

323.

They

are formed of one or

more whorls of

bracts,

and

are therefore modified leaves (274).

324. In anatomical structure they do not


essentially differ
from the leaves, farther than is necessarily
consequent upon
the peculiar modifications of size or
developement to which
they are subject.

325.

When

the floral envelopes consist of but one whorl of

leaves, they are called calyx.

326.

When

two or more whorls are developed, the outer

called calyx, the inner corolla.

is

41

FLORAL ENVELOPES.
327. There

calyx and

is

corolla.

no other essential difference between the


Therefore, when a plant has but one floral

envelope, that one is calyx, whatever


degree of developement.

may

be

its

colour or

328. It is necessary, however, to be aware, that sometimes


the calyx is reduced to a mere rim, either in consequence
of lateral compression, as in the pappus (aigrette, Fr.) of many
Composite, or from other unknown causes, as in some
Acanthacese.
329. If the floral envelopes are of such a nature that it
not
obvious whether they consist of both calyx and corolla,
is
or of calyx only, they receive the name of periantMum or

perigonium.
330. Plants have frequently no floral envelopes
case flowers are said to be naked or achlamydeous.

in that

331. When the floral envelopes are deciduous, they fall from
the peduncle, as leaves from a branch, by means of an articulation ; if they are persistent, it is because no articulation
exists.

When the margins of floral envelopes are united, the


where
the union has taken place is called the tube, and
part
that where they are separate is named the limb.
It frethat
in
an
the
articulation
beforms
quently happens
calyx
tween the limb and the tube.
332.

333. Botanists generally consider that the tube of the calyx


invariably formed by the union of the margins of the sepals.
It is, however, probable that it is in some cases a mere dila-

is

tation

and expansion of the pedicel

334.

as in Eschscholtzia.

When

able from
rities

itself,

the calyx and corolla are readily distinguisheach other, they exhibit the following peculia-

335.

The calyx

consists of

two or more

divisions, usually

green, called sepals, which are either distinct, when a calyx is


said to be polysepalous ; or which unite by their
margins in a

greater or less degree, when


palous, or monophyttous.

it is

called monosepalous, gamose-

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

42

The calyx may be

178

superior

181
182
or inferior 177 ; galeate 176 ; calyptrate
; double
;
188
dilated 179 ; spiny 183 ; oblique 175 18S ; rin;

calcarate 186 ; coroniform 187 ; vesicate

gent

336.

184
.

The

two or more divisions, called


some bright colour, different from that of the
than which they are frequently more developed. When
corolla consists of

petals, usually of
sepals,

the petals are distinct, a corolla is said to he polypetalous ;


when they are united hy their margins, it is called gamopetah>iis or monopetalous.

The

corolla

panulate

may be

195
;

196
134
labiate 189 ; calceolate 193 ;
; papilionaceous
ringent
;
190
197
.
; crisp

funnel-shaped

cam-

43

FLORAL ENVELOPES.

or

337. If the union of the petals or sepals takes place in one


two parcels, the corolla or calyx are said to be one or two-

These

lipped.

lips

are always anterior and posterior with

respect to the axis of inflorescence,

and never

338. If the sepals or petals are of unequal

unequal degrees, the calyx or corolla

right
size,

and

left.

or unite in
1

is

said to be irregular ".

339. If the sepals and petals are unequal in number, or


no multiple of each other, or if the stamens are neither equal
to them in number, nor any power of them, a flower is said to
be unsymmetrical.

When

340.

the petals are so arranged, that of five the


dilated, the two lateral ones contracted and

uppermost is
parallel with each other, and the two lower also contracted,
parallel with each other, and coherent by their anterior mar19
gins, a flower is said to be papilionaceous *.

When

a petal tapers conspicuously towards the base,


1
said to be unguiculate ^ ; its lower part is called the
The former is analogous to the
unguis, its upper the limb.
341.

it

is

petiole, the latter to the

lamina of a

leaf.

The

petals always alternate with the sepals, a necessary consequence of their following the laws of developement

342.

of leaves.
343. If at any time the petals arise from before the -sepals,
is due to the abortion of one whorl of

such a circumstance

between the sepals and those petals which are actually

petals

developed.

344. As petals always alternate with sepals, the number of


each row of either will always be exactly the same. All
deviations from this law are either apparent only, in conse-

quence of partial cohesions,

or, if real,

are due to

partial

abortions.

Whatever

345.

intervenes

between the bracts and the

stamens belongs to the floral envelopes, and is either calyx or


corolla
of which nature are many of the organs
vulgarly
;

called nectaries.

Of

this nature are the horn-like bodies

found beneath the upper galeate sepal of

Aconitum 192, the cup of Narcissus 300, a part of the coronal appendages or
'"
net of
Stapelia

coro-

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

44

But it is to be observed, that as there are no exact


between the corolla and the stamens (348), such bodies
as have been just described are often of an indifferent nature,
and may be referred with equal justice to petals passing into
stamens, and to stamens passing into petals.
346.

limits

with the fringes of Parnassia 198 , some parts of the


is
particularly the case
coronet of Stapelia 203 , the long rays of the Passion-flower.

This

347. If, however, anomalous bodies at this part of the


vegetable system can be shown to belong to any whorl or
series of which a part is certainly petals or stamens, such ano-

malous bodies are to be regarded as belonging to the organ in


whose series they are placed.
in Aconitum 192 , the horn-like
processes belong to the series of the corolla,
and are therefore petals ; in the Ma-

Thus

202
and in the Canna 205 , they
,
evidently appertain to the Andro3ceum
This
(348), and are therefore stamens.
settles the true nature of what has been
called the nectary 206 , in Orchidaceous

hogany

plants,

now termed

the

which, forming a part


series of floral

or labellum,
of the second

lip,

envelopes, is therefore universally recognised as a petal, notwithstanding its singular form.

MALE ORGANS.
MALE ORGANS.

XI.

348.

45

The whorl of organs immediately within

the petals,

is

composed of bodies called stamens, which are considered the


male apparatus of plants, and constitute the Andrceceum.

They consist of a bundle of spiral vessels surrounded


cellular tissue, called t\i& filament, terminated by a peculiar

349.

by

arrangement of the cellular tissue, in a case, finally opening


and discharging its contents, called the anther.
350. There are many instances in which no limits can be
traced between the petals and stamens ; Ex. Nymphsea.
351. In such cases
contracts,

petal

it

is

found that the limb (341) of the

and becomes an anther, while the unguis

assumes the state of a filament.


352. Now as there are no limits between the petals and
sepals (327), nor between the sepals and bracts (323), nor
between the bracts and leaves (274), it follows that the sta-

mens are
and

also

a modification of leaves.

And

as the limb of a petal is analogous to the lamina,


the unguis (341) to the petiole of a leaf, it also follows

353.

that the anther

is

a modification of the lamina, and the

ment of the petiole.


354. The stamens follow the same laws of

fila-

successive de-

velopement as leaves ; and, consequently, if their arrangement


be normal, they will be either equal in number to the petals,

and alternate with them,

or, if

multiple of the petals.


355. If they are twice the

more numerous, some regular

number of petals, two whorls

are

considered to be developed ; and so on.


356. If they are equal in number to the petals, and opposite them, it is to be understood that the innermost
only of

two whorls

is developed, the outermost


being abortive.
357. All deviations from these laws are owing to the abor-

some part of the stamens

Ex. Lamium, Hippuris.


;
the stamens do not contract any union with
the sides of the calyx, they are hypogynous" ; Ex. Ranuntion of

358.

When

culus.

359.

When

they contract adhesion with the sides of the

177
calyx, they become perigynous ; Ex. Rose

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

46

of the calyx
360. If they are united both with the surface
Umbelliferse.
and of the ovary, they are epigynous ; Ex.
217
,
they are
361. When two are long and two are short
are
two
and if out of six
opposite ones

called didynamous ;
shorter than the other four, they are tetradynamous.
362. The filaments (349) are either distinct or united by
are united in one tube, they are called
If
their

margins.

216

monadelphous

they
Ex. Malva

if in

two

19
;

parcels, diadelphous"

13

Ex. Hypericum.
Ex. Pea if in several, polyadelphous"
the
a
solid
in
united
are
363. When they
body, along with
form what is called a column, and are said to be
;

style,

they

06

gynandrous"

Filaments are sometimes apparently forked 209 , in consequence of the separation


of the connective (366), into two arms ; strumose, when a tubercle forms upon
their face 220 ; stupose, if covered with long hairs 212 ; and toothed 214 , if their
214
margin is lengthened on either side beyond the attachment of the anther
.

364.

The

filament

The

anther~

is

not essential to a stamen, and

is

the limb of the stamen, forming within

is

often

absent.

365.

io

substance, and finally emitting a matter called pollen.


366. The two sides of the anther are called its lobes ; and
the solid substance which connects them, and which is in fact

its

a continuation of the filament, as the midrib of a leaf

is

of the

named the connective.


367. The connective is usually simple and uninterrupted
but it is sometimes lengthened into two arms 209 or is articupetiole, is

MALE ORGANS.

47

across which it is placed, and on


which it swings. In the latter case it either bears an anther213
210
or only on one
Ex, Salvia.
lobe on both arms
;

lated with the filament,

The

368.

cavities of the anther containing the pollen are

the cells, and the place by which the pollen is emitted is the
point or line of dehiscence ; the membranous sides of the
anther are named the valves.

369. Dehiscence usually takes place along a line, which


be considered to indicate the margin of the limb out of

may

which the anther is formed ; Ex. Rose.


370. Sometimes a portion only of this line opens, and then
the anther is said to dehisce by pores ; Ex. Azalea.
371. If the line of dehiscence occupies both margins of the
connective, and not the centre of the lobes, the anther opens
by one valve instead of two, which is then hinged by its upper

edge

Ex. Berberry.

372.

The

cells

of the anther are usually two in number

sometimes they are four 239 ; Ex. Tetratheca rarely one ; Ex.
223
Epacris and still more rarely several ; Ex. Viscum
373. The number of cells appears to be determined by no
:

certain rule.

374. Sometimes the


and become sinuous 237 ;
bristles 227

24
,

cells are

folded

down upon themselves

in other cases they are prolonged into


224
or tubes , or even into a spur 211 ; Ex. Mela-

stomacese.

375. Although in most cases the line of dehiscence is


with the anther-lobes, it is occasionally transverse 238

parallel

In Laurus the transverse and hinged (371) dehiscence being


combined 226 the face of the anther breaks up into four hinged
,

lobes.

376. It

an anther

may be
is

conjectured that the transverse dehiscence of


to the transverse articulation of

analogous

petioles (203).

377. The anthers frequently grow together by their marSuch anthers are called syngenesious.
gin ; Ex. Composite.
378. The Pollen is formed by a peculiar modification of the
cellules of the

parenchyma of the anther.

379. It consists of hojlow cases, of extreme smallness, containing a fluid in which float grains of starch and drops of oil.
380. It

is

furnished with apertures 229 , through which

its

STRUCTURAL AXD PHYSIOLOGICAL BOTANY.

48

227

226

225

224

223

222

form of a delicate tube,


lining is protruded in the
with
the stigma.
in
contact
comes
pollen

The shape of

381.

231

spherical
are common
,

pollen grains

the triangular
forms.

382. Its surface

is

228
,

either

is

where the

very variable

the

the polygonal" 22 , the oblong

smooth or studded with

230
,

little

231

points

383.

The

pollen grains are usually distinct from each other,

but in some cases they cohere in definite numbers ; Ex.


Acacia 232 or in irregular masses ; Ex. Orchidacese 23 * or are
:

enclosed within a bag, which seems to be the lining of the


anther (Endothecium) ; Ex. Asclepiadacese 233 23fi
384. In cases where the pollen grains cohere in masses, or
are enclosed within bags, they are connected with a cartilagi.

nous or elastic process, called the caudicle" 3*, which adheres


234
to a gland
belonging to the stigma.
385.

The

function of the

pollen

is

to vivify the

ovules

(396).
XII. DISK.

386.

Whatever

intervenes between the stamens

receives the general name of disk.


387. It usually consists of an annular

and the

pistil

passing the base of the ovary,

when

it is

elevation,

encom-

sometimes called the

cup , Ex. Pseony.


388. Or it appears in the form of a
glandular lining of the

MALE ORGANS.
tube of the calyx

Ex. Rose
Ex. Gesnera,
;

49

or of tooth-like,

hypogynous

Cruciferse.

(358) processes
389. When a fleshy substance occupies the
flower, and bears a single row of carpels, it
;

centre of a
is

called the

Ex. Lamium, Ochna, Geranium, &c. If this subgynobase


stance bears a greater number of carpels than can be arranged
in one row, it is called the torus or receptacle; Ex. Strawberry, Nelumbium.
390. It is certain that the disk is a non-developement of an
inner row or rows of stamens, as is proved by the Moutan
;

Paeony.
391.

The receptacle or torus is the growing point (164)


of the flower-bud in a state of enlargement.
392. The disk is one of the parts which Linnsean botanists
call nectary.

XIII.

393.

FEMALE ORGANS.

The organ which

occupies the centre of a flower,


within the stamens and disk, if the latter be present, is called
the pistil.

394. It

the female apparatus of flowering plants, or the

is
241

gynoeceum
395. It
style,

is

distinguished into three parts

viz.

the ovary, the

and the stigma.

396.
(445).

The OVARY

241

a hollow case, enclosing ovules


It contains one or more cavities, called
is

cells.

397.

The STIGMA

is

the upper extremity of the

pistil.

398. The STYLE is the part that connects the ovary


and stigma.
399. The style is frequently absent, and is no
more essential to a pistil than a petiole to a leaf,
or a filament to an anther.

400. Sometimes the style is thin,


and assumes the form of a petal, as in
401.

The

tinuous with

style
it.

is

flat,

and membranous,

Iris.

either articulated with the ovary, or con-

It usually proceeds directly from the


in some cases arises from the side, or

of the ovary ; but


the base of that organ

apex
even

Ex. Alchemilla, Chrysobalanacese.


E

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

50

the
properly speaking, stigma, except
the name is
the
of
Nevertheless,
style.
secreting
often inaccurately applied to mere divisions of the style, as in
Labiatse ; or to the hairy surface of undivided styles, as in
402. Nothing

is,

surface

Lathyrus.
403. Sometimes the stigmas grow to the face of the anthers,
204
which form themselves into a solid mass; Ex. Asclepias
.

In this case the styles remain separate.


404. The pistil is either the modification of a single
of one or more whorls of modified leaves,

leaf,

or

405. Such modified leaves are called carpels.


CARPEL is formed by a folded leaf, the upper sur406.
of
face
which is turned inwards, the lower outwards; and

within which are developed one or a greater number of buds,


which are the ovules.
407. When the carpels are stalked, they are said to be
seated upon a thecaphore, or gynophore ; Ex. Cleome, PassiTheir stalk is analogous to the petiole of a leaf.

flora.

408.

with

When

facility,

solid

the carpels are all distinct, or are separable


they are apocarpous ; when they all grow into

body, which cannot be separated into

parts, they are

its

constituent

syncarpom.

The ovary is the lamina of the leaf.


410. The style is an elongation of the midrib (208).
411. The stigma is the denuded, secreting, humid apex of
409.

the midrib.

412.

Where

the margins of a folded

leaf,

out of which the

formed, meet and unite, a developement of cellular


tissue sometimes takes
place, forming what is called the marcarpel

is

ginal placenta.
413. Every such
parts, one of

placenta is therefore composed of two


which belongs to one margin of the carpel, and

one to the other.


414. But although the
placenta of many plants appears to
derive its origin from the
margin of the carpels, it is certain
that in
other instances the
is a mere deve-

many

placenta

lopement of the centre of the flower-bud, and in reality the


end of the
Such a placenta is called
medullary system.
central.
It is not
impossible that even marginal placentae
and be
reality central.

may

be so in appearance only,

FEMALE ORGANS.

51

415. This law will explain the structure of some anomalous


pistils, in which the carpels are united into a confused mass ;

Ex. the Pomegranate 271

As

the carpels are modified leaves, they necessarily


of arrangement of leaves, and are therefore dethe
laws
obey
veloped round a common axis.

416.

417. And as they are leaves folded inwards, their margins


are necessarily turned towards the axis.
placenta, therefore, formed by the union of those margins, will be invariably

next the axis.


418. So that

if

a whorl of several carpels with a marginal

placentation unite and constitute a


pistil will be all in the axis.

pistil,

the placentae of that

The normal position of the carpels is alternate with


row of stamens, to which they are also equal
number but this symmetry of arrangement is constantly

419.

the innermost
in

destroyed by the abortion or non-developement of part of the


carpels.

The carpels often occupy several whorls, in which case


are
they
usually distinct from each other ; Ex. Eanunculus,
420.

272

Fragaria, Rubus
421. Sometimes,

notwithstanding their occupying more


than one whorl, they all unite in a single pistil ; Ex. NicoIn these cases the platiana multivalvis, Monstrous Citrons.
centse of the innermost whorl of carpels occupy the axis,
while those of the exterior carpels are united with the backs
of the inner ones, as must necessarily happen in consequence
of the invariable direction of the placentae towards the axis.

When

the carpels are arranged round a convex receptacle (389), the exterior ones will be lowest ; Ex. Ru-

422.

bus 272

423. But

if they occupy the surface of a tube, or are placed


upon a concave receptacle, the exterior ones will be uppermost Ex. Rosa 177
424. Whenever two carpels are developed, they are inThis
variably opposite each other, and never side by side.
happens in consequence of the law of alternate opposition of
;

leaves (191).

425. When carpels unite, those parts of their sides which


are contiguous gTow together, and form partitions between the
cavities of the carpels.
E 2

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

52

426. These partitions are called dissepiments.


427. Each dissepiment is therefore formed of two layers.
But these often grow together so intimately as to form but one
layer.

428. Such being the origin of the -dissepiments,

it

follows

that,

429. All dissepiments are vertical, and never horizontal.


430. They are uniformly equal in number to the carpels out
of which the

pistil is

formed.

single carpel can have no dissepiment whatever.


432. It will also be apparent, that as the stigma must bear
the same relation to the dissepiments as the point of the leaf
to the sides of the lamina, the stigma will always be alternate

431.

with (between) the dissepiments.


433. When the dissepiments of a many-celled pistil are
contracted so as not to separate the cavity into a number of
distinct cells, but merely project into a cavity, the placentae,

which occupy the edges of these dissepiments, become what is


r
called parietal ; Ex. Poppy ?
Occasionally the placentae are
diffused over the whole face of the
dissepiments, as in Bu.

tomus.
434.

one-celled ovary

may

also be

formed out of several

carpels, in consequence of the obliteration

Ex. Nut.

of dissepiments

FEMALE ORGANS.

53

of the foregoing diagrams explain these laws : a is a leaf ; b, a leaf rolled


to its conversion into a carpel ; c and k, a carpel ; d and I,
three carpels approximated, but not united ; e and m, the same united at the
and , these completely united into
ovaries, but disunited at the styles ;

Some

up preparatory

one ovary, one

style,

and one stigma.

435. All dissepiments whose position is at variance with


the foregoing laws are spurious.
436. Spurious dissepiments derive their origin from various
causes,

and may have

either a vertical or horizontal position.

When

they are horizontal they are called phmgmata,


are formed by the distension of the lining of the ovary ;

437.

and
Ex. Cathartocarpus,

Fistula.

438. If vertical, they either are projections from the back


of the carpel, as in Amelanchier and Thespesia* ; or they are

caused by modifications of the placentae, as in Martynia, Didymocarpus, and Cruciferae ; or they are produced by the turning inwards of the margins of the carpels*.
244
, consisting of five cavities in the axis, sursingular fruit of Diplophractum
five two-celled cavities at the circumference, must be composed of
carpels constructed as just described, and arranged in several series (420).

The

rounded by

244
explained by the following cut, where

is a section of the fruit of


shows an ideal arrangement of fifteen carpels in three
rows, five being external and perfect, with the margins of the carpels turned
inwards (406) ; five being altogether imperfect, and the five in the centre
243
shows the transverse section of this ideal figure.
being less imperfect.
In the ripe fruit we must suppose the intermediate carpels to be obliterated,
and the spurious dissepiments of the external carpels to be pressed up against

This

is

Diplophractum

242

their back, so as to bisect the cavity of each carpel.

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

54

439. Sometimes the central placenta extends beyond the


base of the carpels, rising up between them, and either forming an adhesion with the styles, as in Geranium, or a central
distinct axis, as in

Euphorbia.

440. This elongation of the placenta is more apparent in


the fruit than in the pistil.
It is analogous to the cellular

apex of the spadix (304) of Arum.

The styles of different carpels frequently grow tointo


a solid cylinder'; Ex. Lilium. There are various
gether
441.

degrees of union between the styles.


442. The style is incorrectly said to be divided in different

ways, in consequence of this adhesion.


443. If the ovary adheres to the sides of the calyx
called inferior, and the calyx is said to be
superior ;

it

is

Ex.

Apple.
444. If
it is

it

contracts no adhesion with the sides of the


calyx
and the calyx inferior.

called superior,

XIV. OVULE.

The OVULE is a body borne by the placenta


(412),
and destined to become a seed (531).
446. It is to the carpel (406) what the
marginal buds are
to leaves
(185), and to the central placenta what buds are to
445.

branches.

447. It does not, however,


appear to bear any other
analogy to a bud than what is indicated by its position.
448. The ovule is
usually enclosed within an ovary (396) ;
but in Coniferse and
Cycadaceae it is destitute of any covering,
and is exposed, naked, to the influence of the
pollen.
449. It is either sessile, or attached
by a little stalk called
the funiculus, or
The point of union of the funipodosperm.
culus and ovule is the base of the
latter, and the opposite extremity is its apex.
450. It consists of two
sacs, one enclosed within the other,
and of a nucleus within the sacs.
451. These sacs are called the
primine and secundine.

452.

The

primine, secundine,

and nucleus, are

with each other


by a perfect continuity of

point of their surface.

453.

When

all

tissue,

connected
at

some

the parts of the ovule


undergo no alteration of

55

OVULE.

position during their growth, thfe two sacs and the nucleus are
connected at the base (449) of the ovule, which is orthotropous or atropous.
all

454.

And

then the base of the nucleus and that of the

ovule are in immediate connection with each other.

455. But the relative position of the sacs and the base of
the ovule are often entirely altered during the growth of the
latter, so that it frequently happens that the point of union of

the sacs and the nucleus

456.

And

is at the apex (449) of the ovule.


then the base of the nucleus is at the apex of the

ovule.

457. In such cases, a vascular connection is maintained between the base of the ovule and the base of the nucleus, by

means of a bundle of vessels called a raphe.


458. The normal position of this raphe is on the

side of the

ovule, next the placenta.

459. The expansion of the raphe, where it communicates


with the base of the nucleus, gives rise to the part of the seed
called the chalaza (548).

460.

When

the ovule

proach the placenta,

it is

curved downwards so as to apcampylotropous ; when curved downis

wards and grown to the lower half, anatropous ; when attached by its middle so that the foramen is at one end and the
base at the other, it is amphitropous.
461. The mouths of the primine and secundine usually contract into a small aperture called the foramen of the ovule, or
the exostome.
462.

The apex of the nucleus

is

always applied to

this

foramen.
463. In consequence of the relation the base of the nucleus
bears to the base of the ovule, the foramen will be at the
apex of the ovule when the two bases correspond, and at the
base

of the ovule

when

the two

bases

are

diametrically

opposite.

464.

The foramen

indicates the future position of the radi-

of the embryo (555) ; the radicle being always next the


foramen.
This is a fact of great importance in practical
cle

Botany.
465. Within the nucleus

is

a cavity or bag, called the sac

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

56
of

tke

among

named the liquor amnios,


amnios, containing a fluid
which the emhryo is developed.
XV. IMPREGNATION.

466. Impregnation is effected


pollen (378) and the stigma (397).

by contact

between the

ci

emits a tube" of extreme delicacy,


pollen
which pierces the stigma 6 and style 51, and, passing downwards
l
/A enters the foramen
of the ovule"
into the
467.

The

(461)
ovary ,
468. Having reached the foramen, it comes into contact
with the nucleus (450).
.

469. This accomplished, the act of impregnation is over;


a new body gradually appears in the sac of the amnios (465),

and eventually becomes an embryo.


470. Great numbers of modifications of this phenomenon
have been observed, but they^ all resolve themselves into these
facts.

471. In plants, the ovules of which have no pericarpial


covering, such as Cycadaceae and Coniferse, (gymnosperms,)
the pollen falls in the foramen, and there acts as" if it had
struck the stigma.
472. If only one pollen tube enters an ovule, there is but
one embryo in the seed.
But if several pollen tubes pass into
the same ovule, there

seed

may

Ex. Onion, Miseltoe.

be several embryos in the same

57

FRUIT.

XVI. FRUIT.

The

473.
pistil

FRUIT, in the strictest sense of the word, is the

But the term

arrived at maturity.

the

pistil

are

all

and

is

also applied to

envelopes taken together, whenever they


one uniform mass.

floral

united in

474. Hence, whatever is the structure of the pistil, the


same should be the structure of the fruit.
475. But in the course of the advance of the pistil towards

many

maturity,

take

alterations

in

place,

consequence of

and union of

abortion, non-developement, obliteration,


476. Whenever the fruit contains anything

at

parts.

variance

with the laws that govern the structure of the pistil, the latter
should be examined for the purpose of elucidation.
477. Sometimes a pistil with several cells produces a fruit

with but one Ex. the Hazel-nut and Cocoa-nut. This arises
from the obliteration of part of the cells.
478. Or a pistil, consisting of one or two cells, changes to
a fruit having several: the cause of this is a division and
;

doubling of the placentary divisions ; Ex. Martynia or the


expansion of portions of the interior ; Ex. Cathartocarpus,
:

Fistula.

479.
indicate

As

the fruit

upon

its

is

the maturation of the

some

surface

pistil,

traces of a style

true in all cases, except Cycadacese

and

Coniferse,

it

ought to

and this is
which have

no ovary.
480. Hence the grains of corn, and

many

other bodies that

resemble seeds, having traces of the remains of a style, cannot


be seeds, but are minute fruits.
481. That part which was the ovary in the pistil, becomes
the pericarp in the fruit.
482. The PERICARP consists of three parts ; the outer coating called the epicarp, the inner lining called the endocarp, or

putamen, and the intermediate substance named the sarcocarp.


483. Sometimes these three parts are

guished ; Ex. the Peach


substance ; Ex. a Nut.
484.

The

all readily distinfrequently they form one uniform

is the part where it is joined to


where the remains of the style are

base of the fruit

the peduncle.
found.

The apex

is

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

58
485.

The

axis of the fruit

is

often called the columella

the space where two carpels unite is named the commissure.


486. All fruits which are mere modifications of a single
carpellary leaf (406) have always a suture corresponding with
the junction of the margins, or with the placenta?, and often
another corresponding with the midrib of the carpellary leaf
:

the former

is

called the ventral, the latter the dorsal suture.

is

487. If the pericarp neither splits nor opens when ripe, it


if it does split or open, it is said to
,

said to be indehiscent

dehisce, or to

be dehiscent

and the pieces into which

it

splits

are called the valves.

488.

The dehiscence of the

pericarp takes place in differ-

ent ways.
489. If

it takes
place longitudinally, or vertically, so that
the line of dehiscence corresponds with the junction of the
carpels, the dissepiments are divided, the cells remain closed

at the back,

dodendron 264

and the

dehiscence

is

called septicidal

Ex. Rho-

490. Formerly, botanists said that in this kind of dehis; or, that

cence the valves were alternate with the dissepiment


the valves had their margins turned inwards.

491. If it takes place vertically, so that the line of dehiscence corresponds- with the dorsal suture
(486), the dissepiments remain united, the cells are opened at their back, and
the dehiscence

is

called loculicidal

Ex. Lilac, Lily.

492. Formerly, it was said that in this kind of dehiscence


the dissepiments were
opposite the valves.
493. When a separation in the
pericarp takes place across
the cells
horizontally, the dehiscence is transverse ; Ex. Anagallis.

494. If the dehiscence is effected


by partial openings of
the pericarp, it is said to take
place by pores ; Ex. Poppy.
495. Sometimes the cells remain
closed, separating from
the axis formed
by the extension of the peduncle
Ex.
;
Umbelliferae, Euphorbia

(284)

255
.

496. Or the cells


open and separate from the axis, which
formed by a cohesion of the
placentse which separate from
the dissepiments ; Ex. Rhododendron 264
is

497. Sometimes the


dissepiments cohere at the axis, and
separate from the valves (487) or back of the
carpels; Ex.
Convolvulus.

FRUIT.

59

498. All fruits are either simple or multiple.


499. Simple

proceed from a single flower

fruits

Ex.

Poeony, Apple, Nut, Strawberry.


500. Multiple fruits are formed out of several flowers D F ;
Ex. Fir, Pine-apple, Fig.
They are masses of inflorescence

and are also called anthocarpous.


501. Simple fruits are either the maturation of a single
carpel (406), or of a pistil formed by the union of several
in a state of adhesion,

carpels (408).

Of fruits formed

502.

ant

are

the

Achenium

503.

of a single carpel, the most import-

Follicle

(503), Legume (504), Drupe (507),


(508), Caryopsis (511), and Utricle (512).

The

Follicle

a carpel

is

dehiscing

by the

ventral

and having no dorsal suture 260


504. The Legume is a carpel having both a ventral and
E 258
dorsal suture, and dehiscing by both, either, or neither 263
505. The two sutures of a legume sometimes form what is
called a replum ; Ex. Carmichselia.
506. When articulations take place across the legume, and
suture,

it falls

507.
cent,

into several pieces,

The Drupe

and

differs

in its pericarp

it is

said to be lomentaceous" 51

from the

follicle in

262
.

being indehis-

having a distinct separation of epicarp


59

(482), sarcocarp, and endocarp'


508. The Achenium is an indehiscent,
.

bony, one-seeded

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

60

pericarp,

which does not contract any degree of adhesion with


251

249
the integument of the seed
a
the
It
is
509.
drupe,
pericarp of which does not separate into three layers.
.

is pappose when it bears the remains of a calyx at its apex ; Ex.


245
46 while the
245
, or rostrate*
,
,
pappus is setaceous
Compositae : and is truncate
If the style remains and becomes
double 245 , plumose 246 , or paleaceous 251 .
249
kind
of
is
caudate
.
a
the
achenium
tail,
feathery, forming

The Achenium

achenium is elevated on a large


Anacardium 250

510. Occasionally the


fleshy receptacle, as in

511.

The Caryopsis

is

an indehiscent, membranous, one-

seeded pericarp, which adheres firmly to the integument of


the seed ; Ex. Corn.
512. The Utricle is a caryopsis, the pericarp of which has
no adhesion with the integuments of the seed ; Ex. Eleusine,
Ohenopodium.
513. Of fruit formed of several carpels, the principal are

Samara (517), Cremocarp


Nuculanium
(518),
(519), Siliqua (515), Nut or Gland (517),
(522),
Berry
Orange (523), Pome (524), Pepo (525), and

the Capsule (514), Pyxis (520),

Balausta (526).
514. The Capsule
carp

is

a many-celled, dry, dehiscent peri-

253 256 264 269

It i* stellate 256 , toothed at the


265
or spiral"*
,
apex
alter separation 268 ,
they are named cocci.

if its

cells

remain close

61

FRUIT.

515. The Siliqua consists of two carpels fastened together,


the placentae of which are parietal, and separate from the
valves, remaining in the form of a replum (505), and con266

nected by a membranous expansion


516. When the siliqua is very short, or broader than
.

long,

it is

it is

called a Silicula.

The Nut

Gland

is a dry, bony, indehiscent, onefrom


a pistil of three cells, and enproceeding
closed in an involucre called a Cupule ; Ex. the Hazel, Acorn.
It is a sort of compound achenium.

517.

or

celled fruit,

In some Palms, Ex. Sagus, it is covered by scales turned downwards 273 . It is


often bordered by expansions or wings which surround it longitudinally, as
261
or proceed from the apex or
in the Elm 247 ; or transversely, as in Paliurus
;
back only, as in Sycamore 252, in which case it receives the name of Samara.

518. The Cremocarp is a pair of Achenia, then called


mericarps, placed face to face, and separating from a central
55
Their planes of union constitute
axis ; Ex. Umbelliferar .
the commissure.

The Nuculanium is a capsule, which, being fleshy,


Ex. Grape, Arbutus 570
520. The Pyxis is a capsule whose dehiscence takes place

519.

does not dehisce

253 254

transversely

521.

Ex. Hyoscyamus, Anagallis.

The Efario

is

a collection of

distinct,

indehiscent

272
carpels, fleshy or dry, within a calyx ; Ex. Rubus
522. The Berry is a succulent fruit, the seeds of which lose
.

their adhesion

berry.

when

ripe,

and

lie

loose in pulp

Ex. a Goose-

62

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

The Orange is a berry having a pericarp separable


an epicarp, an endocarp, and a sarcocarp, and the cells
filled with pulpy bags, which are cellular extensions of the
523.

into

sides of the cavity.

524.

The Pome

is

a union of two or more inferior carpels,

the pericarp being fleshy, and formed of the floral envelope


and ovary firmly united 274
.

525.

The Pepo

526.

The Balausta

composed of about three carpels, forming


a three-celled, fleshy, indehiscent fruit, with parietal placentae ;
Ex. Cucumber.
is

is

a many-celled

fruit,

with the seeds

63

FRUIT.

arranged in an irregular manner on the backs of the cells, and


is formed
by more whorls of carpels than one, enclosed within
a tough rind ; Ex. Pomegranate 271 .
527. The most remarkable modifications of multiple or
anthocarpous fruits are, the Cone (528), Pine-apple (529),

and Fig (530).


528.
Pinus.
cohere,

529.
all

When
it is

A B

an indurated amentum (305)


Ex.
in size, and its scales
firmly
called a Galbulus ; Ex. Thuja.

The Cone

it is

is

The Pine-apple

grow together
530. The Fig

much reduced
is

a spike of inferior flowers, which

into a fleshy mass.

is the
fleshy, hollow, dilated apex of a
peduncle, within which a number of flowers are arranged,
each of which contains an achenium ; Ex. Ficus, Dorstenia 275
.

Of

the terms above explained only a few are in common use, and it seenis to
be found by systematic botanists more convenient to describe a given fruit by
exact words than to use any particular term.
The names most employed are

the Achenium, Nut, Caryopsis, Drupe, Capsule, Siliqua, Legume, and Cone.

XVII. SEED.

The SEED is the ovule (406) arrived at maturity.


532. It consists of integuments (540), albumen (551), and
embryo (555) ; and is the result of the reciprocal action of
531.

the sexual apparatus.


533. In general, seeds are, like ovules, enclosed within a

covering arising from a carpellary leaf (406) ; but all Gymnosperms are an exception to this. Moreover, some ovules

rupture the ovary soon after they begin to advance towards


the state of seed, and thus become naked seeds ; Ex. Leontice.

Others are imperfectly protected by the ovary, the carpels


not being perfectly closed up ; Ex. Reseda.
534. The seed proceeds from the placenta (412), to which
attached by the funiculus 280 , which is sometimes very
long, but is more frequently not distinguishable from the

it

is

placenta.

535. Sometimes the funiculus, or the placenta, expands


about the seed into a fleshy body ; Ex. the Mace of a nut76 281 283
meg, Euonymus. This expansion is named aril*
.

never developed until after the vivification of the


ovule, and must not be confounded with tumours or dilatations of the integument of the seed.
536. It

is

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

537. Sometimes there are tumours of the testa near the


hilum or at the opposite end such are called StropMolte or
Caruncula
538. The precise nature of these is unknown ; sometimes
;

or they are
they are dilatations of the chalaza ; Ex. Crocus
caused by a fungous state of the lips of the foramen ; Ex.
:

Eicinus
539.

or they arise from unknown causes.


scar, which indicates the union of the seed with

The

the placenta, is called the hilum or umbilicus


540. The integuments are called collectively testa, and
consist of membranes resulting from the sacs of the ovule
.

(451).
541. Sometimes the testa

whole surface

sions of its

is

covered by hair-like expan-

as in the Cotton

or these hairs

occupy one or both ends, when they constitute what is called


the coma'287
This must not be confounded with pappus (328),
which is calyx.
.

542.

The integuments

which are either

284

single

are

often

expanded into wings,


and appear intended

or several 277

render seeds buoyant.


Very often they are corky or
282
and
not
,
spongy
unfrequently consist of spiral cells (19).

to

543. In the seed these membranes are called


by various
names, of whiph the most frequently used are spermoderm or
testa for the primine ;
mesosperm, for the secundine ; and endopleura for the coat of the nucleus (450).

65

SEED.

The mouth of the foramen (461) is often distinctly


is named the micropyle ; Ex. Pea.
545. The raphe 279 285 occupies one side of the seed in all
544.

visible,

and

cases in which

becomes much
546.

pre-existed in the primine


ramified.
it

The raphe

in

is

no

way

but

it

frequently

connected with impregnation

being apparently confined to maintaining a


vascular connection between the placenta and the base of the
nucleus, for the purpose of nourishing the latter.
functions

its

547. Spiral vessels are found in the raphe and

its

ramifica-

tions.

548.

Where vessels of the

raphe expand into the mesosperm

(543), the chalaza (459) appears as a discoloured thickening


of the integuments 285 .

549. The micropyle always indicates the point in the circumference of a seed towards which the radicle (561) points.
550. And the chalaza is as constant an indication, when it
present, of the situation of the cotyledons (559) ; it being
always at that part of the circumference organically opposed
to the radicle.
is

551. Between the integuments and the embryo of some


1 * 293
plants lies a substance called the albumen or perisperm""
299,

552. It consists of a peculiar matter deposited during the


growth of the ovule among the celullar tissue of the nucleus
(450).
553.

When

the cellular tissue of the nucleus combines with

the deposited matter so completely as to form together but


one substance, the albumen is called solid ; Ex. Wheat,

Euphorbia.

When

verted, the

albumen

554.

Albumen

a portion of the tissue remains unconruminated ; Ex. Anona, Nutmeg.


usually wholesome, and may be fre-

is

is

quently eaten with impunity in the most dangerous tribes


Ex. Omphalococca, a genus of Euphorbiacese.

555. The organised body that lies within the seed, and for
the purpose of protecting and nourishing which the seed was
8
created, is the Embryo"*
.

556.

The embryo was

the amnios (465).


557. The latter

is

originally included within the sac of

usually absorbed or obliterated during

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

66

the advance of the

embryo

to maturity

but

it

sometimes

form of Vitellus
remains surrounding the ripe embryo, in the

Ex. Saururus, Piper

The embryo

consists of the

cotyledons (559),
collar (562).
the
and
the
radicle (561),
plumule (560),
558.

297 .-

the

The cotyledons represent undeveloped leaves 295 ".


560. The plumule, or gemmule, is the nascent ascending
559.

axis (64) 2956

561.

The

radicle

is

the rudiment of the descending axis

295c

(7l)
562.

The

collar is the line of separation

between the radicle

and the cotyledons.


563. The space that intervenes between the collar and the
base of the cotyledons is called the cauliculus. (Tigelle, Fr.)
564. In some seeds the embryo is furnished with a
300

suspensor from the point of the radicle


565. The embryo is usually solitary in the seed, but occasionally there are two or several (472).
.

When

several embryos are produced within a single


sometimes
seed,
happens that two of these embryos grow
in'
which
case a production analogous to animal
together,
monsters
is formed.
dicephalous

566.

it

In form, position, and direction, the embryo varies in different species. In


291
in others heliacal 294
in others
general it is straight ; in some it is spiral
;
;

vermicular 290

in others arcuate 301 .

It usually occupies the axis of the


:
but it is also excentrical 292 ; and unilateral 299 . In
either erect with
respect to the seed, or inverted or transverse.

albumen or seed 278


direction,

it is

293

SEED.

67

567. The number of cotyledons varies from one to several.


The most common number is either one or two. In the latter
case,

they are always directly opposite each other.

The cotyledons

are semiterete 895 "

foliaceous 278

flat,

convolute 288

with each other, or divergent 300 . When there is but one cotyledon,
assumes peculiar forms: it is, for instance, fungous 289 ; spheroidal 298

parallel
it

often

lenti-

cular 299 a.

The direction of the embryo, with respect to the seed,


depend upon the relation that the integuments, the raphe,
chalaza, hilum, and micropyle, bear to each other.
569. If the nucleus be inverted, the embryo will be erect,
or orthotropous ; Ex. Apple.
570. If the nucleus be erect, the embryo will be inverted,
or antitropous ; Ex. Nettle.
571. If the micropyle is at neither end of the seed, the em568.

will

will be neither erect nor inverted, but will be in a more


or less oblique direction with respect to the seed ; Ex. Primrose ; and is said to be heterotropous.

bryo

572. Plants that have but one cotyledon, or, if two, with
the cotyledons alternate with each other, are called MONO2
".
COTYLEDONOUS 293

two opposite each other, or a greater


a whorl, are called DICOTYLEDONOUS 288 29 C92

573. Plants that have

number

placed in

297 300 301

574. Endogenous plants are monocotyledonous.


575. Exogenous plants are dicotyledonous.
576. Plants that have no cotyledons are said to be ACOTYLE-

DONOUS 294

577. But

term is usually applied only to cellular


plants which, having no sexual apparatus, can have no seeds
this

(587).
578. Acrogenous plants are acotyledonous.
579. Those seeds of flowering plants, which appear to have
no cotyledons, owe their appearance to the cotyledons being
consolidated

Ex. Lecythis, Olynthia

or abortive

Ex. Cus-

cuta.

580.
action

The plumule is very often


by the germination of the

latent, until it is called into

seed.

Sometimes

distinguishable from the cotyledons; sometimes


developed, and lies in a furrow of the cotyledon

it
;

it is
is

un-

highly

Ex. Maize
F2

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

68
2

".

In the monocotyledonous embryo

it

frequently happens

margins of
which grow together, so that the whole embryo forms one
uniform mass 293 ; but as soon as germination commences the

that the plumule

is

rolled

up

in the cotyledon, the

margins separate.
581. The radicle elongates downwards, either directly from
the base of the embryo, or after previously rupturing the integument of the base. Plants with the first character are
called ExoRHiz^s 295

582.

with the second, ENDORHiz^; 298 2 ".

The endorhizal embryo

is

very

common

in

monocoty-

the exorhizal, in dicotyledons.


583. When the' seed is called into action, germination takes
The juices of the plant, which before were insipid,
place.
ledons

immediately afterwards abound with sugar ; Ex. Barley ; and


growth commences.
584. This growth is in the first instance caused by the
absorption and decomposition of water, whose oxygen combines with the superfluous carbon of the seed,
in the form of carbonic acid
gas.

585.

As

this

and

phenomenon does not take place

plants, except in the

expelled

in full-grown

dark (258), so neither can

seeds, except under the same condition.


exposed to constant light, would not

is

it

occur in

Hence an embryo,

germinate at all ; and


hence the care taken by nature to
provide a covering to all
embryos in the form of the integuments of the seed or of a

pericarp.

586.

As

soon as the
necessary proportion of carbon is reseed by the expulsion of carbonic acid, the

moved from a

young plant begins


cesses of assimilation

and to grow by the proand respiration already described (254).

to absorb food,

ACROGENS, OR FLOWERLESS PLANTS.


587. Many plants not
being increased by seeds, the result
of the mutual action of sexual
apparatus (531), are flowerless,
and destitute of
organs of fructification.
588. Such are
propagated by what are called organs of reproduction, which have no other
analogy with the organs of
fructification than that both
perpetuate the species.

ACROGENS, OR FLOWERLESS PLANTS.


589.

The reproductive organs of flowerless

69

plants vary ac-

cording to the tribes of that division of the vegetable kingdom ;


and have so little relation to each other, that each principal
tribe may be said to have its own peculiar method of propagation.

590.

They

all

agree in their reproductive parts or spores,


seeds, not germinating from any fixed

which are analogous to

point, but producing root or stem indifferently from any point


of their surface.
This germination is therefore vague.

591. The principal tribes are Ferns (592), Mosses (598),


Lichens (605), Algacea (607), and Fungacea (610).
592. FERNS are increased by, little bodies, called spores,
enclosed within cases named tliectz or sporangia 302 303 which
often grow in clusters or sori 3M , from the veins of the under
,

from beneath the epidermis. The


termed the indusium 308

sides of the leaves, or

when

it

593.

encloses the

The indusium

in consequence of the

594.

one

thec<e, is

The

side,

thecse

and

latter,

separates from the leaf in various ways,


growth of the thecse beneath it.

have frequently a stalk which passes up

finally,

curving with their curvature, disappears

on the opposite side 307


595. The part where the stalk of the theca
.

its side, is

called the annulus.

is

united with

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

70

596. These thecse

may

be considered minute leaves, having

the same gyrate mode of developement as the ordinary leaves


of the tribe ; their stalk the petiole, the annulus the midrib,

and the

theca; itself the lamina, the edges of

which are united.

597. They would, therefore, be analogous to carpels,


appeared that they were influenced by the action of any

if it

vivi-

fying matter.
598. MOSSES

are increased by spores (590), contained


within an urn, or theca, or sporangium 31 * 316, placed at the
apex of a stalk or seta, bearing on its summit a kind of loose
31
hood, called a calyptra *, and closed by a lid or operculum.

599.

and the

The

inside of the theca has a central axis or columella,

orifice

processes, or a

600.

At

beneath the operculum

membrane

is

closed

called the peristome 313

the base of the theca

is

by

teeth- like

318
.

sometimes found a tumour

or struma 31 *, or an
equal expansion named apophysis 319 .
601. The number of the teeth of the
is

peristome

some multiple of four.

always

The calyptra
originally grew from the base of the
when the stalk lengthened, the
calyptra was torn
away from its base and carried up,
surrounding the theca.
603. The
be understood to
calyptra
602.

stalk; but

may

at;

be a convolute

the operculum,
another; the peristome, one or

more

ACROGENS, OR FLOWERLESS PLANTS.


whorls of minute

flat

leaves

and the theca

71

be the

itself to

excavated distended apex of the stalk, the cellular substance


of which separates in the form of sporules.
604. There are also in mosses certain organs, called anthers

by some, which do not appear analogous to the male apparatus of flowering plants, and the nature of which has not
been demonstrated.

They

are jointed filaments, staminidia or

antheridia, containing vibrios lodged in


round the rudiment of the future theca.

At

figure 315 the flask-like figure is a


tillidium ; and the club-shaped body

lated threads

may

mucous

cells,

young sporangium, or
on

its left,

and

sur-

in this state pisThe articu-

a staminidium.

be abortive staminidia.

605. LICHENS are cellular expansions, usually horizontal,

but occasionally perpendicular, consisting of a tkallus 331 , or


combination of stem and leaves, upon which shields, apotliecia,
331

or reproductive organs, appear


606. The shields consist of a margin, enclosing a kernel,
nucleus, in which tubes containing sporules, and called asci,
.

are imbedded.
They vary a little
scutellum 337

gyroma ;

whence they have received the following other names


which is the same thing ;pelta 339 tuberadum 335 ; tricaor

in nature,

orbilla,

if covered

with sinuous concentric furrows, lirella

333
;

patellula

334
.

Be-

sides the foregoing, some other peculiar terms are used by writers on Lichens.
Asci are tubes of the nucleus, containing sporules ; the latter are sometimes
named gongyli : periikedum is the part in which asci care immersed hypothe;

a substance overlying the pcrilluvinin.


339
tions of the thallus
;
scypha or oplarium

cium

is

J'oi/ctiu are stalk-like elongais

a cup-like expansion of a

STRUCTURAL AND PHYSIOLOGICAL BOTANY.

72

on the margin
podetium, having shields
they are also called globuli and glomtruli.

338

Soredia, or

3
powdery masses ^

.
Lacuna, are pits of the thallus
which forms the rim and base of shields.
Excipulm is that part of the thallus
Tftallodes signifies formed of the thallus.

of any
607. ALGACEJS are submersed plants, equally destitute
kind of tissue, except the cellular, and propagated by spores

of the system.
(590) lodged in various parts
of
608. The sporules either lie freely in the whole substance
3
or
cells
",
occupy
or are collected in particular
such
plants,

321

324

in spheres
occupying
jointed filaments , or are placed
thallus (605).
of
the
circumferences of expansions
609. There are also other modes of multiplication.
,

the

the special terms employed by writers on this order, the following may
be enumerated as the principal. Among their reproductive organs are gongyli,
or hard round deciduous bodies ; granula, or large spores ; sporidia, or bodies
321 325
.
resembling spores, but not such ; sporangia or coniocysta, or spore-cases
Hypha is a filamentous thallus ; phycomatcr is the gelatine in which the
spores of some begin to vegetate ; peridiolum is a membrane immediately

Among

covering the

spores

vesicula are air-bladders that enable

some species

to

float.

610. FUNGACE.E, which are the lowest form of vegetation,


also cellular, some of their cells however containing
spiral threads, and are propagated by spores.

are

611. In the highest forms, two kinds of organs are de545


are conical naked elevations ; the
one, cystidia
,

tected

other, basidia

3*5
,

spores in definite

are also conical

number on

elevations, but

their apex.

they bear

ACROGENS, OR FLOWERLESS PLANTS.

73

612. The highest forms of the fungaceous order consist


of a stipes 6, an annulus or collar 346 , a pileus 346 or cap, and

an hymenium.
613.

Lower forms

are reduced to a

mere peridium or

in-

342
tegument, containing the reproductive system .

614.

615.
to end,

Some have the sporules enclosed in asci (606).


The lowest consist of nothing but cells, placed end
and enclosing spores

in the terminal cells

344
.

Of

the special terms employed by Mycologists (writers on Fungaceous plants),


The volva is the wrapper which covers over
the principal are the following.
many of them, as Agarics, in their youngest state. Thattus is the spawn
under
Velum is a
ground, or amongst decaying matter.
usually generated
membrane that connects the pileus and collar. Cortina is that part of a velum

which adheres to the margin of the pileus. Flocci are wool-like threads found
mixed with sporules ; and stroma is the body on which flocci grow. Orbiculi
little disks contained within the peridia of certain genera.
Sporangium
Peritkecium is the
the external coating of such genera as Lycoperdon 343 .
348
Ostiolum is the mouth of the bag.
bag of fructification in Sphaeria .
34
Capillitium is a kind of purse or net containing spores *. Mycelia are
nascent fungi, or fragments of their spawn.

are
is

II.

SYSTEMATICAL BOTANY.

616. SYSTEMATICAL

BOTANY

is

the science of arranging plants

such a manner that their names

in

affinities

determined, their true

may

place

be ascertained, their
a natural system

in

judged of, and their whole hisand


with
elucidated
tory
certainty
accuracy.
617. Anything short of this is not a system, but an artificial
fixed, their sensible properties

scheme.

The latter is intended to enable a person


name of a plant, and goes no further.
619. But as the name of a plant conveys no
618.

to ascertain

the

by

itself,

the power thus acquired

by

artificial

information

schemes

is

of

and cannot be considered as anything


beyond a very imperfect and elementary mode of investibut

little

real value,

gation.

What

620.

scheme

is

knowledge is gained by the use of an artificial


mere
a
collection of isolated facts, without mutual

dependence, or any distinct bearing upon general views.


621 In a natural arrangement, on the other hand, the name
.

of a plant

the least object that

gained. Any investigation


is, of necessity, attended
with the discovery of the relationship a given plant bears to
others ; and as plants which are most closely akin in struc-

upon

is

its principles,

is

when completed,

ture are also most similar in their sensible properties, it often


enables us to judge of the use of an unknown plant whose
place is determined in the system, by the ascertained uses of
those species in whose vicinity it takes its place by virtue of
its natural affinities.

622. The only artificial schemes in general use are, 1, that


of Linnseus (623), called the SEXUAL System, in consequence
of its characters being dependent upon variations in the sta-

mens and
method.

pistil,

or sexes, of plants

and

2,

the Analytical

SYSTEMATICAL BOTANY.

76

I.

623. 'This

LINN^AN SEXUAL SYSTEM.

now

is

disused

by men of science

books
employed have been arranged upon
a student to understand it.
for
necessary
still

but, as

many

its plan, it is

624. Its divisions, called classes and orders, depend upon


of the stamens and pistils, and have Greek

modifications

names expressive of
Class

their distinctive characters.

2.

Monandria. Stam. 1.
Diandria. Stam. 2.

3.

Triandria. Stam. 3.

1.

4.

Tetrandria. Stam. 4.

5.

Pentandria. Stam.

6.

7.

Hexandria. Stam. 6.
Heptandria. Stam. 7.

8.

Octandria. Stam. 8.

9.

Enneandria. Stam.

5.

9.

10. Decandria. Stam. 10.

12 19.
20 or more, perigynous (359).
13. Polyandria. Stam. 20 or more, hypogynous (358).

11. Dodecandria. Stam.


12. Icosandria. Stam.

Each of these classes


by the number of styles

Orders.
terized

nia signifies

style

is

divided into orders charac-

or sessile stigmas.

Digynia, 2

4 ; Pentagynia, 5 ;
Octogynia, 8 ; Enneagynia, 9 ; Decagynia, 10
nia, &c. about 12 ; Polygynia, many.

nia,

Class 14.

Didynamia
Orders:
2.

1.

Monogy-

Trigynia, 3 ; TetragyHexagynia, 6 ; Heptagynia, 7 ;

Dodecagy-

Stamens 4, two long and two short.


Gymnospermia, seeds apparently naked ;

Angiospermia, seeds evidently in a seed-vessel.

Class 15. Tetradynamia

Stamens

6,

four long

and two

short.

Siliquosa, with a long pod; 2. Siliwith


a
short pod or pouch.
culosa,
Class 16. Monadelphia: Filaments united into a
cup or column.
Orders: 1. Pentandria; 2. Decandria, &c.

Orders:

1.

as before.

Class 17. Diadelphia:


fraternities.

Filaments united into two parcels or


Orders : 1 Hexandria, Sic. as before.
.

LINN-ffiAN

Class 18. Polyadelpliia

than two.

SEXUAL SYSTEM.

77

Filaments united into more parcels


1. Dodecandria;
2. Icosan-

Orders:

dria, SEC. as before.

Class 19. Syngenesia: Anthers united into a tube.


Orders:
1. Monogamia, flowers
solitary; 2. Polygamia,
flowers in heads.

Sub-orders of the latter:

1.

u3qualis, florets all equal ; 2. Superflua, florets


of the disk complete, of the ray female ; 3. Frustranea, florets of the disk perfect, of the ray

neuter

4. Necessaria, florets of the disk male, of


;
the ray female ; 5. Segregata, florets each with its

own

proper involucre.

Class 20. Gynandria


Stamens and styles consolidated.
ders: 1. Monandria, &c. as before.

Or-

Class 21. Monoecia: Stamens in one flower, pistils in another,


on the same plant. Orders: 1. Monandria, &c. as
before.

Stamens in one flower, pistils in another,


on different plants. Orders: 1. Monandria, &c.

Class 22. Dicecia

as before.

Class 23. Polygamia: Stamens and pistils separate in some


flowers, united in others, either on the same plant
or on two or three different ones. Orders: 1. Moncecia,

&c. as before.

Class 24. Cryptogamia


lices,

no apparent flowers.

Orders

Musci, Hepaticse, Algae, Fungi (592).

Fi-

SYSTEMATICAL BOTANY.

78

ANALYTICAL METHOD.

II.

625. THIS

is

founded upon the

common

process of analysis

In all
unconsciously employed by the human mind.
cases the mental operation by which one thing is distinguished
from another, consists in a continual contrast of characters.
that

is

For instance, in a mass of individuals we distinguish one set


which is coloured, and another which is colourless of those
that are coloured we distinguish red, black, blue, and green
of the round,
of the red, some are square, others are round
some are sculptured on their surface, others are even
and so
;

we

proceed, analysing the subject by a constant series of contrasts, until we have arrived at a point beyond which no
analysis can go.

The

following pages contain such an analysis of the


natural
orders of plants.
The method may be
principal
and
to
applied
genera
equally
species, and is an instructive

626.

process if employed by way of exercise to the mind,


the purpose of rendering distinctions definite

and

for

1.

Plants with visible

seeds

flowers and

.2

Plants with visible flowers and


spores

Plants without flowers


2.

Wood

Leaves netted.

centric layers

in

confused

in concentric
layers
3.

HOMALIACEJE.

ANONACE^.

Carpels consolidated
11. Placenta

over the

spread

.3

...'

239

12. Petals definite in

Wood

237

13.

LOASACE.E.

CACTACE^E.

Leaves

with

transparent

Petals united

dots

MYRTACE^.

.4

.12
.13

number

Petals indefinite

Flowers having a calyx only, or


none
.
.
119
4. Petals distinct

septa

Placenta parietal
Placenta in the axis

Flowers having both calyx and

corolla

.11

con-

....

Leaves straight-veined.

LECYTHIDACEJE.

Placenta parietal

Wood

Leaves straight-veined.

Placenta central

10. Carpels distinct

RHEANTHS.
.261

.-

9.

Leaves dotless

14. Petals indefinite

.14

MESEMBRYACE^;.

Petals definite
PHILADELPHACE.*;.
Leaves with stipules
.
Leaves without stipules
24
.
.

15.

186

.16

5.

Stamens more than 20

Stamens fewer than 20


6.

7.

Ovary

inferior, or partially so

Ovary

superior

15

Leaves furnished with


stipules
Leaves without stipules
.

8.

Carpels more

or less disunited

Carpels consolidated

16. Carpels disunited

36

.10

POME.E.
9

Carpels consolidated
17.

Stamens hypogynous
Stamens perigynous

MAGNOLIACE^E.

18. Placenta parietal

Placenta in the axis

.17
.18

ROSACES.

BIXACE^J.

19. Estivation of
calyx imbricated

Estivation valvate

.19
.

20

22

ANALYTICAL METHOD.
EUPHORBIACE,E.

20. Flowers unisexual

Flowers hermaphrodite
21.

Ovary

one-celled

Ovary

wit<h

PORTULACACE^:.

more

than one

cells

and

enlarged

irregular

DlPTERACE^E.
Calyx not enlarged
23. Stamens monadelphous

Stamens

MALVACEAE.

distinct

24. Carpels disunited or solitary

30

NELUMBIACE^E.
26
the disk

Carpels clear of

Placentas axile

.
27
ANACARDIACE.E.

Stamens hypogynous
27. Carpel solitary

ROSACES.

43. Disk double

Stamens

free

29. Herbs

Carpels several

on

46. Parasites

48. Calyx valvate

.31

34

Stamens

32. Sepals

51. Leaves dotted

PORTULACACE.S.

Sepals more than 2

seeds

flat,

few,

33

phous

stamens

flat,
.

MYRTACE.E.
52
.
Anthers curved downwards
.

52.

34. Placentas spread over the lining


of the fruit
FLACOURTIACE.E.
.
.

35.

Ovary stalked

36.

Ovary sessile
Ovary more or

.
.

35

CAPPARIDACE.E.

PAPAVERACE.E.

less inferior

37

quite superior

55

37. Leaves with stipules

38

Ovary

Leaves without stipules


.41
38. Flowers unisexual
.
BEGONIACE^E.
.

Flowers hermaphrodite

MELASTOMACE^E.
.

53. Flowers tetramerous

.53

ONAGRACE^E.
Flowers not tetramerous
54
.
.

54. Petals always distinct

SAXIFRAGACE^:.

HUMIRIACE.33.

Placentas in lines

BRUNIACE.E.

monadel-

50

CORNACE^E.

Leaves not dotted

Anthers erect

leaves

.
.
CLUSIACE-E.
leathery .
Petals crumpled, seeds numerous,
leaves membranous
.
CISTACE..E.

Petals

downwards

Sepals united into a tube LYTHRACEJE.


33. Petals

.49

MEMECYLACE^E.

.32

RHAMNACE^E.

altern. petals

Anthers erect

...

COMBRETACE^E.

50. Anthers curved

SARRACENIACE^E.
.

.47

31. Stigma large, umbrella-shaped

Stigma simple

LORANTHACEJE.

Calyx imbricated
49. Stamens opp. petals

Placentas dissepimental

NYMPH-EACE.E.

.44
.45
.51
.46
.48

ANACARDIACE^E.

ANONACE.E.

Placentas parietal

47. Leaves balsamic, acrid

Leaves insipid

30. Placentas in the axis

trees

Root plants

APIACE.E.

Seeds numerous
45. Carpels solitary

RANUNCULACE^E.

Trees or Shrubs

.43

CUCURBITACEJE.

.29

Disk simple
44. Seeds few

.28

.
Carpels several .
28. Stamens polyadelphous HYPERICACE^E.

42

Flowers hermaphr. or polyg.


GROSSULACE.E.

42. Flowers unisexual

26. Stamens perigynous

RHAMNACE.E.

41. Placentas parietal

25

.
Carpels consolidated
25. Carpels plunged in a tabular disk

Calyx imbricate, stamens alternate


with petals
HAMAMELACE.S.
.

TILLAGES.

40. Calyx valvate, stamens opposite

.23

HOMALIACE.E.
40
.

Placentas in the axis

the petals

ClSTACBLE.
22. Calyx

39. Placentas parietal

.21

79

39

Petals at

first

united into a tube

55. Leaves with stipules

ESCALLONIACRS.
56
.
.

Leaves without stipules

56. Carpels disunited

Carpels consolidated

81

.57

.
.

59

57. Anther valves recurved

BERBERACE.E.
Anther valves

straight

58. Fruit leguminous

.58

FABACE^E.

Fruit drupaceous or capsular

ROSACES.

SYSTEMATICAL BOTANY.

80

'

59. Placentas parietal


Placentas in the axis

62

81. Carpels disunited


Carpels consolidated

Flowers without a coronet

Anthers with straight valves

61

83. Fruit leguminous

when young

Fruit not leguminous

DROSERACE-E.
Leaves straight when young
VIOLACE.E.

63

.64

Sepals more than 2

65

65. Styles distinct to the base

66

Styles more or less united

72

66. Petals minute

Petals conspicuous
68. Stigmas capitate

Stigmas simple

.68

Leaves not dotted


86. Carpels solitary

EL.SOCARPACEJE.

Calyx imbricated

CUNONIACE^;.

88.

Hypogynous

Calyx simple
75. Leaves simple

Trees or shrubs

73

Flowers hermaphrodite

91

CALYCANTHACE.S.
Stamens hypogynous
92. Stamens indefinite
Stamens

definite

ZYGOPHYLLACKE.
.

CORIARIACE^E.

NYMPH.EACE.E.

95.

Stamens not tetradynamous

95

Hypogynous disk

large

96

Hypogynous disk

small or

97

96. Stamens indefinite

Stamens

STAPHYLEACE.E.

VITACE^E.

Sepals distinct

98. Sepals 2-3

Sepals 5
99.

Brown

parasites

80

CHAILLETIACE.E.

BURSERACE.E.

100. Styles distinct

Styles consolidated

101. Stain,

98

PAPAVERACEJE.
TURNERACEJE.

MONOTROPACEJE.

Green rooting plants

EL^EOCARPACE^E.

RESEDACILE.

FRANKENIACE.E.

'

.
78
Stamens alternate with petals
.
79
78. Stamens perigynous .
RHAMNACE^E.

slits

CAPPARIDACE^E.

definite

97. Sepals tubular

77. Stamens opposite the petals

.94
.99

BRASSICACE^E.

76

SAPINDACE^E.

Stamens hypbgynous
79. Anthers opening by pores

92

ANONACE.E.

Placenta parietal
Placenta axile .

.75

93. Placenta dissepimental

Flowers symmetrical

Petals undivided

.90

94. Stamens tetradynamous

double

89

MENISPERMACEJE.

MALPIGHIACELE.

Anthers opening by

88

FRAXCOACE.E.
RANUNCULACE^E.

90. Flowers unisexual

.74

Leaves compound
76. Flowers unsymmetrical

80. Petals split

double

scales

.77

distinct

.,

scales simple

89. Herbaceous plants

OXALIDACEJJ.

surrounded with
.i

.87

CRASSULACE^.
Hypogynous

SAXIFRAGACE.E.

73. Stamens monadelphous

glands

.71

72. Calyx imbricated


Calyx valvate or open

74. Calyx

.86

MALPIGHIACE^;.
Stamens perigynous
71. Leaves opposite

Stamens

FABACE^.

91. Stamens perigynous

70. Stamens hypogynous

Leaves alternate

Carpels without ditto

.70

85

87. Carpels with hypog. scales

ILLECEBRACE^E.

ANACARDIACE^S.

69

ELATINACE.E.

69. Calyx valvate

84

AMYRIDACE.E.

Carpels several

Fruit without a beak

83

CoNNARACEiE.
85. Leaves dotted

64. Fruit with a long beak

GERANIACE^.

Radicle remote from hilum

PORTULACACE.E.

84. Radicle next hilum

EUPHORBIACE.E.

Flowers hermaphrodite or polyg.


63. Sepals 2

.93

BERBERACEJ3.

PASSIFLORACE^.

62. Flowers unisexual

82. Anthers with recurved valves

60. Flowers with a coronet

61. Leaves circulate

82

.
.

.100
.101
.105

polyadelphous HYPERICACE.E.

Stamens

free

.102

ANALYTICAL METHOD.
with

102. Carpels
scale

an

Carpels without

124. Carpels triple

hypogynous
CRASSULACE-E.

do.

103

Stamens

106. Seeds wingless

PIP BRACED.
Flowers unisexual

Sepals more than 2

inferior

Ovary

superior

108. Anthers opening by pores

slits

109

.110
.112

110. Fruit succulent


.

.111

111. Flowers hermaphrodite

Flowers not
1

BALSAMINACE^E.

.113

do.
.

Calyx imbricated

115. Stam. opposite petals

Stamens more
petals

RHAMNACE.E.

numerous

134. Leaves

Stamens perigynous
118. Seeds comose

Seeds naked

120

121

126

SALICACE^E.
.

.128

Leaves without stipules


121. Ovules numerous
.
122. Flowers hermaphrodite

.122
.

123

.124

CHLORANTHACE^E.
Stamens whorled
SAURURACE^E.
.

37

.138
.142

Flowers^/
143. Leaves with

.140

SANTALACE^E.

.141

COMBRETACE^E.

Embryo straight
Embryo curved

142. Flowers^/

.139

ONAGRACEJE.

141.

Flowers unisexual

CELASTRACE/E.

Flowers not^/

117

.118

123. Stam. unilateral

2-6-celled

TAMARICACE^E.

Calyx present
120. Leaves with stipules

Ovules few

LORANTHACE<E.

PITTOSPORACE^E.

Ovary

140. Calyx valvate


Calyx not valvate

.'

MYRTACE,E.

than

MYRICACE,E.

38. Parasites on branches

139. Flowers^/

GARRYACE.E.

Leaves not dotted

.135

136. Leaves dotted

EMPETRACE.S.

Flowers hermaphrodite

JUGLANDACE^E.

Leaves simple .
135. Leaves opposite

137. Ovary 1-celled

.134

compound

Terrestrial

117. Stamens hypogynous

36

CUCURBITACE.E.

LYTHRACE.E.

116. Flowers unisexual

119. Calyx none

.114
.115
.116

133. Climbing tendrilled herbs

Leaves alternate

SIMARUBACE.E.

.133

Flowers herm. or polyg.

113. Stamens arising from scales

114. Calyx valvate

BEGONIACE.E.

Trees or shrubs

Flowers regular

Stamens not

do.

CORYLACEA;.

32. Flowers unisexual

RUTACE.E.

XANTHOXYLACE^E.

.131

131. Flowers amentaceous

Flowers polygamous
112. Flowers irregular

132

Flowers unisexual

AURANTIACE.E.

Fruit capsular

Leaves not dotted

ARISTOLOCHIACE^E.
ERICACEAE.

Anthers opening by
109. Leaves dotted .

.129
.143
.130

Leaves without stipules


130. Flowers hermaphrodite

.108

CALLITRICHACE.E.

Ovary

129. Leaves with stipules

PORTULACACE.E.

MYRICACE.E.

Carpels double
128.

CEDRELACE.E.

106

.107

.127

127. Carpels single

MELIACE^E.

Seeds winged
107. Sepals 2

PLATANACE^E.

104

CARYOPHYLLACE<E.

free

MYRICACE^E.

126. Flowers hermaphrodite

LINAGES.

J05. Stamens monadelphous

.125

Ovule pendulous

Stigmas simple

125. Ovule erect

SAXIFRAGACE.E.
Carpels parallel

EUPHORBIACE.E.

Carpels single

103. Carpels two divaricating

104. Stigmas capitate

81

CHENOPODIACE^.
ONAGRACE^E.
ARISTOLOCHIACKE.

stipules

Leaves without stipules


144. Flowers hermaphrodite
Flowers unisexual
145. Sepals 2

Sepals more than 2

Carpels solitary

160

145

47

PORTULACACE.S.

146. Carpels more than

.144

46

consolidated 147
.

.153

SYSTEMATICAL BOTANY.

82
147. Stamens hypogyno us

.148
.150

Stamens perigynous
148. Fruit beaked

170. Stamens hypogynous

CARYOPHYLLACE^-:.
Stamens perigynous

GERANIACELE.

Fruit not beaked

49

Fruit many-celled

149. Calyx biglandular imbricated

MALPIGHIACE.E.
Calyx eglandular valvate TILIACEX.
PASSIFLORACE.E.

150. Placenta parietal

Placenta axile

CUNONIACE^E.

Leaves alternate

153. Calyx

.152

152. Calyx valvate


Calyx imbricate

RHAMNACE.E.
ULMACE^;.

membranous

ILLECEBRACE.E.

Calyx firm and herbaceous

Styles terminal

Fruit not leguminous

Anther valves

sepals

158. Carpels solitary

Calyx tube membranous


1

78. Cal. all hardened

Base only of

cal.

hardened

NYCTAGINACE.E.

THYMELACE^E.

.159
BETULACE^.

Calyx tubeless

.183

62

Calyx herbaceous

CHENOPODIACE^E.

183. Carpels solitary or distinct


.
Carpels consolidated

Carpels solitary or disjoined

Ovules

many

165. Leaves dotted

Calyx open, carpels several

.168

RUTACE^E.

.166

.167

167. Carpels numerous, separable

PHYTOLACCACE^;.
Carpels few, inseparable

MENISPERMACEJE.
XANTHOXYLACEJE.

Leaves not dotted

EUPHORBIACE.E.

Ovary

inferior

187. Flowers regular

Flowers irregular
188.

168. Carpels 2,
divaricating

SAXIFKAGACE.E.
.

170

.187

RUTACE.E.

.190

226

.188
.218
.189
.192

.,

Ovary deeply split


Ovary not split

189. Leaves dotted

Leaves not dotted

CELASTRACE.E.

Carpels not divaricating

186. Ovary superior

OLEACE.E.

185. Leaves dotted

165

compound

Leaves simple

.164

184

.185

MYRISTICACE/E.

172

PAPAVERACEJE.

Leaves not dotted


166. Leaves

184. Calyx tubular, carpel


solitary

162. Carpels more than 2, consolidated 163

.182

AMARANTACE^.

161

PORTULACACE.E.

Sepals more than 2

.181

182. Calyx dry and coloured

Flowers unisexual

.180

EtJPHORBIACEjE.

1S4. Ovules few

179

Leaves not lepidote


181. Calyx tubular
.

URTICACE^E.
.

160. Flowers hermaphrodite

Placenta axile

SCLERANTHACE.*.

EI^AGNACEJE.

Flowers not amentaceous

163. Placenta parietal

.177
.178

180. Leaves lepidote

159. Flowers amentaceous

PROTEACE^E.

do.

PETIVERIACE^E.
.

of

POLYGONACE.E.

Carpels more than one

161. Sepals 2

points

179. Fruit triangular


Fruit rounded

.176

the

177. Calyx hardened

.156

Styles triple

.175

AMYRIDACE.E.

within

FABACE^;.

Leaves not dotted


176. Stamens

.174

.157
ROSACES

Stipules simple or

157. Styles simple

LAURACE.E.

straight

POLYGONACE^E.

156. Stipules ochreate

.173

175. Leaves dotted

.155

LYTHRACE^E.

RANUNCULACE^E.

174. Fruit a legume


Fruit not do.

154

FABACE^E.

Carpels solitary

Stamens not

155. Fruit leguminous

PRIMULACE^:.

173. Anther valves recurved

154. Styles from the base of ovary

CHRYSOBALANACE^J.

172. Carpels several

.151

151. Leaves opposite

.171

171. Fruit one-celled

190. Inflorescence
gyrate BORAGINACE.E.
Inflorescence straight
.
.191
191. Estivation
plaited

Estivation

flat.

NOLANACE*:.

STACKHOUSIACE^E.

ANALYTICAL METHOD.
192. Carpels 4, 5, or more

.193

83

SYSTEMATICAL BOTANY.
235. Leaves alternate

EBENACE.E.

Leaves opposite

254. Stamens very short

Stamens long, weak

236

GALIACKS.
.
236. Fruit didymous
Fruit not didymous CAPRIFOLIACKS:.

255. Floating

237. Stem cylindrical, unbranched

256. Ovules pendulous

Stem

conical, branched

238. Fruit solitary


Fruit in cones

Ovules erect

238

240

246

Ovary superior

MELAXTHACE^E.
Anthers turned inwards
258. Stems herbaceous

Stems woody

Flowers hexapetaloid
260. Flowers coloured

Anthers more than one


242. Anther one-celled

243

COMMELINACE.E.
260
.

MARANTACELE.

Anther two-celled

ZINGIBERACKS.

Flowers scarious

Axis

244. Stamens 3

IRIDACE^E.

Stamens 6

245

Stamens more than 6

HYDROCHARACE.E.
AMARYLLIDACE^E.

245. Sepals petaloid


Sepals herbaceous
.

Flowers regular
247. Sheaths of leaves

slit

247

248

GRAMINACE^.

Sheaths of leaves closed

SMILACE^.
249

Leaves straight-veined
249. Carpels disunited

Carpels consolidated

250

259

250. Placentae dhsepimental BUTOMACBL&.


Placentae axile

251. Flowers imperfect


Flowers perfect
252. Flowers on a spadix
Flowers scattered
253. Fruit drupaceous
Fruit berried
Fruit dry

251

252

257

253

255

PANDANACEJE.
.
ARACE.S.
254
.

LILIACEJE.

JUNCACE^E.
.

262

CHARACE^E.

distinct, leafless

Axis confused

.268

FILJCALES.

Sporangia arising from the stem 263


263. Sporangia involucrate .
264
.

Sporangia naked
264. Involucres uniform

.265

MARSILEACEJE.
Involucres 2-formed

265. Sporangia
valved

SALVINIACE^E.

axillary,
.

sessile,

.266

BRYACE^:.

Sporangia valvate
267.

An operculum
No operculum

2-

LYCOPODIACEJE.

Sporangia stalked
266. Sporangia valveless

CYPERACE.E.
248. Leaves netted

262. Sporangia upon leaves

BROMELIACE.S:.

246. Flowers glumaceous

261. Axis distinct, leafy

243. Veins of leaves diverging MUSACEJE.


Veins of leaves straight
. 244

Veins of leaves netted DIOSCOREACILE.

258

PALMACE^E.

242

...

ALISMACE^E.

259. Flowers seinipetaloid

PISTIACE-E.

ORCHIDACKE.
240. Flowers gynandrous
Flowers not gynandrous
241
.
241. Anther

NAIADACE/E.

257. Anthers turned outwards

PINACE^E.

.256

JUNCAGINACE.E.

TAXACE.E.

239. Ovary inferior

ACORACEJE.

TYPHACE.E.

Terrestrial

CYCADACILE.

267

ANDR.EACE.S.

JUNGERMANNIACELE.

268. Stomates

269

No

270

stoniates

269. Sporangia valvate, operculate

JUNGERMANNIACE.E.
Sporangia valveless, without an

operculum
270. Submersed
Aerial

MARCHANTIACE^E.

271. Thallus superficial


Thallus buried

ALGACK^E.

.271

LICHENACKS.

THE NATURAL SYSTEM.

THE NATURAL SYSTEM.

III.

627.

THE

85

true Natural System, whenever

it

shall

be dis-

covered, will represent the species, genera, orders, alliances,


groups, subclasses, and classes of plants, or whatever other
divisions

may

shall stand

be admitted into

it,

next those to which

structure than to

any

so arranged that each plant


is more
nearly allied in

it

others.

of man has not yet attained this end ; no


to
this description has been devised, nor
system answering
does there appear any probability that it will be discovered till
our knowledge of plants is much more advanced.
628. But the

skill

629. All so-called natural systems are, to the present day,


The lower and higher
partly artificial and partly natural.
divisions in

them

are natural, the intermediate divisions are

In other words, the stones of the edifice are hewed


and squared, and the general plan is drawn out, but no
artificial.

builder has yet been found with skill to put

them

together, so

as to form a consistent whole.

630. But although in theory no system that can properly be


called natural has yet been devised, yet for practical purposes many answer to the name, and fulfil the principal conditions required of them.

631.

The genera and natural orders can alone be considered


upon by botanists, the other divisions are unsettled

as agreed

and

this is the reason

why

the natural orders seldom follow in

the same manner in the arrangements of two different botanists.


is no such thing as an arrangement which
express the natural relations of plants in a consecutive

632. There
shall
series.

633. It seems to be generally admitted

by those who have

turned their attention to the consideration of the manner in

which organized beings are related to each other, that each


species is allied to many others in different degrees, and that
such relationship

is

best expressed

by rays

(the affinities)

SYSTEMATICAL BOTANY.

86

common

In like
centre (the species).
of
the
several
the
mutual
relationship
manner,
studying
of
distribution
form
the
same
of
the
vegetable kingdom,
parts
and orders
constantly forces itself upon the mind; genera
of
whose
the
centre
be
to
found
spheres,
apparently
being
surface is only defined by the points where the last traces of
proceeding from

in

affinity disappear.

634. But although the

mind may conceive such a

distribu-

tion of organized beings, it is impossible that it should be so


presented to the eye, and all attempts at effecting that object
have failed. If in describing the surface of a sphere we are

compelled to travel in various directions, continually returning back to the point from which we started, and if in preeye at one glance we are compelled to
a
plane, the effect of which is to separate
upon
project
to the greatest distance some objects which naturally touch
senting

it

to the

it

how much more impossible must it be to follow


the juxtaposition of matter in treating of the solid contents of

each other,

a sphere.
635. The fundamental principle of systematic botany is, that
those plants should be stationed in company with each other
which have the greatest degree of affinity, and that those

should be placed most remotely which have


degree of affinity.

the smallest

636. Affinity is an accordance in all essential characters.


637. From this is distinguished analogy, which is a conformity in one or two characters only.
638.

What we

call

the characters of plants are


merely the
affinity, and all the groups into

by which we judge of
which plants are thrown are

signs

in

one sense

as nature recognises no such


groups.
639. Nevertheless, consisting in

artificial,

inasmuch

cases of species
very
closely allied in nature, they are in another sense natural.
all

640. But as the classes, subclasses,


groups, alliances, natural
and genera of botanists have no real existence in

orders,

follows that they have no fixed


limits, and consethat
it is
quently
impossible to define them.
641. They are to be considered as
more than

nature,

it

the
nothing
expression of particular tendencies (nixus), on the
part of the

THE NATURAL SYSTEM.

87

plants they comprehend, to assume a particular

mode of deve-

lopement.
642. Their characters are therefore nothing more than a
declaration of their prevailing tendencies, and are liable to

numerous exceptions.
This

liability, it

must be remarked,

the natural system

exists as

much

in all artificial

schemes as in

itself.

643. If a system

is

ever to be devised which shall

by com-

mon consent be admitted to be natural in all its parts,


as human means can make it so, this will be brought

as 'far

about

by settling the relative value of the characters by which


plants are limited, and by introducing uniformity and consistency into the distinctions of the groups, whether inferior,
superior, or intermediate.
to the present time, no attempt at settling these points has been successful,
and consequently the characters employed in denning the limits of groups,
of all denominations except the highest, are arbitrary and inconsistent.

Up

644.
1.

The

following propositions seem incontrovertible


is constant can be regarded as
unimportant.
:

Nothing that

2. Every thing constant must be dependent upon or conTherefore all constant


nected with some essential function.

characters,

of whatever

nature,

require

to

be taken into

account in classifying plants according to their natural

affi-

nities.
nature are the internal structure of stems and leaves, the anatomical condition of tissue, the organization of the anther, pollen, and female apparatus,
and the interior of the seed.

Of this

645.

On

the other hand, whatever points of structure are

same species, or in species nearly allied to each


other, or in neighbouring genera, are unessential to the vital
functions, and should be set aside, or be regarded as of compavariable in the

rative unimportance.
Hence the badness

of the Monopetalous, Polypetalous, and Apetalous divisions


upon the presence or absence, and union or disunion, of

of Jussieu, depending

The genus

Fuchsia, for example, has petals highly developed ; but in


F. excorticata they are absent, and yet the plant differs no otherwise from the
rest of the genus : the same is true of species of Rhammis.
Again, the Rue
has the petals separate ; and Correa, very nearly allied to it, has them competals.

bined.

SYSTEMATICAL BOTANY.

88

are connected
646. Those peculiarities of structure which
is
a
which
in
developed are physiowith the manner
plant
logical.

connected
647. Those peculiarities of structure which are
structural.
are
are
which
in
manner
with the
arranged
parts
648. Physiological characters are of two kinds; 1, those

which are connected with the mode of growth (or organs of


and, 2, those which regulate reproduction (or
vegetation),

organs offructification).
649. Physiological characters are of greater importance in
of plants than structural.
regulating the natural classification
650. All modifications of either are respectively import-

with the
ant, in proportion to their connection

phenomena of

life.

allow ourselves to be steadily guided by these


considerations, we shall find that the internal or anatomical
651. If

we

structure of the axis,

and of the

foliage, is of

more importance

than any other character.


Because these are the circumstances which essentially regulate the functions of
growth, and the very existence of an individual.

652. That next in order

is

the internal structure of the

seed, by which the species must be multiplied.


of an embryo, or its absence, the first indicating a true
seed (531), the latter a spore (590), are most essential circumstances to conAnd so also the existence of albumen in abundance round the embryo,
sider.
or its absence, must be regarded as a physiological character of the highest
value : because, in the former case, the embryo demands a special external
provision for its early nutriment, as in oviparous animals ; while, in the latter
case, the embryo is capable of developing by means of the powers resident in

Thus the presence

itself,

and unassisted, as

in viviparous animals.

653. Next to this must be taken the structure of the organs


by whose united action the seed is engen-

of fructification,

dered

for

without

some

certain,

uniform,

able action on their part, the race of a plant

and invarimust become

extinct.
Thus we

find that the structure of the anthers, placenta?,

and ovules, are more


while their numbers are
;
and
the
condition
of
the filament, which appears of so little import;
ance in a physiological point of view, is also inconstant.
So also the texture
and surface and form of the pericarp, which acts as a mere
covering to the seeds,
is not to be regarded in these
in
differs
from
inquiries, and,
fact,
genus to genus ;
as, for instance, between Pyrus and Stranvaesia, or Rubus and
Spiraea, in the
truly natural Rosaceous order.

uniform than that of the parts surrounding them


variable

THE NATURAL SYSTEM.

On

654.

89

the other hand, of the floral envelopes (322), the

number, form, and condition, the" presence or absence, the


regularity or irregularity, seem to be unconnected with functions of a high order, and to be designed rather for the decoration of plants, or for the purpose of giving variety to the
aspect of the vegetable world ; they are consequently of low

and doubtful

value, except for specific distinctions.

There seems, indeed, reason to expect that every natural order will, sooner or
later, be found to contain within itself all the variations above alluded to.
Even in the cases of regularity and irregularity we already know this to be so ;
witness Veronica and Scoparia in Scrophul.iriaceae, and Hyoscyamus in
Solanaceae, Delphinium in Ranunculaceae, and Pelargonium in Geraniaceae.

The

consolidation of the parts of fructification is a circumstance but little attended to in a general point of view,
except in respect to the corolla ; but as it seems to indicate

655.

change that the parts can undergo, or,


occurs between important and unimportant organs,
that in such cases the latter are essential to the former, it proeither the greatest

where

it

bably deserves to be regarded with great attention.


For instance, the presence or absence of the corolla is often a point of little
moment, and is, we know, a very fluctuating circumstance. This is especially
true of those natural orders in which the stamens and petals are separated ; as

On the other hand, when the


Rhamnaceae, Onagraceae, &c.
stamens, which are indispensable organs, adhere to the petals, the latter are
more constantly present, as in Scrophulariaceae, Acanthaceae, Solanaceae, &c.
in Rosaceae,

656. If consolidation is, on the one hand, to be regarded as


a character of high importance, so must disunion also be so
considered on the other.
is indicated by those natural orders of plants, which, like the Rosaceous,
the Ranunculaceous, and the Magnoliaceous, are called apocarpous.

This

657. If

tremely

we descend lower than those points, we find it exwhen we enter into details, to comprehend

difficult,

what gives some of the subordinate


value

we

covered for judging of this


characters

No

assign to them.
is in

and

peculiarities of plants the


fixed rule has yet been disthe employment of secondary

a great degree arbitrary.

SYSTEMATICAL BOTANY.

THE NATURAL SYSTEM OF DE CANDOLLE.

IV.

MANY

natural

systems have been proposed by different

Ray, Linnaeus, Jussieu, De Candolle, Bartling,


Reichenbach, Schultz, Endlicher, myself, and many others,
botanists.

have each had their own system


racter that can be given of them

and, perhaps, the best chathat while they are all far

is,

from the truth, each has some merits which the others want.
The system of De Candolle, however, having been taken
as the basis of the most perfect enumeration of plants that has
ever been made, has so great a reputation, that for the conAnd it
venience of students it most requires explanation.
seems the more deserving of illustration, because the University of

London have declared that

conducted with reference to

be necessary to introduce into an illustration of

It will not
this

their examinations shall be

it.

system every natural order

known, and only

for

interest the botanist

many

are imperfectly

when he extends

inquiries into the minutiae of the science.

his

All, however, of

importance, will be found in the succeeding pages.


Plants are either furnished with visible flowers, or they
are multiplied in some other way.
Hence the two great diviof FLOWERING (Phanogamous or
Phanerogamous), and
FLOWERLESS (Cryptogamous)

sions,

EXOGENS (95) or ENDOGENS


Flowering plants
with
which
(95),
Dicotyledons (573) and Monocotyledons
are

either

(572) respectively correspond.


4.
Flowerless plants are either TETHEOGAMOUS
(Semivascular),
that is, furnished with stomates and vascular tissue ; or
they
are

AMPHIGAMOUS

(Cellular), that

is,

destitute of stomates

entirely cellular.

Hence

arise four CLASSES.

I.

Class

Class 2.

FLOWERING PLANTS.

Exogens or Dicotyledons.
Endogens or Monocotyledons.

and

EXOGEN^l THALAMIFLOR^:.

91

FLOWERLESS PLANTS.

II.

Class 3.

^Etheogamous or Semivascular.

Class

Amphigamous

4.

CLASS

or Cellular.

I.

EXOGEN.E.

the largest class in the vegetable kingdom, comprehending more species than all the others put together. The
subclasses are the following

This

is

Thalamiflora.

calyx and corolla.

Petals distinct

Petals distinct

Stamens hypogynous.
2.* Calycifloree.

calyx and corolla.

Stamens perigynous.

3.

Corottiflorte.

4.

Monochlamydea.

calyx
bearing the stamens.

SUBCLASS
I-

Order

climbing.

Petals

corolla.

united,

calyx only, or none.

J.

Ranunculaceee.

1.

and

THALAMIFLOIU3.

Herbs or shrubs, occasionally

Leaves with the petiole generally dilated, and the

blade very often palmate or digitate.


Sepals 3-6, usually
deciduous.
Petals 3-15, or none.
Stamens indefinite ; anCarpels numerous, or united into a single pistil.
Seeds either erect or pendulous.
USES.
Generally acrid, bitter, narcotic plants, with vesi-

thers adnate.

cating leaves, as Aconite, Stavesacre, Crowfoot. Some however


have the bitter principle predominant and the acridity slight,
as Hydrastis canadensis, Coptis, Xanthorhiza, which are
tonics.
* These are not
exactly the characters given by De Candolle, who includes all
monopetalous orders with an inferior ovary in Calyciflorae, and limits Corolliflorae to
But it seems to me more easy in practice to
the hypogynous monopetalous orders.
regard Corolliflorae as equivalent to the Monopetalae of Jussieu, while Thalamiflorse

and

of that

and MonochlamydejB

author,
Calyciflorae correspond to the Polypetalso
his Apefcihe ; and in a series so very artificial as this, we may
to consult convenience.

be pennitted,

to

think,

DE CANDOLLE'S SYSTEM.

92

This order divides into two principal sections.


TYPICAL GENERA. Ranunculus, CleFlowers

/.

regular.

matis, Adonis.

2. Ripe
1. Flower.
Ceratocephalus orthoceras.
4. Section of carpel and seed of the same.

fruit.

3.

Ovaries of Ranunculus

Krapfia.

//. Flowers

irregular.

TYPICAL GENERA.

Delphinium,

Aconitum.

Delphinium

2.

tricorne.

Anonacea.

without stipules.

Petals and stamens.

2.

Carpels.

3.

A branch

Leaves
and dull-coloured.

Trees or shrubs generally tropical.

Flowers

axillary, large,

of ripe fruit.

EXOGEN^E THALAMIFLORJE.
Sepals 8-4.

Stamens

93

Petals 6, coriaceous, with a valvular aestivation.


anthers adnate ; filaments angular.
;

indefinite

Ovaries numerous.
pels

Fruit succulent or dry, with the caror many-seeded, separate or consolidated.


Embryo

Albumen ruminate.

minute.

The fruits
Aromatic and fragrant in most cases.
of some are succulent and eatable, as the Custard Apple,
Anona squamosa, and the Cherimoyer, Anona Cherimolia;
USES.

those of others are hard, dry, and often jointed, as Habzelia


aromatica, the Piper ^Ethiopicum of the shops, and are used as
peppers.

Some

species are

employed as
Anona, Uvaria.

TYPICAL GENERA.

Anona

furfuracea.

1.

4.

expanded flower. 2. A vertical section oftheandroeforms a central and terminal tuft.


3. A vertical
vertical section of a ripe seed, showing the embryo and

An

ceum and gynoaceum, which


section of a carpel.

febrifuges.

latter

ruminated albumen.
3.
Menispermacea. Shrubs with a sarmentaceous habit.
Leaves alternate. Flowers small. Flowers unisexual, usually
Stamens monSepals in one or several rows.
very small.
Anthers turned outwards.
Ovaries
adelphous or distinct.
numerous, each with one style, sometimes soldered together
into a many-celled body, which is occasionally, in conse-

quence of abortion,

1-celled.

embryo curved, lying

in

albumen

Drupes
;

berried,

radicle superior.

1-seeded

DE CANDOLLE'S SYSTEM.

.94

USES.

Roots of many bitter and

tonic, as

Cocculus palma-

the Calumba root ; of others also diuretic,


tus, which yields
the latter a
as Cissampelos Pareira, and Cocculus Balds,
the negroes of Senegal against intermittents.
used

by

remedy

In the seeds a poison is formed, which in Anamirta Cocculus,


danthe Cocculus Indicus of the shops, becomes extremely
gerous.

TYPICAL GENERA.

Menispermum, Cocculus.

2. A female flower.
3. The vertical
1. A male flower.
which gradually curves the apex downwards, till, when it becomes
vertical section of a drupe, show5.
the drape 4, it acquires a horseshoe form.
ing the embryo and albumen ; a. is the true apex of the fruit, brought to the base

Cissampelos Pareira.

section of an ovary,

as just described.

Berberacea.

4.

Shrubs or herbaceous perennial plants.

SeLeaves alternate, compound, usually without stipules.


Petals sometimes with an
pals 3-4-6, in a double row.

Stamens equal in number to the


appendage at the base.
anthers opening elastically with
petals, and opposite to them
a valve from the bottom to the top.
Ovary solitary, 1;

Seeds attached to the bottom of the cell, 1, 2, or


albumen between fleshy and corneous.
USES. Bark astringent, and in Berberis yielding a
yellow

celled.

die.

Fruit of Berberis acid

TYPICAL GENERA.
5.

leaves,

Nympli<eacea.

growing

tubers of Bongardia eatable.

Berberis,

Epimedium.

Herbs with

in quiet water.

peltate or cordate fleshy

Sepals and petals imbricated,

EXOGENJE THALAMIFLOR^E.

95

Stamens numerous, inpassing gradually into each other.


serted ahove the petals into the disk ; filaments petaloid ;
disk large, fleshy.
Fruit many-celled.
Seeds very numerous,
attached to spongy dissepiments.
Embryo on the outside of
the base of the albumen, in a bag.
USES.
Of little moment. Euryale seeds are eaten. Rhi-

zomata slightly astringent and sedative.


TYPICAL GENERA. Nymphsea, Nuphar.
6.

Herbs with

Nelumbiacea.

Sepals 4 or

peltate,

floating

leaves.

Petals numerous. Stamens numerous.

5.

Disk

fleshy, enclosing in hollows of its substance the monospermous


ovaries.
Nuts numerous, half buried in the disk.

Nuts and creeping rhizomata eatable.


Nelumbium.

USES.

TYPICAL GENUS.

Dilleniaceae.
7.
Trees, shrubs, or under-shrubs, rarely
Flowers often yellow.
herbaceous, leaves without stipules.
Petals 5.
Stamens indefiSepals 5 ; 2 exterior, 3 interior.

"2.

nite.

Ovaries definite.

surrounded by a pulpy
USES.

Generally

Seeds

Carpels baccate or 2-valved.


aril.

Embryo
The
astringent.

in solid

albumen.

leaves of

many

species

are covered with asperities, which render them useful mechanically as polishing substances.
Nothing deleterious known

among them.

Flowers occasionally intolerably fetid.

TYPICAL GENERA.
[^

8.

Dillenia, Tetracera, Hibbertia.

Trees or shrubs with convolute

Magnoliacete.

Flowers

pules.

imbricated.

large,

Stamens

solitary.

Sepals

indefinite.

3-6.

Carpels

sti-

Petals 3-27,

numerous,

dis-

tinct or consolidated.

Bark tonic and febrifugal that of the root of Magglauca and Liriodendron in great repute in North

USES.
nolia

America.

Flowers often very fragrant.

TYPICAL GENERA.

Magnolia, Liriodendron.

Shrubs or small trees. Woody tissue


Leaves
alternate, dotted, with convolute deciduous
glandular.
often brown.
Petals 2-30,
Flowers
Sepals 2-6.
stipules.
Ovaries 1-celled with
imbricated.
Stamens hypogynous.
Fruit consisting of a single row
suspended or erect ovules.
9.

Winteraceee.

of carpels.

Seeds with or without

aril.

DE CANDOLLE'S SYSTEM.

96

Aromatic stimulants.

USES.

Illicium yields the Star

Winters Bark, of the shops.


Drimys Winteri, the

Anise, and

TYPICAL GENEKA.

Fumariea.

10.

An

Illicium,

Tasmannia.

Herbaceous plants with


Petals 4

Sepals 2.
outer one, or both saccate at the base.

and a watery

juice.

brittle

stems

parallel

Stamens

6,

the
in

parcels.

Unimportant. Species slightly diaphoretic.


TYPICAL GENERA. Fumaria, Corydalis.
USES.

Fumaria
3.

The

anthers.

11.

flower seen from below.


2. The same from the side.
1.
4.
stamens, and a portion of the bagged upper sepal.
parcel of

officinahs.

pistil,
5.

The

fruit.

Sarraceniacea.

Herbaceous perennial plants, living

Leaves with a hollow urn-shaped petiole. Scapes


one
Petals 5,
bearing
large flower.
Sepals 5, imbricate.

in bogs.

97

TITALAMIFLORJE.
unguiculate, concave.
5-celled

Stamens

indefinite,

hypogynous.

Ovary

Capstigma very large, umbrella-shaped, peltate.


sule crowned by the stigma.
Seeds very numerous, minute.
USES.
Unknown. Petiole-like leaves remarkable.
;

TYPICAL GENUS.

Sarracenia.

Herbaceous plants ; rarely


Flowers without bracts.
Petals 4, cruciate.
Stamens

Brassicaceee or Crucifera.
Leaves alternate.

12.

under-shrubs.

Sepals 4, deciduous, cruciate.


6, of which two are shorter (tetradynamous).
Ovary superior, with parietal placentae, meeting in the middle, and forming a

spurious

dissepiment.

Fruit a silique

or

silicule.

Seeds attached by a funiculus, generally pendulous.


with the radicle folded upon the cotyledons.

Embryo

A very large and difficult natural order, the subdivisions


which are now made to

in

depend upon the structure


of the embryo.

They

are the following

when

1. Pleurorhizea,

the

embryo has the

dicle applied to the

of the cotyledons

ra-

edge

fig. 17.

when
NotorMzeez,
has
the
radiembryo

2.

the

cle applied to the

the cotyledons

back of

Jig. 14.

Orthoplocea, when
embryo has the radicle applied to the back
of cotyledons which are
3.

the

hollowed out;
4.

fig. 12.

when

Diplecolobea;,

the cotyledons are three


times folded, and the radicle
|)*1OK

applied
"

'

H.fl
"*

their

'

USES.
!

to

1 fi

Erucastrum Canariense.

All the Species

harmless; Some

ailtlSCOr-

The

stamens.

3.

The

1.

siliqua,

flower.

2.

with the valves

4.
transverse
separating from the replum.
5.
of a seed.
perfect seed.

I!

DE CANDOLLE'S SYSTEM.

98

more or less pungent. Radishes, Turnips, Mustard,


are wellCabbage and all its varieties, Rape, Charlock,

butic, all

Cress,

known

plants of the order.


Brassica, Sinapis, Draba.

TYPICAL GENERA.

6. Siliqua of Mathiola livida.


7. Silicula of Thlaspi
Fruits of various genera.
9. Silicula of Schiverukia podolica.
8. Silicula of Alyssum spatlmlatum.
13. Silicula
10. Silicula of Farsetia.
12. Seed of Didesmus -/EgjT>tius cut across.
of Menonvillea linearis.
14. Seed of Lepidium Africanum.
15. Silicula of jEthio-

latifolium.

nema

cristatum.

16.

Seed of Heliophila

18. Siliqua of Mathiola oxyceras.


Silicula of Senebiera serrata.

ceras.

13.

crithmifolia.

Seed of Mathiola oxyDidesmus -(Egyptius. 20.

17.

19. Silicula of

Herbaceous plants or shrubs with a

PapaveracefE.

milky juice. Leaves alternate, without stipules. Sepals 2.


Petals either 3 or 4, or some multiple of that number.
Stamens hypogynous, generally numerous.
Fruit 1-celled,
with parietal placentae.
Seeds numerous.
USES.
A narcotic milk pervades the species ; that of
The
Papaver somniferum becomes opium when inspissated.
roots of Meconopsis
are
a
SanNepalensis
deadly poison.
guinaria Canadensis is emetic and purgative in large doses,
stimulant and diaphoretic in smaller.

TYPICAL GENERA.
14.

Papaver, Glaucium.

Capparidacece.

without true stipules.


cruciate.

Stamens

Herbaceous plants, shrubs, or trees,


Leaves alternate. Sepals 4. Petals 4,

definite or indefinite.

Disk hemispherical,

EXOGENJE THALAMIFLORJE.
or

Fruit

stalked.

Ovary

elongated.

99

most

1-celled,

fre-

quently with two polyspermous placentae ; embryo incurved.


USES.
pungent principle exists in some, as the flowerbuds of Capparis spinosa, which are the Capers of shops, and

several Cleomes

used

as

sometimes so

is

acridity

The

species dangerous.
blister like

substitutes

much

for

This

mustard.

concentrated as to render the

root of Cratseva gynandra

said to

is

Cantharides, and that of Cleome dodecandra

is

used as a vermifuge.
TYPICAL GENERA. Cleome, Capparis.

Physostemon lanceolatum.
mens, and ovary.

The same

15.

small

3.

cut vertically,

1.

A flower

of the natural size.

2.

with one valve separating.


to show the incurved embryo.

The

ripe fruit,

The
4.

calyx, sta5.
seed.

Herbaceous plants with alternate leaves,


and gland-like stipules.
Calyx
Petals lacerated, unequal. Disk large, 1-sided.

Resedacea.
colourless

many-parted.

Stamens

definite,

flowers,

inserted

into

the

disk.

Ovary

sessile,

H 2

DE CANDOLLE'S SYSTEM.

100
3-lobed,

1 -celled,

many-seeded,

with 3 parietal

placentae.

Fruit opening at the apex.


Embryo incurved.
well
known for its fragrance,
so
USES.
Mignionette,

is

Reseda odorata. R. luteola yields a yellow dye.


TYPICAL GENERA. Reseda, Ochradenus.
Shrubs or

Flacourtiacea.

16.

Leaves alternate,

trees.

Petals equal to them


Stamens occasionally changed into nectariferous
in number.
scales.
Ovary roundish ; stigmas several, more or less dis-

without stipules.

Fruit

tinct.

Sepals from 4-7.

1-celled,

capsular or

fleshy,

the centre

filled

with a thin pulp. Seeds few, attached to the lining of the


fruit in a branched manner.
Embryo in albumen.
USES.

The

fruit

TYPICAL GENERA.
Bixacete.

17.

of some eatable and pleasant in India.


Flacourtia,

Roumea.

Trees or shrubs.

Leaves alternate, with

deciduous stipules and pellucid dots.


Sepals 4-7, imbricated.
Petals of a like number. Stamens indefinite, distinct.
Ovary
sessile

placentae 4-7, parietal

styles 1-2-4.

fleshy or capsular, many-seeded.

Fruit 1-celled,

Seeds enveloped in pulp.

Albumen hardly present.


USES.
The seeds of Bixa Orellana are covered with a
pulp, which, when dry, is the Arnotta of shops, used for
Otherwise the properties uncertain.
TYPICAL GENERA. Bixa, Prockia, Azara.

colouring cheese.

18.

Cistacea.

Shrubs

or

herbaceous

Leaves

plants.

usually entire, stipulate or exstipulate.


Sepals 3 or 5, persistent, unequal, in a broken whorl, the three inner twisted.
Petals 5, often crumpled, twisted in a direction

contrary

to that of the
Stamens indefinite.
sepals.
celled ; ovules with their foramen at their

stigma simple.
or

1- or

many-

apex ; style single ;


Fruit either 1-celled with
parietal placentae,

imperfectly 5- or 10-celled.

inverted, either spiral or curved,

bumen.

Ovary

Seeds indefinite.
in the midst of

Embryo
mealy

al-

Radicle remote from the hilum.

USES.
Unimportant. The balsamic Gum Ladanum is a
spontaneous secretion from Cistus Creticus and others.
Many
are beautiful garden
plants, with large delicate flowers.
TYPICAL GENERA. Cistus, Helianthemum.

EXOGEN.E THALAMIFLOR.E.

Cistus Berthelotianus.
2.

off.

4.

1.

A calyx

and

cross section of the ovary.

seed cut through

pistil,

3.

101

the petals and stamens having fallen


of ovary and calyx.

vertical section

the pointed end being the true apex.

Herbaceous plants, often covered with


Leaves
with
glands.
stipulary fringes and a circinate vernation.
Peduncles circinate. Sepals imbricate. Petals 5, hy19.

Droseraceae.

pogynous.

Stamens

petals, or 2,
3-5 valves.

3,

USES.

distinct, either equal in

or 4 times as

many.

number

Styles 3-5.

to the

Capsule of

Embryo in fleshy or cartilaginous albumen.


The herbage of some Droseras is acrid. The bulbs

of others abound in a rich purple dye, and are filled with


It is probable that
starch, which renders them eatable.
would
of
value
to
many species
prove
dyers.

TYPICAL GENERA.

Drosera, Dionsea.

20.
Tamaricacea. Shrubs or herbs, with rod-like branches.
Leaves alternate, resembling scales. Calyx 4- or 5-parted,

Petals withering.

persistent.

Stamens

definite, distinct, or

monadelphous. Stigmas 3. Capsule 3-valved, 1-celled, manyseeded.


Seeds ascending, comose ; embryo straight.
USES.

Ornamental bushes or

resembling

Manna oozes

trees.

sweet substance

out of the stem of Tamarix Gallica, in

The bark

is bitter, astringent, and tonic.


employed in medicine and dyeing, in
India,
yielded by some oriental species.
TYPICAL GENERA. Tamarix, Myricaria.

hot,

dry countries.

very astringent
i.s

gall,

DE CANDOLLE'S SYSTEM.

102
21.

Leaves

Herbaceous plants, or shrubs, or trees.


with an involute vernation. Sepals 5, perPetals 5, regular or irregular, one someimbricate,
Violaceee.

stipulate,

sistent,

Stamens definite in number ; filaments ditimes spurred.


lated ; connective elongated beyond the anthers.
Ovary
with 3 parietal placentae style with a hooded stigma.
Capsule of 3 valves, bearing the placentae in their axis.
Embryo large, straight, in fleshy albumen.
1-celled,

USES.
Roots emetic. Those of the common Sweet Violet
and other species have been employed medicinally.
lonidium Poaya yields one sort of Brazilian Ipecacuanha.
Viola canina and some others have the power of removing
some cutaneous affections, and have been employed as cosmetics.

TYPICAL GENERA.

Viola, Alsodeia.

1.
Corynostyhs Hybanthus.
set of stamens, each
having the connective lemrthI
beyond the anther in the form of a scale.
2.
3.
transspurred petal.
thlee Parietal
Tary '
WiDg
4'
rie frrit- 5 An
Placen

22.Polygalacea.
alternate,

which 1

'

Shrubs or herbaceous
Leaves
plants.
Pedicels with three bracts.

destitute of stipules.

lowers unsymmetrical.
Sepals
Petals consolidated,

laceous.

5,

very irregular, often glu3, of

hypogynous, usually

is anterior and
larger than the rest.
Stamens usually
tube; anthers innate,
1-celled, and opening at their apex.

103

EXOGEKLffi THALAMIFLOR^l.

Ovary with 2 or 3 cells ; ovules solitary, pendulous. Seeds


pendulous, with a caruncula next the hilum ; albumen abundant.
USES.
Leaves bitter, root milky. Polygala Senega, the
Rattlesnake root, is stimulant, diaphoretic, emetic, and emmenagogue ; it has been employed successfully in croup. Many
other species have similar properties.
Polygala Poaya is one
of the Brazilian emetics.
The bark of Monnina polystachya,

a Peruvian plant,

is

detersive,

and used as a

substitute for

soap.

Polygala, Mural tia, Mundia.

TYPICAL GENERA.

1. An entire flower seen from the side.


2. The same cut
Polygala erioptera.
section of a ripe seed ; in the
open to exhibit the stamens. 3. The pistil. 4.
middle is the embryo ; at the apex, which represents the real base, is seen a carun-

cula.

Herbaceous plants or under-shrubs.


Leaves small, opposite, exstipulate,
with a membranous sheathing base. Sepals 4-5, in a furrowed
tube.
Petals hypogynous, unguiculate, with appendages at
FranJceniacea.

23.

Stems much branched.

Stamens definite.
Style 2- or 3-fid.
enclosed in the calyx, 2- 3- or 4~valved, manySeeds attached to the margins of the valves, very

the base of the limb.

Capsule
seeded.

minute

USES.

1 -celled,

embryo

in the

midst of albumen.

Unknown.

TYPICAL GENUS.

Frankenia.

DE CANDOLLES SYSTEM.

104

Elatinaceee.

24.

with

site,

stipules.

Stamens

hypogynous.

weedy

Lijttle

definite.

Sepals 3-5.
3-5-celled;

Ovary

Petals
styles

Seeds numerous,

Fruit capsular.
stigmas capitate.
little albumen.
with
but
straight,
embryo
3-5;

Unknown.

USES.

TYPICAL GENERA.
25.

Leaves oppo-

annuals.

Flowers minute.

Elatine, Bergia.

Herbaceous plants

CaryopTiyllaceae.

with

undivided exstipulate leaves, and tumid nodes.


Petals often

Stamens

slit.

definite.

opposite
Sepals 4-5.

Ovary usually many-

seeded, with a free central placenta.


Stigmas sessile, 2-5.
2-5-valved.
with
Seeds
the embryo curved
Capsule
usually

round mealy albumen.


USES.

Some

Unimportant.

bear gay flowers;

species

more are weeds; a few are


is

fragrant, as the Pink.


said to have an anthelmintic root.

Virginica
There are two sections of this order
1.

Alsinea.

Silene

Sepals disjoined.
Stellaria, Cerastium.

TYPICAL GENERA.
2.

Sepals united into a tube.

Silenete.

TYPICAL GENERA.

iun a
1

vrtv

o
rtically, to

PW\

show

Lychnis, Silene, Dianthus.

2 A flower of Stellaria
f *?**'
US Placenta loaded with seeds
-

J
the

mcdi <%

5 A
\
embryo curved round mealy albumen'
-

'

*<*-

3.

through

EXOGEN.K THALAMIFLOR.E.
26.

Malvacea.

Herbaceous

plants,

105

trees,

or

shrubs.

Leaves alternate, stipulate, very often covered with stellate


hairs.
Flowers generally showy.
Calyx with a valvate
aestivation.

Petals twisted.

Stamens

indefinite,

monadelph-

anthers 1-celled, reniform.


Ovary formed by the union
of several carpels ; styles the same number as the carpels.

ous

Fruit either capsular or baccate ; albumen in small but variable quantity; embryo curved, with twisted and doubled
cotyledons.
USES.
Mucilaginous

low.

The unripe

fruit

as

Marsh Mallow and common Mal-

of Hibiscus esculentus

is

used as an

The liber of several affords a tenacious


ingredient in soups.
fibre ; the hemp-like substance called Sun in India is obtained
from

Hibiscus

The hairy

cannabinus.

seeds of

Many

are

beautiful

objects.

furnish cotton.

Gossypium
TYPICAL GENERA. Malva, Lavatera, Hibiscus.

Abutilon macropodum.

1.

An

unexpanded flower. 2. The stamens and styles.


whose upper extremities are free and ra-

ripe fruit, consisting of many carpels,


section of a seed.
diant.
4.
3.

27.
plants.

Tiliaceae.

Leaves

Trees or shrubs, very seldom herbaceous


Flowers often small.
alternate.

stipulate,

Calyx valvate. Petals 4 or 5, usually with a little pit at their


Stamens distinct; anthers 2-celled.
Ovary single,
composed of from 4 to 10 carpels; style one; stigmas as
many as the carpels. Seeds several ; embryo erect in the
axis of fleshy albumen, with flat foliaceous cotyledons.

base.

DE CANDOLLE'S SYSTEM.

106

The leaves
fibres.
SES
Mucilaginous plants with tough
Corchorus capof Corchorus olitorius are eaten as spinach.
From the
sularis furnishes a kind of coarse hemp in India.
Russia mats are made; its
inner hark of Tilia

Europsea
from the hracts, are said to be antispasmodic.
TYPICAL GENERA. Tilia, Triumfetta, Grewia.

flowers, separated

Leaves alternate, with involute


Trees.
28.
Dipteracea.
vernation; stipules deciduous.
Calyx 5-lobed, unequal, beStamens
Petals contorted.
coming enlarged, imbricated.
indefinite, distinct

towards the apex.


in pairs,

anthers

subulate, opening longitudinally


disk, few-celled ; ovules

Ovary without a

pendulous; style

single.

Fruit surrounded by a

having tough, leafy, enlarged, permanent divisions.


Seed single, without albumen. Cotyledons crumpled.
that
USES.
Tropical trees often yielding valuable timber ;

calyx,

called Sal, or Saul, belongs to

The

Shorea robusta.

juice

is

Sumatra Camphor.'

balsamic; Dryobalanops Camphora yields


Dammer pitch
Vateria Indica furnishes Copal.
from species of Shorea.
'

TYPICAL GENERA.
29.

and

Aumntiacea.

filled

is

obtained

Shorea, Dipterocarpus, Vateria.

Trees or shrubs, almost always smooth

with transparent receptacles of

oil.

Leaves

alter-

compound, always articulated with the petiole.


Flowers usually white or green, and fragrant. Calyx urceo-

nate, often

late

or campanulate, short.

Petals 3-5, inserted upon the

outside of an hypogynous disk.


Stamens inserted upon an
filaments
sometimes
combined in one or
disk;
hypogynous
several parcels.

Ovary many-celled ; style 1 ; stigma thickFruit many-celled, filled with pulp.


Seeds usually pendulous ; raphe and chalaza distinctly marked.

ish.

USES.
The Orange, Lemon, Lime, and Citron are species
of Citrus, and are well known for the aromatic rind and
pulpy
flesh of their fruit.
The wood is generally hard and durable.
The unripe fruit of ^gle marmelos, an Indian tree, is prescribed in diarrhrea and
The leaves of the order
dysentery.
generally are regarded as stomachic and tonic.

TYPICAL GENERA.

Citrus, Triphasia, Limonia.

SQTernstromiacea.

Trees or shrubs.

Leaves alternate,

EXOGENJE THALAMIFLOR^l.

107

Flowers
stipules, now and then with pellucid dots.
often large and showy.
Sepals 5 or 7, coriaceous, in a broken
Petals not equal in number to the sepals.
whorl, deciduous.
without

Stamens numerous
with

several

monadelphous or polyadelphous.

cells

styles

Ovary

filiform.

usually with a central column.

Capsule 2-7-celled ;
Seeds large, attached to the

very few ; albumen none ; cotyledons occasionally plaited.


USES.
The Tea of Commerce consists of the leaves of Thea

axis,

viridis

and Bohea.

Camellia oleifera yields excellent

oil.

species of Camellia, common in gardens, are objects of


Leaves of Kielmeyera speciosa are mucilaginous.
beauty.

The

TYPICAL GENERA.

4.

Kielmeyera rosea.
An embryo.

31.

1.

The

pistil.

2.

A transverse

section of

it.

3.

A ripe fruit.

Herbaceous plants, shrubs or trees.


Flowers geneentire, sometimes dotted.

Hypericacea.

Leaves opposite,
rally

Camellia, Gordonia, Thea.

yellow.

Sepals 4-5,

persistent,

imbricated,

unequal,

with glandular dots. Petals 4-5, hypogynous, twisted, oblique,


often having black dots.
Stamens indefinite, often polyaFruit a capsule or berry, of many
delphous.
Styles several.
valves and many cells.
Seeds minute, indefinite ; embryo
straight,

USES.

with no albumen.

The

juice

is

resinous,

purgative, febrifugal or asan essential oil or a

tringent in different species, according as

DE CANDOLLE'S SYSTEM.

108

resemabound. The latter, when concrete,


yellow juice most
the
has
Hypencum
properties.
bles gamboge, of which it
vulneranes.
Androsjemum and perforatum are old-fashioned
for a gargle
In Brazil a species of Hypericum is employed
in cases

of sore throat.

TYPICAL GENERA.

Hypericum, Vismia.

3. A
2. A bundle of stamens.
1. An entire flower.
A seed laid horizontally and cut through, to show the em5. A piece of a leaf with transparent dots.

Hypericum floribundum.
pistil

with 3 carpels.

bryo and netted

4.

testa.

Leaves
Trees or shrubs.
Guttifera.
Flowers sometimes
opposite, coriaceous.
Petals hypogynous,
Sepals 2 to 6, persistent.

Clusiacea or

32.

without stipules,

polygamous.
4 to 10. Stamens numerous, hypogynous.

Disk

fleshy, occa-

sionally 5-lobed.
Ovary 1- or many-celled ; ovules solitary,
or
erect,
ascending, or numerous and attached to central pla-

Seeds
style very short ; stigma peltate or radiate.
in
often
none.
with
an
aril
albumen
;
pulp,
frequently nestling
centae

USES.

The

Gamboge is the juice

delicious

Malacca

of Hebradendron cambogioides.
Mangosteen is the produce

fruit called

of Garcinia mangostana.
The resinous oil Tacamahaca flows
from the root of Calophyllum Calaba. The general properties
of the species are acrid and purgative.
of great beauty on account of their

some thick

They

leaves.

TYPICAL GENERA.

are often objects

large flowers

Clusia, Garcinia.

and hand-

EXOGEN^E THALAMIFLORJE.

109

Hebradendron cambogioides. 1
female flower, with the sterile stamens surrounding the pistil. 2. A male flower. 3. An anther, which opens by throwing off
a cap, in consequence of transverse dehiscence. 4.
transverse section of the ovary.
.

Leaves opposite, without stipules.


Flowers unsymmetrical.
Petals
inserted
round an hyimbricated.
Calyx
an
disk.
Stamens
inserted
upon
pogynous
hypogynous disk,
33.

Aceracea.

Trees.

Flowers small, green, often polygamous.

Fruit of 2 parts,
generally 8.
Ovary 2-lobed; style 1.
which are samaroid ; each 1 -celled ; with one or two seeds ;

albumen none.

The

USES.

kind of sugar
is light,

saccharine sap of Acer saccharinum yields a


North America. The timber of most species

in

and

clean,

useful,

TYPICAL GENUS.
34.

where strength

large,

Leaves opposite, withterminal. Flow-

Trees or shrubs.

^Escnlaceee.

out stipules, quinate or septenate.


ers

not required.

is

Acer.

Racemes

Flowers unsymmetrical.
4 or 5, unequal,

showy.

Calyx

cam-

Petals

5-lobed.

hypogynous.
Stamens 7-8, unequal. Ovary 3-celled ; ovules 2 in each cell.
Fruit 1- 2- or 3-valved.
Seeds large, with a broad hilum ;

panulate,

albumen none

embryo curved, germinating under ground.

Handsome
which renders them
USES.

trees or bushes. Seeds filled with starch,

nutritious

also dangerous.

TYPICAL GENUS.

^Esculus.

but

it

is

said that they are

DE CANDOLLE'S SYSTEM.

110

Small trees or shrubs, sometimes


MalpigUacea.
with stipules. Sepals generally
Leaves
opposite,
climbing.
on the outof
with 5
oblong conspicuous glands
35.

large

pairs

side.
1,

seldom fewer.

Stamens
unguiculate.
less combined
or
more
carpels,

Petals

of S

5,

Ovary

ovules suspended.

Seeds without albumen.


Fruit dry or berried.
Of no moment. The fruit of some Malpighias is
USES.
eaten in the West Indies under the name of Barbadoes
cherries.

The bark appears

and samaroid,

2.

36.

1.

3.

Sapindacete.
tendrils.

Fruit dry

Banisteria.

Diplopteris paralias.
An expanded flower.

have

to be astringent.

Fruit succulent, Malpighia.

TYPICAL GENERA.

flower-bud, showing the double glands of the calyx.


4. Ripe fruit of Ryssopteris timorensis.
carpels.

The

Trees, or shrubs which often climb

Leaves generally compound.

and

Flowers un-

Petals hypoCalyx imbricated.


sometimes
sometimes
with
a
doubled
naked,
gynous,
appendsymmetrical, polygamous.

Disk fleshy. Stamens 8-10,


the
cells containing 1, 2, 3,
rarely
Ovary 3-celled,
seldom
ovules.
Fruit
sometimes
more,
very
capsular, sometimes samaroid, sometimes fleshy and indehiscent.
Seeds

age in the inside, imbricated.


5-6-7.

Albumen 0.
usually with an aril.
USES.
Leaves and branches of

some species of Magonia


and Paullinia poisonous.
The fruit of some Euphorias or
Nepheliums, Pierardias and Hedycarya, eatable and agreeable ; the former is the
Longan and Litchi, which occasionally

EXOGEN^E THALAMIFLOR.E.

Ill

The fruit of Sapindus


arrive in this country from China.
of soap.
others
instead
and
employed
saponaria
TYPICAL GENERA.

Sapindus Senegalensis.
after fertilization.

4.

Sapindus, Paullinia, Serjania.

1.

An

expanded flower.

A vertical section of a ripe

2.

petal.

3.

The

ovaries

drupe, showing the embryo.

Trees with timber which is usually comand beautifully veined. Leaves alternate, without stipules. Calyx 4-5-cleft. Petals 4-5.
Stamens 8-10,
Seeds
either united or distinct.
Style and stigma simple.
37.

Cedrelaceee.

pact, scented,

flat-winged.

USES.

Mahogany is the timber of Swietenia Mahagoni

the

of Cedrela Toona, and Soymida febrifuga, is


valuable as a tonic, in careful hands ; it can only be exhibited

bark of that

tree,

in small doses.

East India Satin-wood

oxylon Swietenia.
TYPICAL GENERA.

is

Cedrela, Swietenia.

produced by Chlor-

DE CANDOLLE'S SYSTEM.

112

Leaves alternate,
without stipules.
numerous,
Calyx 5-parted.
monadelphous ; anthers with a fleshy connective extended
beyond the lobes. Ovary 5-celled; ovules 1-2, suspended;
Fruit drupaceous. Embryo in fleshy albumen.
styles simple.
Trees or shrubs.

Humiriacea.

38.

Stamens

USES.
The liquid yellow fragrant Balsam of Umiri flows
from the wounded trunk of Humirium floribundum. In properties

it

resembles Copaiva.

TYPICAL GENUS.
39.

Leaves alternate, with-

Trees or shrubs.

Meliacete.

out stipules.

Humirium.

Sepals

3, 4,

Petals hypogynous, usually


as many as the petals; filaments
or 5.

Stamens twice
cohering in a long tube ; anthers sessile within the orifice
of the tube.
Ovary with 3, 10, 12 cells; ovules suspended,
valvate.

1-2 in each cell.


Fruit often 1-celled.
Seeds without albumen, not winged.
USES.
The bark of Guarea Aubletia, Trichilia cathartica,
and others, purgative and emetic. Eoot of Melia Azedarach
anthelmintic.
Some of the tropical genera have a wholesome
pleasant fruit.
Febrifugal qualities have been recognized in
the Neemtree, Melia Azedarachta, and some others.
TYPICAL GENERA. Melia, Quivisia.

Ekebergia Senegalensis.
averse section of the

1.

ovary.

flower.

4.

2.

The calyx and

ripe fruit.

5.

staminal tube.

3.

vertical section of the latter.

EXOGEN-E THALAMIFLOR^:.
40.

Vitacece.

separable joints.

113

Scrambling, climbing shrubs, with tumid


Leaves with stipules.
Flowers small,

Calyx small, nearly entire. Petals in aestivation valand often inflected at the point
stamens opposite
them, inserted upon the disk.
Ovary 2-celled ovules erect,
definite.
Berry pulpy albumen hard. Embryo small.
green.

vate,

The common Vine,

USES.

Vitis vinifera,

is

well

known

besides which there are other species, in which reside similar


The leaves of some kinds of
qualities, although very inferior.

being acrid, are used in bringing indolent tumours

Cissus,

to suppuration.

TYPICAL GENERA.

Vitis, Cissus,

41.

Strong-scented herbs or shrubs with

Geraniacea.

Ampelopsis.
sti-

Stems tumid, and separable at the joints.


ribbed.
Stamens definite,
Petals 5, unguiculate.
Sepals 5,
often monadelphous.
Fruit of 5 elastic cocci, rolling back
from a long-beaked gynobase, to which the hardened styles
pulate leaves.

Seeds solitary, pendulous, without albumen. Cotyledons convolute and plaited.


USES.
The root of Geranium maculatum is a powerful

adhere.

astringent.

Otherwise the order

is

of no importance, except

Geranium sylvaticum. 1. The stamens and style. 2. The unripe fruit surrounded
by a calyx. 3. The rostrate gynobase, from which the cocci are rolling back with
elasticity

one has dropped

off.

4.

transverse section of a seed.

DE CANDOLLE'S SYSTEM.

114

for the beautiful flowers of

numerous

species,

especially be-

longing to the genus Pelargonium.

TYPICAL GENERA.
42.

Geranium, Erodium, Pelargonium.


Succulent herbaceous plants.

Balsaminaceae.

Leaves

Flowers usually unsymmetrical.

Sepals 5,
without stipules.
~
StaPetals
4,
odd
the
irregular.
sepal spurred.
irregular;
mens 5. Carpels consolidated into a 5-celled ovary. Fruit
5 elastic valves.
capsular, with

suspended

Seeds solitary, or numerous,

albumen none.

They have generally gay flowers.


Unimportant.
TYPICAL GENERA. Balsamina, Impatiens.
USES.

Linacea. Herbaceous plants or small shrubs. Leaves


43.
Flowers
Petals fugitive.
without stipules, usually alternate.
Petals
imbricated,
3-4-5,
persistent.
Sepals
symmetrical.
united
Stamens
in
a
twisted.
ring.
hypogynous, unguiculate,

Ovary with about


number to the cells

as
;

many

cells as sepals.

stigmas capitate.

Styles equal in

Capsule many-celled.

albumen present.
cell single, inverted
The mucilaginous seeds of Linum usitatissimum
The leaves of L. catharticum are purgative. The
linseed.

Seeds in each

USES.
are

tough fibre of the first is the Flax of manufacturers.


TYPICAL GENERA. Linum, Radiola.
44.
Oxalidacea:, Herbaceous plants, under-shrubs, or trees.
Leaves alternate, compound, often sensitive. Sepals 5, equal.

Petals equal, unguiculate.


Stamens 10, more or less
adelphous.
Styles 5 ; stigmas capitate. Fruit capsular,

mon-

mem-

branous, with 5

cells.
Seeds few, within a fleshy integument,
which expels the seeds with elasticity.
Embryo long, taper.
Albumen between cartilaginous and fleshy.

USES.

They

are generally acid in a


high degree.

Blimbing and Carambola, acid

The

of the Indian ArchipeThe roots of


lago, are the produce of the genus Averrho.a.
Oxalis Deppei form an
esculent.
agreeable

TYPICAL GENERA.
45-

Pittosporaceee.

fruits

Oxalis, Averrhoa.

Leaves simple, alternate, without

sti-

Shrubs, trees, or half herbaceous plants, sometimes


twining.
Petals hypogynous,
Sepals deciduous, imbricated.

pules.

EXOGENJE THALAMIFLORJE.
imbricated.

Stamens

5.

capsular or berried, with

115

Ovary single, many-seeded. Fruit


many-seeded cells which are some-

Albumen fleshy.
times incomplete.
USES.
Unimportant. The species are resinous.
TYPICAL GENERA.
46.

Rutacea.
dotted.

stipulate,

aphrodite,

sionally

Trees or shrubs (or herbs). Leaves exFlowers often very gay.


Flowers herm-

sometimes irregular.

times combined.
like disk.

Pittosporum, Sollya, Billardiera.

Stamens

Ovary few-celled ;
divided near

component parts

the

as the

Petals someSepals 4-5.


on the outside of a cupovules 2-4 ; style single, occa-

definite,

base,

fruit

always separable into

approaches maturity.

capsular, separating into carpels when


or without albumen ; radicle superior.

ripe.

its

Fruit

Embryo with

a cupEriostemon myoporoides.
1.
complete flower. 2. The ovary, seated in
3. The ripe fruit, separated spontaneously into
shaped disk, surrounded by a calyx.
its component carpels.
4.
vertical section of a seed, showing the embryo lying in
the midst of albumen.

i2

DE CANDOLLE'S SYSTEM.

116

U SES

The powerfully scented oil possesses active properRuta graveolens, common Rue, is anthelmintic, sudo-

ties.

Various species of Barosma, called


at the Cape of Good Hope Bucku, are powerful antispasmoThe bark of Cusparia febrifuga, called Angostura bark,
dics.

rific,

and emmenagogue.

a valuable febrifuge ; and that of many other American


trees of the order seems to possess the same quality.
Correa is
TYPICAL GENERA. Ruta, Boronia, Dictamnus,
remarkable for having a monopetalous corolla.
is

Leaves without
Xanihoxylacea. Trees or shrubs.
unisexual.
Flowers
dots.
with
Calyx in 3,
pellucid
stipules,
Petals usually longer than the calyx, convo4, or 5 divisions.
47.

Stamens equal to the petals in number, or twice as


many. Ovary of the same number of carpels as there are

lute.

a smaller number

petals, or

ovules 2

styles

more

or less

Fruit berried or membranous, sometimes consistSeeds solitary or


ing of several drupes or 2-valved capsules.

combined.

and shining; embryo lying


within fleshy albumen ; radicle superior.
USES. Aromatic, pungent, and stimulant.
Xanthoxylum
twin, pendulous, usually smooth

Clava Herculis is a powerful sudorific and aperient.


The
bark of Brucea, of Xanthoxylum caribeeum, and others, is
The capsules of some Fagaras are used as pepper.
febrifugal.
TYPICAL GENERA. Xanthoxylum, Ptelea.
48.
Herbaceous plants, shrubs, or trees ;
Zygophyllacea.
branches often articulated at the joints.
Leaves opposite, with
stipules,

not dotted.

Flowers hermaphrodite.

Calyx con-

Petals unguiculate.
Stamens dilated at the base,
sometimes placed on the back of a small scale.
Ovary with a
volute.

and 4 or 5

disk,

simple.

Seeds few
USES.

cum

cells

ovules pendulous or erect

albumen whitish.
Zygophyllum Fabago is- an anthelmintic.
;

radicle superior

yields the

wood

for its hardness,

and

called
in

TYPICAL GENERA.
49.

style

Fruit capsular, rarely fleshy, with


angles or wings.

Simamlacea.

Lignum Vit, known

medicine for

Guaia-

in turnery

its sudorific
qualities.

Zygophyllum, Guaiacum.
Trees or

shrubs.

Leaves without

EXOGEN.E THALAMIFLORJE.
stipules, alternate,

without dots.

unisexual.

in

117

Flowers hermaphrodite, or
4 or 5 divisions. Petals longer ; aesti-

Calyx
Stamens arising from the back of an hyposcale.
gynous
Ovary 4- or 5-lobed, upon a stalk, each cell
with 1 suspended ovule ; style simple.
Fruit indehiscent
without
albumen.
embryo
vation twisted.

The wood

USES.

intensely bitter.

The

root of

Simaruba

amara, used as a tonic, is bitter, purgative, and emetic. The


wood of Picraena excelsa furnishes the Quassia chips of the shops.
TYPICAL GENEKA. Quassia, Simaruba.

Shrubs with square branches. Leaves


Flowers herbaceous, hermPeaphrodite, monoecious or direcious.
Calyx 5-parted.
tals 5, fleshy, with an elevated keel.
Stamens 10. Ovary
Ovules
5-celled, 5-angled ; style none ; stigmas 5, subulate.
50.

Coriariacea.

opposite, simple,

entire, ribbed.

Carpels 5, drupaceous, indehiscent, 1-seeded, someAlbumen none.


times surrounded by the enlarged petals.

solitary.

The

USES.

fruit

of Coriaria myrtifolia

leaves are used for dyeing black,


drian Senna.

TYPICAL GENUS.

and

is

poisonous

for adulterating

the

Alexan-

Coriaria.

SUBCLASS

II.

CALYCIFLOR*:.

trees. Leaves simple. Flowers in axillary cymes, minute.


Sepals 4 or 5, imbricated, inserted into the margin of an expanded torus.
Petals imbri-

51.

cate.

disk.

Celastraceee.

Shrubs or

Stamens alternate with the


Disk large, expanded, flat,

Ovary with 3 or 4

inserted into the

petals,

closely

surrounding the

ovules ascending

fruit cap;
;
ovary.
sular or drupaceous ; seeds often with an aril ; albumen fleshy.
USES.
Sub-acrid, but apparently unimportant plants in a
medicinal point of view.
yellow die is obtained from the
cells

bark of Euonymus tingens in India.


TYPICAL GENERA. Celastrus, Euonymus.
52.

Shrubs.

Staphyleaceae.

with both
imbricated.

common and
Petals

5,

the petals, perigynous.

Leaves opposite, pinnate,

partial stipules.

imbricated.

Disk

Sepals

5,

coloured,

Stamens alternate with

large, urceolate.

Ovary

2- or

DE CANDOLLES SYSTEM.

118
3-celled,

ovules

superior;

erect;

styles

or

3,

cohering.

Seeds with a bony testa and no

Fruit membranous or fleshy.

albumen none.
Staphylea pinnata and trifolia are cultivated as ornamental shrubs under the name of Bladder-nuts, because their
aril

hilum large

USES.

nut-like seeds are enclosed in a bladdery seed-vessel.

TYPICAL GENUS.

Staphylea.

Rhamwacea. Trees or shrubs. Leaves alternate, with


minute stipules. Flowers axillary or terminal, minute. Calyx
53.

Petals distinct, inserted into the orifice of

4-5-cleft, valvate.

the calyx.
Stamens definite, opposite the petals, to which
they are equal in number. Ovary superior, or half-superior, 23- or 4-celled

ovules solitary, erect

frequently a berry

albumen

fruit

fleshy, in

a capsule, or more

very small quantity

embryo with large flat cotyledons, and a short inferior radicle.


USES.
The berries of Rhamnus Frangula, catharticus, and
others, are active purgatives.

When ripe,

those of some species,

especially R. catharticus and infectorius, yield a yellow dye.

The fruit of Zizyphus communis is the Jujube of the shops,


and that of the Z. Lotus gave their name to the Lotophagous
nation of antiquity ; all the fruit of that
genus seems harmless ; Z. Chinensis, indeed, is cultivated in China as the
apple
is with us.
The bark of Ceanothus americanus and some
others is astringent, and has been
employed in diarrhoea.
TYPICAL GENERA. Rhamnus, Paliurus, Ceanothus.

4.

AA

seed

.,
dmded

8een from above '


vertically.

frait -

3.

The same

cut

EXOGEN/E CALYCIFLORJE.

119

Trees or shrubs, with a resinous causbecoming black in drying. Leaves alternate, without pellucid dots.
Flowers small, green, unisexual. Calyx
small.
Petals perigynous, imbricated.
Stamens usually definite.
Disk fleshy, hypogynous. Carpel simple ; styles 1 or
3, occasionally 4 ; ovule solitary, attached by a cord to the
bottom of the cell. Fruit indehiscent. Seed without albumen.
USES. A hard, black, acrid varnish is obtained from Semecarpus Anacardium and Melanorhoea usitatissima. The Cashew
nut, whose eatable kernel is surrounded by a rind full of
54.

Anacardiaceee.

tic juice,

oil, is produced by Anacardium occidentale.


Rhus toxicodendron and radicans are dangerous acrid poisons.
Rhus Coriaria, Cotinus, and some others, are astringents.
Rhus Typhinum, and some others, are cultivated as plants of

resinous acrid

ornament. Pistacia Atlantica and Lentiscus yield mastich


and P. Terebinthus, Scio turpentine.
TYPICAL GENERA. Rhus, Pistacia.

Pistacia Atlantica.

show the

55.

ovule.

4.

1.

Fdbaceee

shrubs, or trees.

Female

ripe fruit

flowers.

opened

to

2.

An

ovary.

show the

3.

The same

cut open to

seed.

Herbaceous plants,
Leguminosa).
alternate
Leaves
;
petiole tumid at the base.
(or

Flowers usually showy.


Calyx inferior, the
Petals
and
segments
variously combined.
unequal,
Stamens deeither papilionaceous or regularly spreading.
Stipules 2.

often

or indefinite, perigynous, or hypogynous.


Ovary simSeeds
destitute
of
albumen.
a
Fruit
ple, superior.
legume.
finite

BE CANDOLLE'S SYSTEM.

120

there are 3 principal


natural order, of which
very large

divisions:

/O<A\

Flowers papilionaceous (340).


Division l.Papilionace*.
and Sissoo, a species
Laburnum,
The Locust-tree,
USES
wood. The roots of Glycyrluza
of Dalbergia, yield valuable
Beans, Kidney Beans, Vetches,
Peas,
glabra are liquorice.
Clover, Suckarticles of food.
are
of
and other sorts
pulse,
and others, are f
Lucerne, Medick, Saintfoin,
ling, Melilot,
der plants.
furnished

Indigo

is

various

by

plants, especially
tinctoria.
gofera

Indi-

kind of manna oozes

from Alhagi
sists

Maurocon-

Cowhage

runi.

of the

on

hairs

stinging
the pods of

Mucuna pruriens.
tain

Astragali

Ceryield

gum Tragacanth. The


seeds of Laburnum and
several others are narcotic

as

also

the

is

root of Piscidia Erythtincture of


rina, the

which is said
more powerful

to

be

than

laudanum.
TYPICAL GENERA.
Cytisus, Lathyrus, Co-

1. The standard,
Adenocarpus frankenioides.
3.
wings, and keel split open. 2. The stamens.
4.
cross section of a seed.
legume, with a portion of one of the valves turned back.

lutea.

Division 2.

Ccesalpiniea.

Petals regularly spreading, im-

Stamens perigynous.
USES.
Senna is the foliage of different species of Cassia.
The Tamarind fruit comes from Tamarindus Indica. The

bricated.

pods of the Carob-tree (Ceratonia Siliqua)


tritious.

Hsematoxylon

Campeachianum

Csesalpinia Brasiliensis, Brazil

TYPICAL GENERA.

wood.

Cassia, Bauhinia.

are highly nuyields

logwood

EXOGEN;E CALYCIFLOR^:.

Cassia acutifolia.

Division 3.

MimoseeE.

1.

flower

121

somewhat magnified.

Sepals and petals valvate.

Sta-

mens hypogynous.

Gum

Arabic oozes from various species of Acacia,


and arabica.
A.
Verek
Catechu is obtained by
especially
Catechu
and
the
bark
of
A.
boiling
many kinds are employed
USES.

Several are fine timber trees.

for

tanning purposes.
Finally,
The
the leaves and branches of some kinds are poisonous.
a
of
coarse
kind
of
stems
Desmanthus
natans
supply
spongy
rice paper.

Most

are objects of great beauty.

TYPICAL GENERA.

J.
Acacia Verek.
same. 4. Half a seed.

Acacia, Mimosa, Inga.

flower magnified.

2.

The

pistil.

3.

section of the

DE CANDOLLE'S SYSTEM.

122
56.

Trees, shrubs, or herbaceous plants.

Rosaceee.

Leaves

alternate, usually with conspicuous stipules, more frequently


Flowers large, showy, arranged
compound than simple.
in
terminal.
but
most
cases
Calyx lined with a
variously,

Stamens usually indefinite.


Carpels
equal.
or
disunited
or consolidated.
several,
Styles distinct,
solitary
and more or less obliquely placed upon the ovary. Fruit
disk.

Petals

Seeds without albumen.

various.

Embryo

Tube of calyx

straight.

and covering
over the achsenia with a false pericarp.
USES.
Fruit astringent.
Petals fragrant and astringent.
Division 1.

Rosea.

fleshy,

Flowers in all cases beautiful.


TYPICAL GENUS. Rosa.
Division 2.

Potentillete.

Carpels numerous, superior, inde-

hiscent.

Usually gay flowers. The fruit of Fragaria is the


The
Strawberry, of Rubus the Bramble and the Raspberry.
roots of Tormentils and some Geums and Potentillas are
USES.

astringent,

and have been used as

TYPICAL GENERA.

febrifuges.

Rubus, Fragaria.

t>

1.

Spiraea Arancus, flower cut open.


2.
section of an ovary. 3. Part of flower
4.
tragamlndica
vertical section of the
half-ripe receptacle, covered with
5.
carpels.
6.
single carpel.
section of a ripe
carpel, with the seed inside

Division 3.

USES.

Spiraea.

Carpels few, 2-valved.

Roots of Gillenia emetic, of


Spirsea ulmaria tonic.
TYPICAL GENUS. Spiraea.

EXOGEN^l CALYCIFLOR.E.
Division 4.

Amygdaleee.

123

Carpel single, a drupe.

The fruit of the Peach, Nectarine, Almond, Plum,


and
Cherry,
Apricot, are produced by various species. Many
are of great beauty on account of their gay flowers.
HydroUSES.

cyanic acid

is

yielded

by the

leaves of

all,

especially of the

Prunus Laurocerasus, or common Laurel. The bark of Prunus Coccomilia and some others is febrifugal.
TYPICAL GENERA.
Division 5.

Prunus, Amygdalus.
Carpels adhering to the calyx.

Pome<z.

Beautiful trees or bushes, bearing a fruit which is,


majority of species, eatable. Apples, Pears, Quinces,

USES.
in the

The
Medlars, Services, are the produce of different species.
The
is
hard.
Hawthorn
is a valuable mausually very

wood

terial for fences.

TYPICAL GENERA.
Division 6.

none.

USES.

Pyrus, Cratsegus.

Sanguisorlea. Flowers often unisexual. Petals

Tube of the calyx hardened.


Astringents of

used for sheep pasture


TYPICAL GENERA.

is

little

importance.

Sanguisorba

Common

Burnet

officinalis.

Alchemilla, Sanguisorba, Poterium.

A
A

2. The same with


1 .
flower with a pair of bracts.
Sanguisorba officinalis.
removed.
3.
half the calyx cut away.
ripe fruit, from which the calyx has been
5. Transverse section of a fruit.
vertical section of fruit and calyx.
4.

DE CANDOLLE'S SYSTEM.

124

Resinous
Amyridacete.
with
pellucid dots.
compound,
57.

minal, panicled.

mens

definite.

Calyx minute.

Ovary

superior,

Leaves

shrubs.

or

trees

Flowers axillary and


1-celled;

ter-

Sta-

Petals 4-5, imbricated.

stigma capitate;
Seed with-

Fruit indehiscent, glandular.


ovules pendulous.
out albumen ; radicle superior.

Us ES Fragrant resinous shrubs.


Myrospermum toluiferum yields Balsam of Tolu and Gum Elemi comes from
some species of Amyris. Balsam of Copaiva is furnished by
.

different species of Copaifera.

TYPICAL GENERA.
58.

Amyris, Copaifera.

Chrysobalanacete.

Leaves simple,

Trees or shrubs.

with veins that run parallel with each


other from the midrib to the margin.
Calyx 5-lobed. Petals
alternate, stipulate,

more or

less irregular, either 5 or

none.

Stamens

definite or

usually irregular.
Ovary superior, solitary, cohering more or less on one side with the calyx ; ovules erect.
Seed solitary, erect. EmFruit, a drupe with 1 or 2 cells.
indefinite,

bryo with no albumen.

The

USES.
of the

West

fruit of

Indies.

Chrysobalanus Icaco

The general

the Cocoa

is

Plum

properties appear to be in-

nocuous.

TYPICAL GENERA.

Hirtella, Chrysobalanus.

Shrubs with square stems. Leaves


without
opposite, simple,
stipules.
Sepals and petals conStamens infounded, indefinite, combined in a fleshy tube.
serted in a fleshy rim.
Anthers adnate, turned outwards.
Ovaries several, simple; ovules solitary.
Nuts enclosed in
59.

Calycanthacea.

the fleshy tube of the calyx, 1-seeded.


ledons convolute.

The

USES.

flowers are fragrant, but of no

TYPICAL GENERA.
60.

Albumen none

Lythraceee.

quently 4-cornered.

known

coty-

use.

Calycanthus, Chimonanthus.

Herbs, rarely shrubs.


Branches freLeaves without stipules. Flowers in

cases showy.
Calyx tubular. Petals inserted between
the lobes of the calyx,
very deciduous. Stamens inserted
into the tube of the
calyx below the petals.
Ovary 2- or

many

4-celled

style filiform

calyx, usually 1-celled.

capsule membranous, covered

by the

Seeds numerous, without albumen.

EXOGENJE CALYCIFLORJE.
USES.

125

The Lngerstromias and Lythrums,

as well as

some

others, are species of great beauty.


Lythrum Salicaria has
been employed in diarrhoea on account of its astringency.
Lawsonia inermis furnishes the Henn6 with which oriental

women

stain their nails.

A few are acrid.

Ly thrum, Ammannia.

TYPICAL GENERA.

Trees or shrubs.
Combretaceee,
Leaves without stiFlowers
pules.
generally showy.
Calyx 4- or 5-lobed, deciduous.
Stamens twice as many as the segments of the
calyx, or three times as many.
Ovary 1 -celled, with from 2
to 4 ovules, hanging from the apex of the cavity. Seed with61.

out albumen

cotyledons usually convolute.


Many species are astringent, and are used by the
tanners in the countries where they occur.
Myrobalan nuts,
;

USES.

also tonics, are

produced by Terminalia

bellerica.

The

kernels

of T. Catappa are eaten like almonds.

TYPICAL GENERA.

Combretum, Conocarpus.

Melastomacea.

62.

Trees, shrubs, or herbaceous plants.

Leaves opposite, with several ribs. Flowers generally purple


and very handsome. Calyx cohering with the angles of the
Petals twisted in aestivation ; filaments curved downovary.

wards

in aestivation

anthers 2-celled, elongated beyond the

insertion of the filament (see fig. 211, p. 46).


Ovary with
several cells, and indefinite ovules.
Pericarp with placentae

attached to a central column.

Of no
fruit, when

USES.
their

importance.
succulent,

TYPICAL GENERA.

is

Seeds innumerable.

The

species appear harmless


eatable.

Ehexia, Melastoma, Lasiandra.

Leaves deciduous, oppoPMladelphacea. Shrubs.


without dots or stipules. Flowers usually white. Calyx

63.
site,

persistent,

imbricate.

having from 4 to 10 divisions.

Stamens

definite.

Petals convolute,
or consoli-

Styles indistinct,

dated; stigmas several. Capsule with 4 to 10 cells, manySeeds scobiform ; aril loose, membranous. Albumen
seeded.
fleshy.

USES.
Merely known as plants of ornament, and someThe rough leaves of Deutzia are said to
times of fragrance.
be used by the Japanese as a polishing material.
TYPICAL GENERA. Philadelphus, Deutzia.

DE CANDOLLE'S SYSTEM.

126

Leaves with transTrees or shrubs.


Myrtaceee.
often
with
a
and
vein
running parallel with their
parent dots,
Petals
like a cap.
4or
sometimes
5-cleft,
margin.
Calyx
64.

Stamens indefinite ; anthers ovate,


quincuncial or wanting.
Fruit either dry or fleshy.
l-2-4-5-6-celled.
small.
Ovary
Seeds definite or indefinite ; embryo without albumen.

The

principal divisions are the following

Fruit 2- or more celled, fleshy.

Division 1.

Myrteee.

Division 2.

Leptospermea.

Fruit 2- or more celled, cap-

sular.

Division 3.

Chamadaucieez.

Fruit 1-celled.

USES.
The spices called Cloves and Pimento are the dried
flowers of Caryophyllus aromaticus and the dried fruit of

The New Holland Eucalypti contain a

Eugenia Pimenta.

great quantity of tannin in their bark.

Cajeputi oil is obtained


from Melaleuca Cajeputi.
The bark of the Pomegranate
root is an anthelmintic.
Almost all the species are beautiful
either in foliage or flower.

Eugenia tuberculata.
men. 4.
ripe fruit.

65.

1.

5.

A flower. 2. The same


A leaf with the dots upon

Onagracea.

alternate or opposite.
lar, 4-lobed, valvate.
tion.

Stamens

2,

4,

divided vertically.

3.

sta-

it.

Herbaceous plants or shrubs.


Leaves
Flowers generally showy.
Calyx tubuPetals regular, with a twisted sestivaor 8, inserted into the
calyx.

Styles

EXOGEN.7E CALYCIFLOR.E.
consolidated.
cells.

Stigma 4-lobed.

127

Fruit many-seeded, with four

Seeds without albumen.

Division 1.

Fruit a capsule.

Petals 4.

CEnotherea.

Seeds

numerous.

These are gay flowered plants, of no known use.


TYPICAL GENERA. (Enothera, Epilobium.

USES.

Jussiaeoides.

Ludwigia
2.

A calyx

and

1.

Division 2.

An

5.

Fuchsiea,

Unknown.

USES.

TYPICAL GENUS.
Division 3.

flower with two sepals and all the petals cut off.
transverse section of the ovary.
4.
seed
3,
embryo extracted.

inferior ovary.

with the distinct raphe.

Fruit a succulent berry.

Petals 4.

Beautiful bushes.

Fuchsia.

Circteea.

Petals

2,

4, or none.

Stamens

Fruit a capsule.
USES. Unknown.

or 2.

TYPICAL GENERA.
66.

Small shrubs, or obscure herbaceous

Cercodiaceae.

Calyx minute.

plants.

Ovary

Circaea, Lopezia.

Petals small or none.

Stamens

1-8.

Ovules pendulous, anaStyles distinct.


Fruit nut-like, usually crowned by the rim

1-4-celled.

tropous (460).
of the calyx.
Seed pendulous, with a small quantity of fleshy

albumen.
Division

Stamen
USES.

1.

Hippuridete.

Calyx obsolete.

1.

Unknown.

'TYPICAL GENUS.

Obscure weeds.

Hippuris.

Petals none.

DE CANDOLLE'S SYSTEM.

128

Hippuris vulgaris.
position of the ovule.

Division 2.

1.

3.

A complete flower. 2. A section of the pistil, showing the


A section of the ripe fruit and seed.

Haloragea.

Stamens more than one.

Calyx toothed.

Petals present.

EXOGEN^l CALYCIFLOR.E.

129

Unknown.

USES.

TYPICAL GENERA.

Myriophylluin, Loudonia.

Loasacea.
Herbaceous plants, hispid, with pungent
Leaves without stipules.
Flowers generally showy,

67.
hairs.

white or yellow. Calyx 5-parted. Petals 5 or 10, hooded, with


an inflexed aestivation the interior often much smaller. Stamens indefinite. Ovary with several parietal placentae, or with
;

a free central lobed one.

numerous, without

aril

Unknown.

USES.

TYPICAL GENERA.

Seeds

Loasa, Bartonia.

Cucurbitacea.

68.

Fruit capsular or succulent.

embryo in axis of fleshy albumen.


Usually handsome plants.

Annual or perennial

herbs.

Stem

climbing by tendrils. Leaves palmated, or with palmate ribs,


covered with asperities.
Flowers white, yellow, or brownish
red, unisexual.

Calyx 5-toothed. Corolla o-parted, scarcely


distinguishable from the calyx, with strongly reticulated veins.
Stamens 5, either distinct, or cohering in three parcels ; anthers sinuous.
Ovary with 3 parietal placentas ; stigmas very
thick,

Seeds

Fruit

velvety or fringed.
flat,

in

an

aril

embryo

flat,

more or

less

succulent.

with no albumen.

USES.
The Gourd, Melon, Cucumber, Pumpkin, Vegetable
Marrow, and Squash, are the fruits of various species, in all
which an acrid purgative principle is diffused ; which, when
concentrated, as in the Bottle Gourd, the Colocynth, and the
Bryony, becomes dangerous, unless administered with skill,
-

when

it

is

a useful medicine.
is

Elaterine, or Elatine, an ex-

found in the Spirting Cucumber,

tremely poisonous principle,


Momordica Elaterium. The seeds are nutty and harmless.
TYPICAL GENERA. Cucumis, Bryonia, Momordica.

Passifloracea.
Usually climbing by means of tenLeaves alternate, with leafy stipules. Flowers often
enclosed in an involucre.
Sepals 5, their tube lined with fila69.

drils.

mentous processes. Petals

5.

Stamens monadelphous. Ovary

Fruit
;
;
stigmas simple, clavate.
with 3 polyspermous placentae.
Seeds with a brittle sculptured testa.
Embryo in fleshy albumen.

stalked,

styles 3

The fruit of Passiflora quadrangularis, the Granaof P. edulis, and several others, contains a pleasant sub-

USES.
dilla,

1-celled

DE CANDOLLE'S SYSTEM.

130

acid pulp, on account of which they are served up at dessert.


The root of the first species is emetic and narcotic ; and similar properties are ascribed to that of P. rubra, which is called

P.
Jamaica Dutchman's laudanum.
as
an
emmenagogue.
reputation
TYPICAL GENERA. Passiflora, Tacsonia.

in

Herbaceous plants.

Turnemcea.

70.

foetida

has some

Leaves alternate,

without stipules, with occasionally two glands at the apex


of the petiole.
Calyx often cploured, with 5 lobes, imbriPetals 5, equal, twisted.

cated.

Stamens

distinct.

Ovary

with 3 placentae; ovules indefinite; styles 3 or 6, cohering


more or less. Capsule 3-valved, the valves bearing the plaSeeds with a thin aril on one side ;
centas in the middle.

embryo
USES.

in the

middle of fleshy albumen.

Unknown.

TYPICAL GENUS.

Turnera.

Portwlacacea.

71.

Succulent

shrubs or herbs.

Leaves

without stipules, or sometimes with membranous ones. FlowPetals generally 5.


ers usually ephemeral.
StaSepals 2.

mens

inserted irregularly into the calyx, or hypogynous,


variable in number.
Ovary 1 -celled ; style single, stigmas
several.

centa.

Capsule 1-celled. Seeds attached to a central plaEmbryo curved round the albumen.

USES.
Insipid plants, occasionally employed as esculents,
as in the case of Portulaca oleracea, the common Purslane.

TYPICAL GENERA.
72.

Calandrinia, Montia.

Herbaceous or half shrubby plants, with


Flowers minute, with scarious bracts. Se-

Illvcebracea.

scarious stipules.
pals 3, 4, or 5.

Petals minute.

Stamens

definite.

Fruit dry, 1-3-celled.


Seeds
superior; styles 2-5.
central placenta ;
embryo on one side of the albumen.
USES.

Unimportant weeds

TYPICAL GENERA.

Ovary
upon a

said to be slightly astringent.

Herniaria, Illecebrum.

73.
Scleranthacea. Small herbs. Leaves
opposite, without stipules.
Flowers axillary, sessile, minute, hermaphrodite.

Calyx 4- or 5-toothed.

Stamens from

1 to 10.

Ovary

simple, superior, 1-seeded. Fruit a utricle enclosed within the

EXOGENJE

131

CALYCIFLOR-ffi.

hardened calyx. Seed pendulous from a funiculus


cylindrical, curved round farinaceous albumen.

Unknown.

USES.

TYPICAL GENUS.
74.

Mere weeds.

Scleranthus.

Succulent herbs or shrubs.

Crassulaceee.

Flowers usually in cymes, showy.

none.

Petals either distinct

20.

embryo

or

Stipules

Sepals from 3 to

Stamens inserted

cohering.

with the petals.


Hypogynous scales usually several, 1 at the
base of each carpel.
Ovaries of the same number as the

which they are placed. Fruit of several


opening on their face. Seeds variable in number.
USES.
Sempervivum tectorum, and many others, are refri-

petals, opposite to
follicles,

gerants and somewhat acrid. Some are plants of considerable


beauty, and capable of growing in the most exposed and sunburnt places.
Sempervivum glutinosum is used to impregnate the water in which the fishermen of Madeira steep their
nets, in order to render

TYPICAL GENERA.

them

Succulent shrubs or
Mesembryacea or Ficoidea.
Flowers showy, opening only under bright sunshine.

75.
herbs.

Sepals definite, succulent.

mens

durable.

Sempervivum, Sedum.

indefinite.

Petals indefinite, linear.

Ovary many-celled.

Capsule many-celled, with a starry dehiscence.


curved or spiral, on the outside of mealy albumen.
USES.

Sta-

Stigmas numerous.

Mesembryanthemum emarcidum,

Embryo

the Hottentot's

when

bruised and fermented, becomes narcotic, and is


fig,
used like tobacco.
M. crystallinum and nodiflorum are collected in the countries

where they grow wild,

for the sake of

the alkali they contain.

TYPICAL GENUS.
76.

Mesembryanthemum.

Cactacea. Succulent shrubs, usually destitute of leaves,


Flowers usually very handsome. Se-

and with spinous buds.

Stamens indefipals indefinite, confounded with the petals.


nite ; filaments long, filiform.
inferior,
1-celled, with
Ovary

numerous

Fruit succuparietal placentae ; stigmas numerous.


Seeds without albumen.
USES.
The fruit is eaten under the name of Indian figs.

lent.

TYPICAL GENERA.

Cereus, Mammillaria.

K 2

DE CANDOLLE'S SYSTEM.

132

Bushes with alternate leaves, mem


Flowand
a
plaited vernation, often spiny.
stipules,
4or
ers in axillary racemes.
5-parted, reguCalyx superior,
1 -celled,
Petals 5, minute. Stamens 5.
lar.
inferior,
Ovary
with 2 parietal placentae. Berry 1 -celled, many-seeded; embryo minute, in horny albumen.
USES.
Bibes rubrum is the common garden Currant, B.
Black Currant, and B. Grossularia the Goosethe
nigrum
all
well-known
fruits. Many have beautiful flowers.
berry,
77.

Grossulacea.

branous

TYPICAL GENUS.

Bibes.

78.
Saxifmgacea. Herbaceous plants.
with or without stipules.
Calyx superior or

Leaves simple,
inferior.

Petals

Stamens 5-10, perigynous or hypogynous ; anthers bursting longitudinally.


Disk hypogynous or perigynous, rarely consisting of 5 scales.
Ovary 1-celled, with two
formed
from extended points of
parietal placentae.
Styles 2,
the ovary.
Fruit membranous, with two divaricating lobes.
5,

or none.

Seeds numerous, very minute.


fleshy albumen.
USES.

gent roots.

Embryo

taper, in the axis of

Heuchera Americana, and some

Many

TYPICAL GENERA.

are pretty flowers.

Saxifraga, Heuchera.

others,

have

astrin-

EXOGEN^E CALYCIFLOR.3E.
Shrubs

Escattoniacea.

79.

leaves.

with

133
toothed,

alternate,

Flowers showy.

glandular, exstipulate
Calyx 5toothed. Petals forming a tube, but finally separating ; aestivation imbricated.
Disk conical, epiStamens definite.

Ovary

gynous.
centse

in

with two large polyspermous plaFruit


style simple ; stigma 2-lobed.

2-celled,

the axis

capsular, splitting by the separation of the cells


base.
Seeds minute ; embryo in oily albumen.

Escallonia.

Hamamelacea. Shrubs. Leaves alternate, with deFlowers sometimes unisexual. Calyx in


Stamens 8
4 being sterile.
4, linear.

80.

ciduous stipules.
4 pieces.
Petals

styles 2 ; ovules solitary, pendulous.


Embryo in the middle of horny albumen.

2-celled

capsular.

Araliacea.

81.

habit

5-10.

Fruit

Unknown.

USES.

TYPICAL GENERA.

the

their

Unknown.

USES.

TYPICAL GENUS.

Ovary

at

Hamamelis, Fothergilla.
Trees, shrubs, or herbaceous plants, with
Petals
Calyx entire or toothed.

of Apiacese.

Stamens equal

to the petals or twice as

from without an epigynous disk.

many, arising
Ovary with more cells than

two.

Fruit succulent or dry, consisting of several 1-seeded


Seeds pendulous. Embryo minute, in copious albumen.
USES.
Panax quinquifolium forms the root Ginseng, re-

cells.

sort of
garded by the Chinese as a powerful stimulant.
in
North
America
from
Aralia
nuis
Sarsaparilla
prepared

Common

dicaulis.

TYPICAL GENERA.
82.

Cornaceee.

Ivy,

Hedera Helix, has

irritating leaves.

Hedera, Aralia.
Trees or shrubs, seldom herbs.

Leaves

Flowers
(except in one species) opposite, entire or toothed.
dioecious.
4.
Petals
4, oblong, broad,
Sepals
occasionally

Stamens 4, alternate with the petals.


crowned by the calyx, 2-celled. Seeds pendulous,
valvate.

Albumen

Drupe
solitary.

fleshy.

Cornus mascula, the Cornelian Cherry, and some


C.
others, produce a succulent eatable fruit of bad quality.
florida and sericea have a powerfully tonic bark.
TYPICAL GENERA. Cornus, Aucuba.
USES.

DE CANDOLLE'S SYSTEM.

134

Herbaceous plants with


Apiacea or Umbellifera.
Flowers in umbels.
Calyx entire or 5Stamens
Petals 5, usually inflexed at the point.
toothed.
83.

stems.

fistular

Ovary 2-celled. Styles 2, di5, alternate with the petals.


Fruit consisting of 2 carverging ; disk double, epigynous.
from a common axis. Seed
pels, or mericarps, separable
Embryo minute, at the base of horny
solitary, pendulous.
albumen.
USES.

The

Carrot, Parsnip, Parsley, Fennel, Skirret,

others, are eatable.

Celery

is

poisonous when

and

wild, bland

if

Many species are dangerous poisons, as (Enanthe


Cicuta
crocata,
virosa, Conium maculatum, .ZEthusa Cynacultivated.

pium

Coriander, Anise.

Dill,

nax,

others have aromatic carminative fruits, as Caraway,

foetid

gum

resins,

Assaftetida,

GENERA.

TYPICAL

Ammoniacum, Opopa-

exude from certain Oriental


Pastinaca,

Carum,

species.

Petroselinum,

Daucus.

Athamanta

cervarisefolia.

2.
separate flower, with hairy petals.
petal
itself.
3.
ripe fruit with the two carpels or mericarps separating from the
double carpopod or axis.
4.
seed deprived of its
integuments, and divided vertically, so as to show the position of the embryo.
1.

by

The genera of
terized

cut

is

this large

very much by

and

difficult

order being charac-

peculiarities in their fruit, the following


intended to explain the
principal terms employed in

speaking of them.

EXOGEN.E CALYCIFLORJE.

135

1. Is an ideal plan of a fruit divided transversely ; a a is the commissure, or


2. Is a
plane of contact of the mericarps ; b b primary ridges ; c c secondary ridges.
view of the back and section of the fruit of Laserpitium Siler ; each mericarp has the
secondary ridges winged, the primary obsolete ; there are two vittae on the commissure, and one under each secondary ridge ; these vittae, which are cavities containing
3. Sclerosciadium humile ; the
oil, are represented by dots ; the albumen is solid.
primary ridges are corky ; there are no secondary ridges ; the vittae alternate with
the primary ridges, and there is one at each edge of the commissure ; the albumen is
solid.
4. Discopleura capillacea ; there are 5 very small primary juga, the two
lateral of which are in contact with a thickened accessory margin ; there are 2 vittae
on each face of the commissure, and one between each primary ridge ; the albumen is
solid.
5. Echinophora spinosa ; albumen involute ; vittae alternate with the primary
6. Compressed fruit of Diposis saniculaefolia ; the commissure is very narridges.
row ; there are 5 minute primary ridges ; one along the back, one along each edge,

and two on the indexed

side

the albumen

SUBCLASS
LorantJiacea.

84>.

Leaves opposite, without

and

COROLLIFLOR^E.

Parasitical
stipules.

the

base,

1-celled

half-shrubby

valvate

ovule erect.

more or

less

united

stamens opposite to them. Ovary


Seed solitary ; emFruit succulent.

bryo cylindrical, longer than the fleshy albumen.


USES. Bark astringent ; that of Loranthus tetrandrus
in Chili for a black dye.

ployed
TYPICAL GENERA.

plants.

Flowers either very long


Calyx with 2 bracts at

and green.
Corolla with 3, 4 or 8 petals,

tubular, or small

the base.
at

III.

is solid.

Miseltoe

Viscum, Loranthus.

is

is emViscum album.

DE CANDOLLE'S SYSTEM.

136
85.

Caprifoliacea.

Shrubs or herbaceous plants, with


Flowers usually showy

opposite leaves, destitute of stipules.

Calyx 4-5-cleft, with bracts at its base. Comonopetalous or polypetalous, rotate or tubular, regular
Stamens epipetalous. Ovary with from 1 to 5
or irregular.
cells.
Fruit indehiscent, 1 or more celled.
Embryo straight

and

fragrant.

rolla

in fleshy

albumen.

Honeysuckles, species of Caprifolium, are beautiful,


The Elder has sudorific flowers,
fragrant, twining shrubs.
and drastic foetid leaves. The roots of Triosteum perfoliatum
USES.

are emetic

and

cathartic.

The

fruit

of Symphoria racemosa,

the Snowberry, is a favourite food of pheasants ; that of different species of Viburnum is eatable, but unpleasant.

TYPICAL GENERA.
86.

Sambucus, Caprifolium, Viburnum.

Leaves
Trees, shrubs, or herbs.
or
with
verticillate,
simple, opposite
interpetiolary stipules,
Avhich are simple, bifid, or multifid, and form one of the
prinInflorescence extremely
cipal characteristics of the order.
Cinchonaceee.

varied.

Corolla tubular, regular, valvate, or


Calyx simple.
Stamens all on the same line, alternate with the
lobes of the corolla.
Ovary surmounted by a disk ovules
numerous or few. Fruit either splitting, or indehiscent and

imbricated.

Seeds definite or indefinite


dry, or succulent.
surrounded by horny albumen.
USES.
Foremost among the useful

embryo

small,

species of this large


order stand the species of Cinchona, whose bark is so valuable
on account of its tonic
febrifugal qualities ; in this respect a
large number of other genera correspond, among which are

Buena, Remija, Portlandia, and Exostema. Others are poweremetics ; as Cephaelis


Ipecacuanha, whose roots form the
best Ipecacuanha of the
shops ; Richardsonia scabra, and
ful

species
Manettia, Chiococca, and Spermacoce. A
few have the emetic
principle so concentrated as to be dangerous poisons, as Randia dumetorum.
Coffee is the horny
albumen of Coffea Arabica.
several

of

EXOGEN^E COROLLIFLOR;E.

137

1.
flower magnified.
section across a ripe fruit.
Coffea Arabica.
2.
3.
portion of a seed, stowing the small embryo laid bare in the end of convolute albumen.

Richardsonia scabra.
section of a seed, with

1.

an

An

erect

ovary with

its calyx.
2.
in copious albumen.

embryo

corolla.

3.

.A

vertical

Herbaceous plants or under-shrubs.


87.
Dipsacete.
Leaves opposite or whorled.
Flowers capitate, surrounded

by a many-leaved
surrounded by an
Stamens 4
lous ovule
like calyx,

involucre.
involucel.

anthers distinct.

stigma simple.

embryo

in fleshy

Calyx superior, membranous


Corolla

Ovary

oblique,

1-celled,

imbricated.

with a pendu-

Fruit crowned by the pappus-

albumen.

DE CANDOLLE'S SYSTEM.

138

The ripe heads of Dipsacus fullonum, dried, are


USES.
formed of hard stiff spines, and are employed by fullers, in
dressing cloth, under the name of teasels.
TYPICAL GENERA. Scabiosa, Dipsacus, Knautia.
88.

Valerianacete.

Herbs.

Leaves opposite, without

Flowers corymbose, panicled, or in heads. Calyx


Corolla tubular,
superior, membranous, or resembling pappus.
1 to 5.
calcarate.
or
sometimes
Stamens
regular
irregular,
stipules.

Ovary with

perfect cell, and 2 other abortive ones ; ovule


Fruit dry.
stigmas 1 to 3.
Embryo destitute
1

pendulous ;
of albumen.

Common

USES.

Valerian, and several others, have power-

fully aromatic, antispasmodic, febrifugal roots. The genus Valerianella consists of annual herbs, whose leaves are used as

under the name of Lamb's lettuce.

salad,

The Spikenard of

was Nardostachys Jatamansi.


TYPICAL GENERA. Valerianella, Centranthus, Valeriana.

the ancients

Valeriana Celtica.
calyx.

npe

3.

fruit

The

and

fruit,

An

entire

flower magnified.

with the pappose


full-grown calyx.

2.
4.

The ovary and young

A vertical section

of a

seed.

89.

Asteracea or Composite.
Shrubs, or herbs, extremely variable in appearance.
Flowers in heads, surrounded by an involucrum, and seated on a

from which
a pappus
Co(328).

receptacle,

palese often spring.

Calyx obsolete

EXOGEN/E COROLLIFLOR2E.
regular or irregular.

rolla

Ovary

inferior, one-celled,

Anthers

139
a tube.

united into

with an erect ovule.

Embryo

with-

out albumen.
Division 1.

Cichoracea.

TYPICAL GENERA.
Division 2.
lar.

Florets

all ligulate.

Milky.

Hieracium, Taraxacum.

Corymbifera.

Corolla funnel-shaped.

Florets in part or wholly tubuInvolucrum hemispherical, leafy

or scaly, soft, seldom spiny.

TYPICAL GENERA.

Chrysanthemum, Tussilago.

Division 3.

Cynaracea. Florets wholly tubular. Corolla


with a ventricose throat. Involucrum hard, conical, and generally spiny.

TYPICAL GENERA.
Division 4.

Carduus, Cynara, Onopordum.

Labiatiflvra.

Florets bilabiate.

TYPICAL GENERA.
USES.
ciple is

Among
commonly

Mutisia, Triptilion.
the Cichoraceous division a narcotic prinfound, which in the garden Lettuce is so

diffused as to be bland,

and

in

Lactuca virosa

trated as to render the extract similar to

is

so concen-

opium

in effect.

Succory, Endive, Salsafy, Scorzonera, well-known esculents,


Of the Corymbiferous division, Chamomile is
belong here.
characteristic,

with

its bitter

tonic qualities.

Many

others,

such as Coltsfoot, Elecampane, Feverfew, correspond in proWormwood, Southernwood, species


perties with Chamomile.
of Artemisia, are aromatic and extremely bitter

pungent herb, used for pickling,


Pellitory of Spain,

organs

and
and

is

acrid,

is

;
Tarragon, a
Artemisia Dracunculus.

and

excites the salivary


Pyrethrum ; and similar

Anacyclus
produced by Spilanthus oleracea, Bidens

powerfully,

effects are

which

is

others.
others,

tripartita,

The Sunflower, Guizotia oleifera, Madia sativa,


yield a bland oil when their seeds are pressed.

Jerusalem artichokes, a well-known article of food, are the


tubers of Helianthus tuberosus. The Cynaraceous division consists principally

of bitter plants.

Centaurea calcitrapa, Sily-

bum

(or Carduus) Marianum, Cnicus Benedictus, and the


common Burdock, are all stomachics of some importance.
The flowers of Carthamus tinctorius are dried for the use of

The fleshy receptacles of


the dyers, and resemble Saffron.
the
bottoms
of our kitchens.
are
artichoke
Cynara Scolymus

DE CANDOLLE'S SYSTEM.

2.
ligulate
tubular floret of the disk.
1.
involucrum and
5.
4. An anther.
stigmas.
ach^mum cut
6.
lp e
the lorets have fallen.

Argyranthemum Jacoteifolium.
3. Style and

florel^Tthe ray.

An

which
receptade, from
with toothed coronetted pappus.
through vertically,

Intl

The

old divisions of this large order are adhered to be-

cause they appear more likely to be permanent than the more


recent suborders, &c. proposed by De Candolle and others,
in

which

peculiarities in the stigma are chiefly employed.


who desires to become acquainted with the de-

The

student

tails

of this

enormous order, numbering more than 7000


De Candclle' s Prodromus, Vols. V. VI.
(

species, will consult

EXOGEN^

COROLLIFLORJE.

141

The preceding wood -cut will assist him in understanding the distinctions of that author.

and VII.

1. Tubular floret of Webbia aristata, with double pappus (Vernoniacea,


2.
D.C.)
Tubular floret and stigma of Anisochaeta mikanioides, with pappus of 4 setae (Eupaof
Tubular
floret
Berthelotia
3.
with
toriacetB, D.C.)
lanceolata,
silky pappus (Aste-

4. Stigma of Blumea senecioides (Asteroidete, D.C.)


5. Ligulate floret
roidea, D.C.)
and stigma of Lipochaeta umbellata ; pappus of two unequal winged paleae (Senecio6. Stigma of Dunantia achyranthes (Senecionidcce, D.C.)
nulece, D.C.)
7. Tubular
floret with ventricose throat and the
stigma of Aplotaxis Nepalensis (Ci/narete,

D.C.)

8.

Ligulate bilabiate floret of Oreoseris lanuginosa (Mutisiacetz, D.C.)


Brachyramphus obtusus (Cictioracete, D.C.)

9.

Ligulate

floret of

90.

Galiac&e,

or

Herbaceous plants, with


Stems usually angular.
stipules.

Stellate.

whorled leaves, destitute of

Corolla valvate, rotate or tubular,


Calyx 4-5- or 6-lobed.
Stamens
in
number to the lobes of the corolla,
regular.
equal
and alternate with them.
Ovary 2-celled ovules solitary,
;

erect.

Fruit a didymous, indehiscent pericarp.

Embryo

minute, straight, in horny albumen.


USES.
The roots of Bubia tinctorum yield madder, a
in
which others participate, though in a less degree.
quality

The yellow flowers of Galium verum are used to curdle milk.


The fragrant Woodruff has the reputation of being diuretic
;

Asperula cynanchica is said to be astringent. Except the


species used for dyeing, none are of any real importance.
TYPICAL GENERA. Asperula, Galium, Rubia.
91.

out

Goodeniacea. Herbaceous plants, rarely shrubs, withLeaves scattered, without stipules.


Flowers

milk.

Calyx superior, equal or unequal. Corolla more or


folded inwards
split at the back ; the segments
in aestivation.
Stamens 5, distinct. Ovary with indefinite
Fruit
ovules; stigma surrounded by a membranous cup.
a capsule ; albumen fleshy.
Unknown. TYPICAL GENERA. Goodenia, Euthales.
USES.
showy.

less irregular,

92.

Scawlacea.

axillary or

terminal,

Herbaceous plants with the flowers


and never in heads. Calyx superior,

sometimes obsolete. Corolla irregular, split at the back,


Stathe edges of the divisions folded inwards in aestivation.
mens 5, distinct ; anthers distinct or united. Ovary fewwith solitary erect ovules ; stigma surrounded by
Fruit drupaceous or nucamentaceous.
USES.
Unknown.

celled,

cup.

TYPICAL GENERA.

Sceevola,

Dampiera.

DE CANDOLLE'S SYSTEM.

142
93.

Glandular herbs.

Stylidiacea.

Calyx superior,

StaCorolla irregular, imbricated.


2-6-parted, permanent.
which
with
mens 2, connate into an elastic slender column,

the style

is

consolidated.

many-seeded. Seeds albuminous.


Unknown. Eemarkable
USES.

column of stamens.
TYPICAL GENERA.

for

the

irritable

elastic

Stylidium, Leuwenhoekia.

Herbaceous milky plants or shrubs.


Flowers usually showy.

Lobeliacea.

94.

Capsule 2-valved,

2-celled.

Ovary

Leaves alternate, without stipules.

Calyx

or entire.
superior, 5-lobed,

Corolla irregular, 5-lobed,

Stamens 5; anthers cohering. Stigma fringed.


Fruit capsular, 1 or more celled, many-seeded ; embryo in

or 5-cleft.

the axis of albumen.

USES.

The

abound in a milky juice of extreme


is an emetic, but dangerous from

species

Lobelia inflata

acridity.
its violence.

hypercathartic.

Many

TYPICAL GENERA.
95.

(or Isotoma) longiflora


are plants of great beauty.

Hippobroma

is

fatally

Lobelia, Clintonia, Siphocampylus.

Herbaceous plants or under-shrubs.


Flowers
stipules.

Gesneraceee.

Leaves opposite, rugose, fleshy, without

Calyx half superior, valvate. Corolla tugenerally showy.


Anthers cohering, with
bular, with an imbricate sestivation.
a thick connective. Ovary 1 -celled, surrounded by glands,
with 2-lobed polyspermous placentae ; stigma capitate. Embryo in the axis of albumen.
USES.

The

succulent fruit

is

eatable.

Some

species yield

a dyeing substance. The species are, however, of no real


importance but they are generally gay flowers.
TYPICAL GENERA. Gloxinia, Gesnera, Columnea.
;

96.

Campanulacea.
yielding a white milk.

Herbaceous plants or under-shrubs,


Leaves alternate, without stipules.

Flowers usually showy.

Calyx superior, permanent. CoStamens


regular, valvate.
alternate with the lobes of the corolla.
Anthers distinct.
Style covered by collecting retractile hairs ; stigma naked.
Fruit dehiscing by apertures, or valves.
Seeds numerous ;

rolla usually 5-lobed,


withering,

embryo

in the axis of

albumen.

EXOGENJE COROLLIFLORJE.
USES.

Slightly acrid, but not dangerous.

used like Radishes,


TYPICAL GENERA.

is

Rampion, a root
Campanula Rapunculus.
Campanula, Phyteuma, Roella.

Wahlenbergia procumbens.
4.

143

1.

An

Transverse section of the ovary.

entire
5.

flower.

A vertical

2.

Stamens.

3.

section of a seed,

stigma.

showing the

embryo.

97.

ovary is
USES.

Are the same

Vaccinacea.

as Ericacese, only the

inferior.

The bark

is

slightly astringent,

and

fruit succulent.

Cranberries are the fruit of species of Oxycoccus, Bilberries


and Whortleberries of species of Vaccinium.

TYPICAL GENERA.
98.

Ericaceae.

Vaccinium, Thibaudia.

Shrubs or under-shrubs.

without stipules.

Leaves ever-

4- or 5-cleft,

Calyx
green, rigid,
Corolla hypogynous, 4- or 5-cleft, imbricated.

inferior.

Stamens de-

hypogynous; anthers 2-celled, dehiscing by a pore.


Ovary many-celled, many-seeded style 1. Fruit capsular.
Seeds indefinite, minute ; embryo in the axis of albumen.
USES.
Loiseleuria procumbens, Rhododendron ferrugineum, and others, are astringents. Arctostaphylus Uva Ursi
Rhododendron Chrysanthum is a powerful naris diuretic.
cotic, and this seems to be a general character of the order,
some of which, as Rhododendron maximum, Kalmia latifolia,
and Azalea Pontica, are dangerous poisons. Most of the
finite,

species are plants of great beauty.

TYPICAL GENERA.

Rhododendron, Kalmia, Erica.

DE CANDOLLE'S SYSTEM.

144

Rhododendron
save

one.

2.

albiflorum.

An

1.

anther.

3.

A corolla and pistil, with all the stamens removed


A ripe capsule burst. 4. A vertical section of a

seed.

99

Leaves
Trees or shrubs without milk.
or
divisions.
in
6
3
Calyx inferior,

Ebenacea.

alternate,

coriaceous.

Stamens
Corolla hypogynous, usually pubescent, imbricated.
definite ; twice as many as the segments of the corolla, four
times as many, or the same number.
Ovary several-celled,
the cells having 1 or 2 pendulous ovules;
style divided.
Albumen cartilaginous ; embryo in
Fruit fleshy, few-seeded.
the axis

radicle turned

Subdivision.

nous.

towards the hilum.

Styraceae.

Ovary

inferior.

Stamens perigy-

Style simple.

The fruit of Diospyros Lotus, Kaki, and others, is


USES.
extremely astringent when* green, but becomes bletted and
sweet after a time, when it is eaten.
Diospyros Virginiana and
others have a febrifugal bark.
Ebony is the wood of Dios-

The
pyros Ebenus and several other species of that genus.
fragrant gum resins, Storax and Benzoin, are produced by
species of Styrax.

TYPICAL GENERA.
100.

Aquifoliacea.

Flowers small.

Trees or shrubs.

Leaves coriaceous.

Sepals inferior, 4 to 6, imbricated.

Corolla

Stamens alternate with its segments. Disk


Ovary with from 2 to 6 cells; ovules solitary, pen-

hypogynous.
none.

Diospyros, Maba, Styrax.

EXOGEN^E COROLLIFLORJE.

145

Fruit indehiscent, with from 2 to 6 stones.

dulous.

Seed

suspended ; albumen large embryo small, 2-lobed.


USES.
Ilex Aquifolium, the common Holly, has leaves
;

emetic,

and

fugal.

The

berries purgative
fruit

have similar

Some

its

leaves are powerfully febriverticillatus and others

and bark of Prinos

qualities.

Paraguay tea

is

Ilex

Paraguensis.

are diuretic.

TYPICAL GENERA.

Ilex, Prinos, Cassine.

101.

Trees or shrubs with milky juice.


Sapotaceee.
Leaves alternate, without stipules, coriaceous.
Calyx inferior, regular,

permanent.

Corolla hypogynous;

its

segments

usually equal in number to those of the calyx, seldom twice


or thrice as many.
Stamens arising from the corolla, definite.

Anthers
present.

each

turned outward; sterile stamens usually


with
several cells, and one erect ovule in
Ovary
1
Fruit
baccate.
Seeds nut-like. Testa
Style

usually

cell.

bony, shining. Embryo large, usually in fleshy albumen.


USES.
The species are generally astringent and febrifugal.
Achras Sapota and others are the Sapodilla plums, whose
fruit is

much esteemed
West Indian

in the

West

Indies.

The

Star-apple

Chrysophyllum Cainito.
vegetable butter is yielded by some species of Bassia.
TYPICAL GENERA.
Achras, Chrysophyllum, Mimusops.
(another

102.

fruit)

is

Trees or shrubs.

Myrsinacea.

Leaves alternate,

Calyx 4- or 5-cleft.
Stamens opposite the segments of the

serrated, coriaceous; stipules wanting.

Corolla hypogynous.
corolla; sometimes 5 sterile, petaloid, additional filaments.
Ovary 1, with a free central placenta ; style 1. Fruit fleshy,

mostly 1-seeded.

Seeds peltate, albumen horny

lying across the hilum.


USES. Embelia robusta

TYPICAL GENERA.

is

embryo

said to have purgative berries.

Ardisia, Myrsine.

Branches usually dichoCalyx monophyllous, permanent.


Stamens 2. Ovary
Corolla hypogynous, 4-cleft, valvate.
without any disk, 2-celled
ovules pendulous ; stigma bifid
or undivided.
Seeds with dense
Fruit often 1-seeded.
103.

tomous.

OleaceaE.

Trees or shrubs.

Leaves opposite.

albumen.
USES.

Olive

oil

is

obtained from the fruit of Olea eu-

DE CANDOLLE'S SYSTEM.

146

exudes from the trunk of Ornus europsea


bark of the Olive is a powerful febrifuge.

Manna
ropsea.
and others. The
Phyllireas are

handsome evergreen shrubs.

TYPICAL GENERA.

Olea, Phyllirea, Syringa.

Leaves opposite or alternate,


mostly compound. Calyx divided or toothed, permanent.
Corolla regular, with from 5 to 8 divisions, imbricated and
Shrubs.

Jasminacea.

104.

Stamens

twisted.

2.

Ovary

destitute of a disk, 2-celled

Seeds with no albumen.


ovules erect ; stigma 2-lobed.
USES.
The flowers of most species are fragrant.
leaves and bark are bitter, but of little moment.
TYPICAL GENERA.

The

Jasminum, Nyctanthes.

Apocynacea. Trees or shrubs, usually milky. Leaves


opposite, quite entire, often having glands upon the petioles,
with no stipules. Calyx inferior, permanent. Corolla reguStamens 5.
Filaments distinct.
lar, 5-lobed, contorted.
105.

Ovaries 2, or 1 which is 2-celled, polyPollen granular.


spermous.
Stigma 1. Seeds with fleshy albumen.
USES.
Often dangerous poisons, but in some cases simply
The root of Nerium, the kernel of Tanghinia
purgatives.
the seeds of various kinds of Strychnos, called
first class ; the leaves of Cerbera

venenata,

Nux

vomica, belong to the

Manghas, Allamanda cathartica, to the second.


elastica, and others, abound in Caoutchouc.
TYPICAL GENERA. Vinca, Echites, Nerium.

Vahea,

Urceola

106.
Shrubs or herbaceous plants, milky,
Asclepiadacea.
and often twining. Leaves entire, opposite, having cilise between their petioles. Calyx inferior, permanent. Corolla 5lobed, regular, imbricated, very seldom valvular. Stamens 5 ;
filaments connate

anthers 2-celled

and sticking by 5 glands


Ovaries

to as

pollen cohering in masses,

many

processes of the stigma.

Stigma common to both styles, 5cornered.


Follicles 2.
Seeds comose ; albumen thin.
USES.
The roots of many are emetic, sudorific, acrid, and
2.

purgative.
indicus.

Styles 2.

Indian Sarsaparilla is the root of Hemidesmus


Asclepias tuberosa and Curassavica are employed

as cathartics in the United States


leaves of

Senna ;

and West

Cynanchum Argel are used


The extract
they are acrid.

in

Egypt

Indies.

The

to adulterate

of Calotropis gigantea,

EXOGEN^l COROLLIFLORJE.
the

Mudar

plant,

147

powerfully alterative and purgative.

is

species have a tough fibre, which renders


cordage ; others yield abundance of Caoutchouc.

Many

TYPICAL GENERA.

them

fit

for

Periploca, Stapelia, Physianthus.

Schubertia multiflora.
1. The anthers united to the
2. The ovary and
stigma.
3.
stigma, from the latter of which the pollen masses have been removed.
pair of
pendulous pollen masses, with their gland. 4. The ripe follicles.

Trees or shrubs, often twining or


Leaves
climbing.
opposite, usually compound, without stiand showy. Calyx inferior, sometimes
Flowers
large
pules.
107.

Bignoniacea.

Corolla irregular.
Stamens 5, of which 1 alare sterile.
Ovary in a disk, 2-celled,

spathaceous.

ways and sometimes 3


polyspermous
capsular
pressed.

style 1

stigma of 2 plates.

Fruit berried or

the latter, 2-valved, 2-celled, long and


Seeds often winged ; albumen 0.
if

com-

Some have hard timber,


USES. Usually beautiful plants.
and a red fecula is obtained from the leaves of Bignonia CheThe genera with berried fruit form a pecurere and others.
liar division, and include Crescentia Cujete, the Calabash-tree,
and Parmentiera edulis, both of which have eatable fruit.
TYPICAL GENERA. Bignonia, Tecoma.
108.
cal.

Cyrtandraceee.

Flowers showy.

Leaves opposite, often radi-

Herbs.

Calyx

Corolla irregular, imbricated.

inferior,

campanulate, equal.

Stamens didynamous.
i 2

Disk

DE CANDOLLE'S SYSTEM.

148
annular.

Ovary

1 -celled,

with 2 double placentae

stigma

Fruit capsular and siliquose, or succulent, manySeeds minute, often with tails ; albumen absent.

2-lobed.

seeded.

Unknown.

USES.

TYPICAL GENERA.

^Eschynanthus, Streptocarpus.

Leaves opposite,
Flowers showy.
Calyx inferior, permanent. Corolla regular, with an imbriStamens inserted upon
cated, twisted, or plaited aestivation.
the corolla, some of them occasionally abortive.
Ovary 1-

Herbaceous

Gentianaceee.

109.

plants.

without stipules, usually 3-5-ribbed.

entire,

Capsule or berry many-seeded ; the


stigmas 1 or 2.
of
the
turned
inwards.
valves
margins
Embryo in the axis of
soft albumen.
celled

USES.
tensely

so.

All the species are more or less bitter ; many inThe Gentian root of the shops is obtained from

Gentiana lutea chiefly the leaves and stems of Agathotes


Chirayta furnish the Gentian of India.
Menyanthes trifoliata
;

is

the Buck-bean, employed advantageously as a tonic.


TYPICAL GENERA. Erythrsea, Gentiana, Chironia.

Gentiana amarella.
the ripe

fruit.

site.

3.

1.

seed.

Section of the ovary of Chironia baccifera.


4.
vertical section of it.

Polemoniacea.

Calyx

inferior,

Herbaceous
5-parted.

plants.

Corolla

2. Section of

Leaves oppo-

regular,

5-lobed.

EXOGEN.E COROLLIFLOR^!.
Stamens
celled

5,

unequal, on the tube of the corolla.


Ovary 33-lobed.
Capsule 3-celled ; 3-valved, the

stigma

valves separating from the axis.

Unknown

USES.

Embryo

in

horny albumen.

TYPICAL GENERA.

Polemonium, Phlox,

Ill

Herbaceous plants, or shrubs, usually


Leaves alternate. Calyx permanent, in-

Gilia.

GonvolvulacetK,

twining and milky.

in 5 divisions, remarkably imbricated, often unequal.


Corolla hypogynous, plaited.
Stamens 5, inserted towards
the base of the corolla.
Ovary with 2 to 4 cells, few seeded ;
ferior,

ovules erect

1.

style

Disk annular.

Capsule with the

valves fitting at their edges to the angles of a loose dissepi-

Seeds with mucilaginous albumen

ment.

embryo curved

cotyledons shrivelled.

The

Scammonia yield Scamtrue


mony
Exogonium Purga,
Jalap ; of Ipomoea Batatoides, a kind of false Jalap, called Purga Macho ; and a great
many more possess similar properties. The Batatas, or Sweet
USES.
;

roots of Convolvulus

of

Potatoe, has the purgative quality so much diffused as to be a


valuable article of food ; the great roots of others have also

been found eatable.

TYPICAL GENERA.

Ipomoea Batatoides.

1.

Ipomoea, Convolvulus, Calystegia.

The

pistil

and annular disk.

3.
the ovary.
capsule of Convolvulus tricolor.
seed of that species.

4.

2.

transverse section of
section of the

vertical

DE CANDOLLE'S SYSTEM.

150
112.

Leafless parasites.

Cmcutacea.

Calyx permanent,

Corolla permanent, imbriinferior, 4-5-parted, imbricated.


Stamens
cated.
Scales alternating with segments of corolla.
opposite the last.
styles 2.

USES.

Capsule.

Embryo

ovules in pairs,

spiral, in fleshy

erect

albumen.

Unknown.

TYPICAL GENUS.
113.

2-celled

Ovary

Cuscuta.

Cordiacete.

Trees.

Leaves scabrous, without

sti-

pules.
Calyx inferior, 5-toothed. Corolla regular. Stamens
alternate with the segments of the corolla.
Ovary 4-celled,

Fruit
with 1 pendulous ovule in each cell ; stigma 4-cleft.
Seed pendulous by a funiculus cotyledrupaceous, 4-celled.
dons plaited ; albumen 0.
;

USES.

Unimportant.

Sebesten plums, an emollient muciby Cordia Myxa, and Sebestena.

laginous fruit, are produced


TYPICAL GENUS. Cordia.

114.

Herbaceous plants, or shrubs. Stems


Boraginaceee.
Leaves alternate, covered with asperities. Flowers in
gyrate racemes (scorpioid).
Calyx inferior, permanent. Coround.

rolla

hypogynous, regular.

corolla.

simple or
USES.

Ovary

Stamens

4-parted, 4-seeded

Nuts

5,

inserted

style simple

upon the
;

stigma

Seed without albumen.


The dye called Alkanet is obtained from the roots
of Anchusa tinctoria and several other
The foliage is
species.
insipid and harmless.
bifid.

TYPICAL GENERA.

4, distinct.

Myosotis, Anchusa, Lithospermum.

151

COROLLIFLORJE.

1. Throat cut
2.
3. Ripe fruit with two of the nuts
Myosotis.
open.
pistil.
4.
remaining, and the scars of two that have dropped off.
perpendicular section
of a nut.

115.

Solanacete.

alternate,

sometimes

Leaves

Herbaceous plants or shrubs.


collateral.

Inflorescence often out of the

;
pedicels without bracts.
Calyx permanent, inferior.
Corolla regular, or somewhat unequal, plaited.
Stamens in-

axil

serted

upon the

corolla.

Pericarp with 2 or 4 cells.


curved in fleshy albumen.

2-celled

stigma simple.

Seeds numerous

embryo usually

Ovary

USES.
Many are narcotic, as Tobacco, Henbane, Stramonium, Bitter-sweet, and Deadly Nightshade, or Belladonna.
The fruit of others is almost free from deleterious qualities,

and eatable as the Aubergine, Solanum esculentum, Tomatoes,


Solanum Lycopersicon, Physalis edulis, and many others.
In some species starch is collected in great quantity, and
renders them fit for food, as in the tubers of the Potatoe, Solanum tuberosum.
TYPICAL GENERA. Solanum, Datura, Physalis.
;

or

Petunia violacea.
1.
cross section of the ovary.
Dulcamara. 3.
section of one of its seeds.

2.

116.

Hydrophyllacea.

Herbaceous

Ripe

fruit

of Solanum

plants.

Leaves

Inflorescence often gyrate.


usually lobed.
Calyx inferior,
Corolla regular.
Stamens
5-cleft, with reflexed appendages.
5, epipetalous.
Ovary simple, 1-celled ; placentae 2.
2-valved.
Seeds reticulated ; embryo cartilaginous.

Fruit

DE CANDOLLE'S SYSTEM.
Unknown.

USES.

TYPICAL GENERA.

Corolla irregular.

Calyx permanent.
1 -celled, in

brown leafless herbs.


Stamens didynamous.
with 2 or more parietal pla-

Parasitical

Orobanchaceee.

117.

Ovary

Nemophila, Phacelia.

a fleshy disk,

Fruit capsular, many-seeded, enstigma 2-lobed.


closed within the withered permanent corolla ; seeds very
centse

minute

USES.

embryo extremely

known

Scarcely

TYPICAL GENERA.
118.

small, in the

apex of albumen.

they are astringent plants.

Orobanche, Lathraa.

Scrophulariacea.

Herbs or shrubs with opposite or

rolla irregular.

Calyx tubular, permanent. CoStamens didynamous, or 2. Ovary 2-celled;

ovules numerous

alternate exstipulate leaves.

stigma 2-lobed.

Fruit 2-celled

seeds in-

is

so lower-

definite or definite, albuminous.

USES.

Foxglove, whose action upon the pulse

ing, is Digitalis purpurea.

and

Gratiola

officinalis,

some Calceo-

are purgative and emetic.


and sub-aromatic. Vandellia
powerful antibilious emetic and febrifuge.
larias,

others,

officinalis is bitter

TYPICAL GENERA.

Scrophularia, Antirrhinum, Pentstemon.

I.
Digitalis purpurea.
corolla split open.
2.
pistil.
tion ot it.
4.
5.
ripe capsule.
vertical section of a seed.

Euphrasia
a

diffusa is

3.

transverse sec-

153

COROLLIFLORJE,

Herbaceous plants or underLeaves opposite, often replete


Flowers in axillary cymes; sometimes

Lamiacete or Labiate.

119.

Stem

shrubs.

with aromatic

4-cornered.
oil.

Calyx tubular, permanent.

solitary.

Corolla bilabiate.

mens didynamous, the 2 upper sometimes wanting.


4-lobed

style

Seeds with

little

Fruit 1

stigma bifid.
or no albumen.
;

to 4

Sta-

Ovary

small nuts.

The

species are always harmless, and in many cases


useful for their tonic aromatic qualities.
Lavender is Lavan-

USES.

for coughs, is Marrubium vulMint,


Marjoram, Thyme, Sage, are all potSavory,
herbs used in cookery.
Teucrium Marum is a powerful and

dula vera

Horehound, used

gare.

The cordial Peppermint is presingular stimulant of cats.


kind of stearoptene resempared from Mentha piperita.
bling Camphor, is found in many species.
TYPICAL GENERA. Lanaium, Salvia, Scutellaria.

3.

Marrubium
The pistil.

An

1.
vulgare.
nut.
5.
4.

2.
corolla slit open.
entire flower seen in profile.
vertical section of the latter, showing the embryo.

Trees or shrubs, sometimes herbaceous


Verbenaceee.
Flowers in opLeaves
opposite, without stipules.
plants.
in dense
sometimes
or
alternately
spiked
posite corymbs,
Stamens didyCorolla irregular.
heads.
Calyx tubular.
120.

namous, occasionally

2.

Ovary

2- or 4-celled

ovules erect

DE CANDOLLE'S SYSTEM.

154

Fruit composed of 2
or pendulous ; style 1 ; stigma bifid.
or 4 nucules in a state of adhesion ; albumen none.
USES.

few are slightly aromatic and bitter.


Verbena, Aloysia, Callicarpa.

TYPICAL GENERA.
121.

Herbaceous plants or shrubs.

AcantJiaceee.

opposite, without stipules.


fascicles, or even solitary.

Leaves

Inflorescence in spikes, racemes,

Flowers usually opposite, placed


Calyx very much imbricated, permanent, inCorolla irregular, 2-lipped.
Stamens mostly 2, some-

within bracts.
ferior.

times didynamous.
Ovary in a disk, 2-celled, 2- or manyseeded ; stigma 2-lobed.
Capsule 2-celled, bursting elasti-

Seeds hanging by hard, usually hooked processes of


cally.
the placentae ; albumen none.
USES.
Acanthus spinosus is accounted emollient. The

and roots of Adhatoda Vasica are supposed to be


antispasmodic. Justicia paniculata is bitter and stomachic.
TYPICAL GENERA. Ruellia, Justicia, Eranthemum.

leaves

122.
Lentibulacea.
Herbaceous plants. Leaves undiFlowers
vided, or resembling roots, and bearing vesicles.
single, or in spikes.
Calyx permanent, inferior. Corolla

with a spur. Stamens 2 ; anthers simwith a free central placenta; stigma


bilabiate.
Seeds without albumen.
Capsule 1-celled.
USES.
Of no importance.
irregular, bilabiate,

Ovary

ple.

1-celled,

TYPICAL GENERA.

Pinguicula, Utricularia.

123.
Plumbaginacea. Herbaceous plants or shrubs.
Leaves alternate, undivided, somewhat
sheathing.
Calyx
inferior, tubular, plaited.

Ovary

superior, 1-seeded

cord

styles 5.

USES.

Statice

astringent roots.
cant.

Corolla regular.
Stamens definite.
ovule pendulous from an umbilical

Fruit a utricle.

Seed inverted.

Limonium, and others, have extremely


The bark of Plumbago is acrid and vesi-

TYPICAL GENERA.

Armeria, Plumbago.

l24.Globulariace<e. Shrubs or herbs.


Leaves alternate.
Flowers in heads.
Calyx inferior, permanent, 5-cleft, sometimes 2-lipped.

Corolla hypogynous,
bilabiate,

made up

of

EXOGENJE COROLLIFLORJE.
Stamens

5 parts.

155

from the tube of the

4,

corolla.

Ovary

Albumen fleshy.
superior, 1-celled, with a pendulous ovule.
USES.
The species are said to be bitter, tonic, and pur;
they appear to be of little importance.
TYPICAL GENUS. Globularia.

gative

Herbaceous plants. Calyx 4-5-cleft,


Stamens inserted upon
permanent,
the corolla opposite its segments.
Ovary 1-celled, with a
125.

Primulaceee.

Corolla regular.

inferior.

free

central placenta

style

with a central placenta.


fleshy albumen.
USES.

The

root of

stigma capitate.

Embryo

Cyclamen

Capsule

lying across the hilum in


is

acrid

the

flowers of

Cowslips sedative. Anagallis arvensis is powerfully acrid.


TYPICAL GENERA. Primula, Anagallis, Lysimachia.

1.

Aretia Vitaliana.

of the latter,

126.

flower cut open.

3.

free central placenta.

5.

2.

showing the

inconspicuous
4-leaved, imbricated.

Stamens 4

USES.

ovules peltate or erect, solitary, twin,

stigma hispid,

in fleshy

The

Corolla membranous, hypogynous, 4filaments flaccid ; anthers versatile.

Ovary without a disk


or indefinite

Embryo

Herbaceous plants, with spiked


and ribbed leaves. Calyx inferior,

Plantaginacea.
flowers,

parted.

4. A vertical section
pistil.
A section of a seed.

The

simple.

Capsule membranous.

albumen.

species are of little importance.

The

seeds of

DE CANDOLLE'S SYSTEM.
and others are mucilaginous; the foliage
Plantago Psyllium
this plant, which
of PI angustifolia is slightly astringent ;
Plantain or Ribgrass, is of some value
is commonly called
for sheep-feed in

dry exposed places.


TYPICAL GENEBA. Plantago, Littorella.
SUBCLASS IV. MONOCHLAMYDE^:.

127.

Under-shrubs or herbaceous plants.


without stipules, often with pellucid dots.

Phytolaccacea.

Leaves alternate,

Stamens indefinite,
of 4 or 5 petaloid leaves.
the calyx, alterof
divisions
the
of
number
or, if equal to the
several
to
1
cells, each
from
of
them.
nate with
Ovary
baccate or dry, 1- or
Fruit
ovule.
1
ascending
containing
Calyx

inferior,

Seeds solitary, with a cylindrical embryo curved


round mealy albumen.
The succulent fruit of Phytolacca decandra is said
USES.

many-celled.

to be useful in chronic
is

and

siphylitic

rheumatism

its

juice

and dangerously purgative.


TYPICAL GENERA. Phytolacca, Eivina.

acrid, emetic,

Under-shrubs or herbaceous plants,


Petiveriacea.
128.
with an alliaceous odour. Leaves alternate, with distinct
with minute pellucid dots.
Calyx of several
stipules, often
distinct leaves.

Stamens perigynous,

indefinite, or, if equal

segments of the calyx, alternate. Ovary superior,


Fruit 1-celled, indehiscent, dry. Seed
1-celled ; ovule erect.

to

the

without albumen
USES.

radicle inferior.

Petiveria alliacea

is

acrid, sudorific,

and emmena-

gogue.

TYPICAL GENERA.

Petiveria, Seguiera.

Herbaceous plants or under-shrubs.


Flowers small. Calyx
stipules.
sometimes tubular at the base, persistent. Stamens inserted
129.

Chenopodiacea.

Leaves alternate without

the base of the calyx, opposite its segments.


Ovary
superior, with a single ovule attached to the base of the

into

cavity.

Fruit membranous.

Embryo curved round

farina-

ceous albumen, or spiral, or doubled up without albumen.


USES.
Spinach, Garden Orach (Atriplex hortensis), Chard
Beet, and Sea Beet, are delicate esculents whose leaves are
eaten boiled.
The roots of common Beet and Mangel Wurzel

EXOGEN;E MONOCHLAMYDEJE.

157

are succulent, sweet, and valuable for food.


The seeds of
are
consumed
for food in
Chenopodium Quinoa
extensively
Peru.
On the other hand, Chenopodium olidum and bary-

osmon

are

emmenagogues Ch. anthelminticum furoil of wormseed


Ch. ambrosioides is

foetid

nishes the anthelmintic

a fragrant expectorant. Several species of Atriplex are reVarious kinds of Salsola and
ported to have emetic seeds.
Salicornia supply the sodas of the shops.

TYPICAL GENERA.

Chenopodium, Atriplex, Blitum.

1.
portion of the spike of Salicornia herbacea, with the flowers lodged in the
flower separate.
flower of Salsola Kali.
2.
3.
4. Its
notches of the axis.
5. The same magnified, with a portion of the leafy dilated calyx torn
ripe fruit.
flower
of
album.
8.
section of the
Chenopodium
away. 6. Its embryo. 7.
same, showing the superior ovary. 9. Its seed cut through to show the embryo.

Stem either herbaceous, shrubby, or


130. Nyctaginaceee.
Leaves opposite, and almost always unequal ;
arborescent.
Flowers having either a common or
sometimes alternate.
Calyx tubular, sometimes coloured ; beproper involucre.
coming indurated at the base. Stamens definite, hypogynous.

Ovary

superior, with a single erect ovule.

Fruit a utricle,

enclosed within the base of the calyx.


Embryo with
aceous cotyledons, wrapping round floury albumen.

USES.

The

fleshy roots of the

species of Mirabilis

slightly purgative.

TYPICAL GENERA.

Mirabilis,

Oxybaphus.

foli-

are

DE CANDOLLE'S SYSTEM.

158

Leaves simple,
Amarantacea. Herbs or shrubs.
in
heads
or
without stipules. Flowers
spikes, usually coloured.
in
immersed
dry coloured bracts.
Calyx scarious, persistent,
131.

Stamens hypogynous.
Ovary superior, 1- or few-seeded ;
Fruit a utricle.
central funiculus.
from
a
free
ovules hanging
Seeds lentiform albumen farinaceous ; embryo curved round
;

the circumference

radicle next the hilum.

Unimportant. The species are insipid, on which


account some species of Amaranthus have been employed as
Their dry richly coloured flowers render some of
spinach.
USES.

the species beautiful objects of cultivation.

TYPICAL GENERA.
132.

Amaranthus,

Celosia, Trichinium.

Herbaceous plants or under-shrubs.

Begoniaceae.

Leaves alternate, oblique. Stipules scarious. Flowers unisexual.


Stamens
Sepals in the males 4 ; in the females 5.
anthers collected in a head, the connective very
Ovary winged, 3-celled, with 3 double polyspermous

indefinite

thick.

Fruit 3placentae in the axis ; stigmas 3, somewhat spiral.


celled, with an indefinite number of minute seeds; embryo

without albumen.
USES.

Unknown.

TYPICAL GENUS.
133.
nate.

Lauracea.

Begonia.
Trees.

Leaves without

imbricated.

stipules,

alter-

Stamens

Calyx
definite, perigynous; anthers 2-4-celled, bursting by recurved valves.
Glands at the base of the inner filaments.
Ovary superior,
with one or two pendulous ovules. Fruit
Seed withfleshy.
out albumen ; embryo
amygdaloid, with peltate cotyledons.
USES.
All appear to be aromatics,
although some, as Oreofoetens
and others, have the aromatic
daphne
so con4-6-cleft,

The

centrated as to be acrid.

principle
seeds of Nectandra Puchury

and Aydendron Cujumary are the Pichurim beans or Sassafras


nuts, used as a substitute for nutmegs.
Cimiainomum zeylanicum yields cinnamon, and a bark of like nature is

supplied

by many other plants of this order. Camphor is obtained


from Camphora officinarum and the aromatic Sassafras
bark,
used by the
people of the United States as a powerful sudois taken from the root of
rific,
Sassafras officinale.
The
;

EXOGEN.E MONOCHLAMYDE.E.
Avocado

pear,

an eatable West Indian

Persea gratissima.
TYPICAL GENERA.

Laurus,

borne by

is

Cinnamomum.

fruit,

159

A
A

1.
male flower. 2.
female.
3.
stamen, with a gland at
cluster of fruit.
5.
6.
anther, with the recurved valves.
cotyledon seen from within, with the plumula adhering to the inner face.

Litssea Baueri.

the base.

4.

An

Herbaceous plants, rarely shrubs.


134.
Polygonacea.
Leaves alternate, their stipules usually cohering in the form of
an ochrea. Calyx inferior, imbricated. Stamens definite.
Ovary superior, with a single erect ovule. Nut triangular.
Seed with farinaceous albumen
remote from the hilum.
USES.

Rumex

embryo inverted

scutatus, Acetosa,

and

radicle

others, are the Sorrel

plants used in cookery. Rhubarb is the root of several species


of Rheum similar properties, only more feeble, are found in
Rumex alpinus. In addition to acid and purgative qualities,
;

a great degree of astringency manifests itself, as in the roots


of Rumex, and the bark of Coccoloba uvifera, which is said
to yield a kind of Kino. It is reported that the seeds of Poly-

gonum aviculare are emetic, notwithstanding


P. Fagopyrum and tataricum are employed as
places

the leaves of

that those of

food in some

Polygonum hydropiper and

acrid.

TYPICAL GENERA.

Rumex, Polygonum, Rheum.

others are

DE CANDOLLE'S SYSTEM.

160

Polygonum
seed.

1.

lapathifolium.

A flower of Polyg.

3.

2.

flower cut open.

Convolvuli.

4.

The same

vertical section of the

cut open.

5.

A transverse

section of a seed.

135.
juice.

-Myristicacea.

Tropical trees, often

Flowers unisexual. Calyx

trifid.

yielding a red

Ovary

superior, with

Seed enveloped in a
albumen ruminate.
aril ; embryo very minute
many-parted
USES.
Myristica moschata yields the well-known spices
Fruit 2-valved.

a single erect ovule.

mace and nutmeg.

Similar aromatic qualities pervade

the

order.

TYPICAL GENUS.
136.

Myristica.

Proteacea.

Shrubs or small

trees.

Leaves hard,

Calyx valvular. Stamens 4, opposite


dry, without stipules.
the segments of the calyx, and usually imbedded in their
points.
Ovary superior, simple ; style simple. Fruit dehisSeed without albumen.
These are often handsome bushes with densely
capitate flowers, and in Australia are regarded as indications
of bad land but they are of little use.
The seeds of Gue-

cent or indehiscent.

USES.

vina are large, almond-like, and sold as nuts in the markets


of Chili.

TYPICAL GENERA.
137.

Leaves
grant.

Protea, Banksia, Grevillea.

Trees or shrubs with a scurfy surface.


without stipules.
Flowers axillary, often fraMales: calyx 4-parted; stamens 3 to 8, sessile.
Elaagnaceee.

entire,

EXOGEN^E MONOCHLAMYDEJE.

161

Female calyx inferior, tubular, persistent. Ovary 1-celled ;


ovule ascending ; stigma subulate.
Fruit enclosed within the
:

calyx ; embryo surrounded by fleshy albumen.


USES.
The succulent fruit of Elseagnus hortensis and orien-

forms a part of an oriental dessert. That of Hippophae


rhamnoides, the Sea Buckthorn of England, may be eaten.
TYPICAL GENERA. Elaeagnus, Shepherdia.

talis

138.

Stem shrubby.

Thymelacetz.

Leaves without

sti-

pules.
Calyx inferior, tubular, often coloured. Stamens defiin
the orifice of its tube.
nite,
Ovary with one pendulous
ovule.
Fruit nut-like or drupaceous.
Albumen none, or

thin

embryo straight radicle superior.


The bark of the species is generally
;

USES.

caustic

that

Daphne Laureola, the Spurge Laurel, acts as a vesicant ;


the succulent black fruits are dangerous.
Lace Bark, the
of

lintearia, derives its name from the delicate


which are tough, and easily separated by a little
violence.
The same toughness of the fibre is found in many
;
species
Daphne cannabina derives its name from being as
as
Cannabis (Hemp).
tough
TYPICAL GENERA. Daphne, Gnidia, Struthiola.

liber of

white

3.

Lagetto

fibres,

Daphne Mezereum.
The fruit.

139.

1.

Santalacea.

flower cut

Trees,

Leaves alternate, without

open.

2.

shrubs,

stipules.

vertical section of

an ovarv.

or herbaceous plants.
Flowers small. Calyx
M

DE CANDOLLE'S SYSTEM.

162

Stamens

half-coloured, valvate.

the calyx.
tral

Ovary

placenta.

Embryo

1-celled.

Style

single.

in the axis of

4-5, inserted in the base of

Ovules 1-4, attached to a cenFruit 1-seeded, indehiscent.

albumen.

The wood is sometimes fragrant


USES.
obtained from several species of Santalum.
TYPICAL GENERA.

Sandal wood

is

Thesium, Nyssa, Santalum.

AristolocMacea.

140.

Herbaceous

plants

or

shrubs.

Leaves alternate, often with leafy stipules. Wood without


Flowers brown, or some dull colour, hermconcentric zones.

Calyx superior, valvate.

aphrodite.

inferior, 3- or 6-celled

Ovary

Stamens epigynous.
;
stigmas radiatSeeds with a minute

style simple

Fruit 3- or 6-celled, many-seeded.


embryo in the base of fleshy albumen.
ing.

Many are tonic and stimulating. Aristolochia serand


pentaria
fragrantissima are employed as powerful aromatics; others, as A. Clematitis, indica, &c. are emmenaThe Asarums seem to have similar qualities, but
gogues.
more feeble; A. Canadense is called Wild Ginger in North
USES.

America.

In

consequence of their stimulating properties some


are employed as alexipharmics ; the Guaco of the Oronoko,
said to be a specific against the bite of
snakes, is a species of
Aristolochia.

TYPICAL GENERA.

Aristolochia,

Asarum.

141.
Empetracea. Small acrid shrubs with heath-like
evergreen leaves and minute flowers, which are unisexual.
Sepals hypogynous imbricated scales. Stamens equal in number to the inner sepals, and alternate with them.
3:

Ovary

6- or 9-celled

ovules solitary,
ascending ; stigma radiating.
Fruit fleshy, 3- 6- or 9-celled ; the
coating of the cells bony ;
embryo in the axis of fleshy watery albumen.

USES.

Unknown.

TYPICAL GENERA.

Empetrum,

Ceratiola.

l4>2.Euphor1)iacea:.
Trees, shrubs, or herbaceous plants,
abounding in acrid milk. Leaves opposite or alternate,
usually with stipules.
Flowers sometimes enclosed within
often

EXOGENJE MONOCHLAMYDEJE.

163

an involucre, monoecious or

dioecious.
Calyx lobed, someCorolla consisting of petals, or scales, or
Stamens definite or indefinite.
Ovary superior,

times wanting.
absent.

or

2-

ovules solitary or twin; suspended ; styles


to the cells; stigma compound or single.
generally consisting of 3 dehiscent cells, separating
3-celled

equal in

Fruit

number

with elasticity from their


albumen.
USES.

Castor-oil

communis

common

axis;

in

embryo

fleshy

obtained from the seeds of Ricinus

is

Tiglium-oil from that of Croton Tiglium ; and


a similar purgative quality seems to be general in the seeds
;

of the order.

Cascarilla

is

the bark of Croton Eleutheria

and the same aromatic

principle occurs in many species.


are
as
Manchineel, Hysenanche, Sapium
Many
deadly poisons,
aucuparium, Sec. The drastic drug Euphorbium flows from

the stem of some


so

Boxwood,
Buxus sempervirens.
nutritious

substance

Euphorbias in North Africa.


engravers, is the timber of
Cassava, or Mandioc, or Tapioca, a

succulent

useful

to

wood

consisting of starch,

is

obtained from

the stem of Jatropha Manihot, a poisonous plant

but

it

is

purified by washing and torrefaction.


TYPICAL GENERA. Buxus, Andrachne, Cluytia.

1.

The

involucre of a Euphorbia, containing monandrous male florets, surrounding


2. 3. 4. Male florets of different species, with the articulation

a long-stalked female.

6.
vertical
5.
that separates the filament from the pedicel.
carpel separate.
vertical section of a ripe seed, showing the central column
7.
section of an ovary.
and an embryo in the midst of albumen.

DE CANDOLLE'S SYSTEM.

164

Andrachne
the scales at

telephioides.
base.
4.

its

vertical section of

143.

1.

male flower.

A transverse

2.

section of

A female

flower.

an ovary.

5.

3.

A ripe

A pistil with
seed.

6.

it.

Herbaceous plants.

Chlomntkacete.

Leaves opposite with

intervening

Stamens

aphrodite or unisexual.

with a fleshy connective.


Fruit drupaceous.

lous.

stipules.

Stems jointed.
Flowers herm-

lateral; anthers 1-celled,

Ovule penduat
the apex of
minute
Embryo

Ovary

1-celled.

fleshy albumen.

USES.

Chloranthus

and others are powerful arowhich have been used with suc-

officinalis

matics, especially the roots,


cess in dangerous typhus.

TYPICAL GENUS.

Chloranthus.

Leaves
144.
Piperacea. Shrubs or herbaceous plants.
without stipules. Flowers usually sessile in spikes, hermaphrodite.

Stamens

1-celled, containing

definite or indefinite.

a single erect ovule

Ovary

stigma

superior,

sessile,

sim-

Fruit somewhat fleshy, indehiscent.


Seed erect, with
the embryo lying in a fleshy sac or vitellus placed at that
ple.

end of the seed which

is

opposite the hilum, on the outside

of the albumen.

USES.
The pungent aromatic peppers of the shops are
obtained from different species; Piper
nigrum yields black
and white pepper; P. longum the long pepper. Cubebs is
the pepper of P. caninum and others.
P. Betel and me-

thysticum are both intoxicating.


TYPICAL GENERA. Piper, Peperomia.

EXOGEN^E MONOCHLAMYDE.E.

Serronia Jaborandi.

1.

vertical section of the same,

cluster of flowers magnified.

2.

165

ripe fruit,

3.

showing the seed and the position of the embryo.

Herbaceous marsh or water plants.


Flowers hermaphrodite. Stastipules.
mens 6, clavate, persistent.
Ovaries 4, distinct, with solitary
or a 3- 4-celled pistil.
Nuts 4, indeascending ovules
minute
in a fleshy
hiscent or a 3- 4-celled capsule.
Embryo
sac or vitellus, on the outside of hard mealy albumen.
145.

Saururaceae.

Leaves alternate, with

USES.

Unknown.

TYPICAL GENERA.

Saururus, Aponogeton.

Trees or shrubs. Leaves alternate, simFlowers


unisexual, amentaceous.
ple,
stipules.
Ovary
1-celled
erect.
ovules
Fruit coriaceous,
;
numerous,
superior,
Seeds comose ; albumen 0.
1-celled, 2-valved, many-seeded.
146.

Salicaceee.

with

USES.

whose

Various species of Salix are the Willows from


S. alba is a
shoots wicker-work is made.

flexible

tree, and its bark abounds in tannin


and purpurea yield a good febrifugal bark.
The same property resides in Populus tremula, and other
the young buds of Populus candicans
species of that genus
and balsamifera exude a fragrant resin used in medicine
finally, the timber of Poplars is light, clean, and very useful
for purposes in which hardness and strength are not required.

very large fast-growing

S. Russelliana

TYPICAL GENERA.

Populus, Salix.

DE CANDOLLE'S SYSTEM.

166

Populus.

1.

ripe capsule.

2.

Nigra.
6.

Tremula.

3.

male flower.

4.

female flower.

5.

seed.

Leaves alternate,
Platanacea. Trees or shrubs.
147.
with scarious sheathing stipules. Flowers amentaceous, in
round unisexual catkins. Stamens single. Ovaries terminated by a thick style, having the stigmatic surface on one
ovules solitary, or two, suspended.
Nuts clavate.
;
Seeds solitary ; embryo in the axis of fleshy albumen.
USES.
The large species yield a beautiful but brittle and

side

perishable timber.

TYPICAL GENUS.
148.

Platanus.

Trees, shrubs or herbs, sometimes lacLeaves alternate, usually covered with asperities
or stinging hairs; with stipules.
Flowers small, monoecious
or dioecious.
Stamens definite, often
Calyx membranous.
turned back with elasticity.
ovule
Ovary superior,

Urticacea.

tescent.

simple;

EXOGEN^E MONOCHLAMYDEJE.
solitary,

167

erect or suspended ; stigma simple.


Fruit, a nut.
or without albumen ; radicle always superior.

Embryo with
Division 1.

Urticea.

TYPICAL GENERA.

Flowers

loose.

Juice watery.

Urtica, Parietaria.

A
A

I. Branch of Procris splendens.


2. Cluster of male and female flowers.
3.
male flower about to expand. 4. The same expanded. 5.
nut of hemp. 6.
vertical section of it.
vertical section of the ovary of Dorstenia.
7.
8. An acci-

dental double pistil of the same.

Division

2.

Flowers consolidated.

Artocarpea.

Juice

milky.

TYPICAL GENERA.
USES.

The

Morus, Artocarpus.

leaves of

Hemp

are narcotic

and the Upas,

(Antiaris toxicaria,) certain kinds of Fig, and many Nettles,


are dangerous narcotico-acrid poisons.
The deleterious principle

is,

however, so

little

developed in some that they befor food, as the fruit of the

come harmless, and are used

common
several

Fig, the Mulberry, the Bread-fruit, (Artocarpus,) and


Even the milky juice, which is generally

others.

very acrid, is bland in some cases, especially that of the


Cow-tree of Equinoctial America, on which the natives feed ;
it
always abounds in Caoutchouc, which is obtained in large
quantities from many kinds of Fig. The Banyan-tree of India
is
is

Ficus indica.
also

common

The toughness

of fibre found

in other species, especially

some

in

Hemp

nettles

and

DE CANDOLLE'S SYSTEM.

168

Broussonetia papyrifera.
Hops, so valuable for their bitterand
bracts
the
of
ripe fruit of Humulus Lupulus.
ness, consist

Morus

alba.

1.

female flower
3.
2. Clusters of females.
5.
vertical section of
part of the calyx cut away.
cluster of fruit consisting of baccate calyxes.

male flower.

The same with a

4.

separate.

a ripe achEenium.

6.

Trees or shrubs. Leaves alternate, with


Flowers
amentaceous, monoecious ; males
unisexual,
stipules.
sometimes having a calyx. Stamens definite, usually distinct.
Betulacea.

149.

Ovary

superior,

branous,
none.

2-celled

indehiscent,

Fruit mempendulous.
Seeds pendulous; albumen

ovules

1-celled.

Timber trees, furnishing a


TYPICAL GENERA. Betula, Alnus.
USES.

150.

Leafy shrubs, with resinous glands and


Flowers unisexual, amentaceous,
Stamens 6 or 8. Ovary 1-celled, surrounded

alternate.

achlamydeous.

by

kind of wood.

Myricacea.
leaves

dots,

light

several

hypogynous

scales

ovule solitary, erect

Fruit drupaceous, or dry and dehiscent.

2.

erect

Seed

stigmas
solitary,

radicle superior.

Aromatic shrubs.

Sweet Gale, Myrica Gale, is


as a substitute for hops.
The berries of the
M. cerifera, the Candleberry Myrtle, secrete a natural wax
from their surface ; its roots are astringent. Comptonia asUSES.

used

in

Sweden

plenifolia is

used

in

TYPICAL GENERA.

North American medicine


Myrica, Comptonia.

in diarrho?a.

MONOCHLAMYDE^l.
151.

Corylacea or Cupuliferee.

with

Trees or shrubs.

169
Leaves

Flowers unisexual, amentaceous.


stipules,
Stamens 5 to 20. Ovary crowned by the rudiments of a
calyx, within a coriaceous involucre, with several cells and
several ovules ; ovules pendulous.
Fruit a 1 -celled nut, enalternate.

closed in the involucre.

Seed

solitary.

The Beech, valuable for its timber, and its nuts


called mast, is Fagus sylvatica.
Hazel-nuts are the fruit
of Corylus Avellana.
Sweet Chesnuts are borne by Castanea
USES.

vesca.

Oaks, the most valuable of European trees, are va

rious species of Quercus.

TYPICAL GENERA.

Fagus, Quercus.

3. The latter, with the


2. Female do.
1. Male catkins.
sylvatica.
male
4.
of the involucre stripped off to show the ovaries at the apex.
5.
flower.
half-grown female, with the involucre, now consisting of consolidated
7.
the nuts.
fi.
sculps, forced back.
ripe involucre opening .and exposing
8. The same of a young ovary.
transverse section of a ripe nut.

Fagus

scales

DE CANDOLLE'S SYSTEM.

170

Leaves alternate, pinnated,


Trees.
152.
Juglandacea.
Flowers unisexual, amenwithout pellucid dots or stipules.
in the females
membranous
in
the
males
taceous.
;
Calyx
Petals in the males

superior.

Stamens

present.
2- 4-celled

indefinite.

in the females occasionally

Ovary

inferior, incompletely
Fruit drupaceous, 1-celled,
Seed 4-lobed ; radicle superior.

ovule solitary, erect.

with 4 imperfect partitions.


USES.
Trees furnishing excellent timber

and nigra

regia

that of Juglans
;
of Carya alba for purthe former are Walnuts, the

used for gunstocks

is

poses of elasticity and strength

purgative ; that of the common


Walnut when young, made into a preserve with the husk, is
a domestic medicine ; and Juglans cathartica derives its name
latter

from

The

Hickory.

its

fruit is

quality.

TYPICAL GENUS.

Juglans.

53.
Cycadacea. Trees, with a cylindrical trunk, increasing
a
by single terminal bud. Leaves pinnated, gyrate. Flowers
1

dioBcious.

Males monandrous,

in cones.

Females either in
Ovules solitary,

cones, or in the form of contracted leaves.

naked.

Embryo

the midst of albumen,

in

hanging by a

spiral suspensor.

USES.

bitter

starch,

Zamias

which forms
in the

of unknown use exudes from the


the latter contains a great
quantity of
a kind of arrow-root extracted from

gum

trunk when wounded

West

Indies,

and a

sort of

Sago from the

species of Cycas.

TYPICAL GENERA.

Zamia, Cycas.

154.
Taxace/e.
Trees with continuous branches.
Ligneous tissue marked with circular disks.
Leaves usually entire ; sometimes dilated and
lobed, and in those cases having
forked veins.
Flowers mono3cious or
Fidirecious, solitary.

laments monadelphous.
Females ; ovules naked, their outer
skin
hard.
Seed
becoming
hard, either naked or surrounded

by a succulent
ous.

USES.
trees.

cup.

Albumen

fleshy.

Embryo

dicotyledon-

The Yew and several others are valuable timberThe leaves of Yew are foetid and deleterious
;

are said to act

they

medicinally like Digitalis without accumulating

EXOGEN-ffi
in the

system

MONOCHLAMYDE^;.

171

the succulent fruit seems harmless

but the

seeds are said to be dangerous.

TYPICAL GENERA.

Taxus, Dacrydium.

flowers.
1.
male flower. 2. An anther.
female flower.
5. Of a ripe fruit.
4.
vertical section of an ovule.
6. Of a
N.B. 4. and 6. are the same part in youth and age.
ripe seed, showing the embryo.
5. Is the ripe ovule, with an accessory cup.

Taxus baccata loaded with male

3.

155.

Pinacete,

or

Conifers.

branched trunk abounding in


with circular disks. Leaves

Trees or

resin.

entire.

shrubs,

with a

Ligneous tissue marked


Flowers moncecious or

Males monandrous or monadelphous, collected in


a deciduous catkin. Females in cones. Ovary a flat scale.
Ovules naked. Fruit a cone. Seed with a hard integument.
Embryo in oily albumen, with 2 or many opposite cotydioecious.

ledons.

USES.

The timber is of great value Deal, Fir,


wood are produced by various species.
;

Cedar, Larch

Pine,

Tur-

pentine, resin, pitch, and similar substances are obtained from


others ; the resin Sandarach exudes from Thuja articulata.

Juniper-berries are the galbuli of Juniperus communis, and


are diuretic.
Savin, a dangerous emmenagogue, is the Juni-

perus Sabina.

Larch bark

is

equal to that of

ning power.

TYPICAL GENERA.

>t.(

Thuja, Abies, Cupressus.

Oak

for tan-

DE CANDOLLES SYSTEM.

172

Thuja

orientalis.

1.

magnified fragment of a branch bearing a cone of male

portion of a female branch.


vertical section of a ripe seed.

flowers.

2.

3.

4.

Scales with

naked

ovules.

5.

Abies Larix
4.

1.

An

anther.
2.
female scale with ovules.
3.
ripe cone
scale of the latter with a naked seed.
5. Vertical section of seed
and embrm

EXOGE1O3 MONOCHLAMYDE^E.

1.
Cupressus sempervirens.
female cone with naked ovules.

scale of a
3.

173

male cone with pollen.

ripe cone.

4.

2.

scale of

The same with one

of the

scales removed.

CLASS
This class

much

is

II.

ENDOGEN^E.

easy to arrange systematically.


lowing subdivisions may be used
1.

Rhizanthee.

2.

Florida.

3.

Glwmaceee.

last, and much more


For general purposes the fol-

smaller than the

Fungoid parasitical plants.


Leafy plants with the floral envelopes ver-

ticillate.

Leafy plants with the

floral

envelopes im-

bricated.

SUBCLASS

Order 156.

Rafflesiacea.

I.

RHIZANTH^;.

Flowers by abortion

dioecious.

Perianth superior, 5-parted, imbricated ; the throat surrounded by calli. Column adhering to the tube of the perianth ;
anthers numerous, 2-celled, opening by a vertical aperture.

Ovary

inferior, 1-celled,

with many-seeded parietal placentae

styles conical.

USES.

Astringents

TYPICAL GENERA.

scarcely

known.

Rafflesia, Pilostyles.

DE CANDOLLE'S SYSTEM.

174

the top of a stalk


157.
Cytinaceae. Flowers monoecious, at
Perianth tubular, with a spreading
covered with scales.
limb.
Column fleshy, thickened at the point, covered by
anthers.

Anthers

8, 2-celled.

Ovary

inferior, 1-celled,

with

8 parietal placentae. Style simple, joined to the tube of the


perianth by septiform processes ; stigma capitate, thick.
USES.

Unknown.

TYPICAL GENERA.

Cytinus, Aphyteia.

Balanoplioraceec. Flowers monoecious, in dense heads.

158.

Calyx deeply 3-parted, equal, spreading, sometimes imperfect.


Stamens 1-3, epigynous.
Ovary inferior, 1-2-celled, 1-2seeded

convex.

ovule pendulous.
Style 1 ; stigma simple, rather
Fruit 1-celled, containing spores collected in a bag

resembling a seed.
USES.
Cynomorium coccineum was formerly used as an
astringent, under the name of Fungus Melitensis.

TYPICAL GENERA.

fu

"giforme
},Kbetween them
;
hairs

4.

pendulous ovules.

6.

Balanophora, Cynomorium.

SUBCLASS

I59.ffydrocharacex.
herbaceous.

male P lan t- 2.
female.
3. Male flowers with
Females. 5.
vertical section of a
female, with the two
section across a ripe fruit.
7. Seeds.
_}

II.

FLORIDA.

Floating or water-plants.

Petals 3, coloured.
Ovaryl- or many-celled;

Stamens

Sepals

3,

definite or indefi-

stigmas 3-6; ovules often

ENDOGENJE FLORIDA.
parietal.

Seeds without albumen;

175

embryo undivided,

anti-

tropous.

USES.

Unknown

TYPICAL GENERA.

Hydrocharis, Stratiotes.

Aromatic, tropical,
Zingiberacea or Scitaminea.
Leaves with divergent veins.
herbaceous plants.
Calyx
Corolla irregular, with 6 segments in 2
superior, tubular.
160.

whorls.

Stamens

of which

3,

Filament not petaloid.

Anther

the

2 lateral are abortive.

2-celled.

Stigma

dilated,

Fruit usually capsular, occasionally berried.


Seeds
with or without an aril ; albumen floury ; embryo enclosed

hollow.

within a vitellus.
USES.

Zingiber

Aromatic stimulants.
officinale

Cardamomum and

Cardamoms

Ginger
are

the

is

the

fruit

rhizoma of
of Elettaria

Grains of Paradise, or Meleguetta


Turmeric, Galangale,
pepper,
and Zedoary are other products of the order.
TYPICAL GENERA. Alpinia, Hedychium.
others.

are furnished

by Amomums.

Flowers of Kaempferia pandurata. 2. The inner row of the corolla seen in pro4. The
3. The anther, enclosing the apex of the style between its lobes.
5.
transverse section of
style and stigma, with two abortive stamens at the base.
the ovary.
6. Ripe fruit of Ceylon Cardamoms, Elettaria Cardamomum Zeylanicum
of Pereira.
seed.
8. The same cut through to show the embryo seated in
7.
1.

file.

vitellus.

Herbaceous plants, in tropical countries


Leaves often articuoften growing on trees and rocks.
161.

Orchidacea.

DE CANDOLLE'S SYSTEM.

176

Petals 3, of which 2 are


Sepals 3.
Stamens 3, united in
the
undermost.
and
1,
lip,
uppermost,
a column, the 2 lateral abortive, the central perfect, or the
central abortive, and the 2 lateral perfect
pollen powdery,

with the stem.

latecl

or cohering in masses.
Ovary 1-celled, with 3 parietal placentae ; style a part of the column of the stamens ; stigma

a viscid space in front of the column.


minute.
USES.

The

roots of Orchis mascula

Seeds very numerous,

and others contain a

large quantity of hard mucilage, and form a nutritious substance called Salep.
The fragrant Vanilla is the succulent
fruit

of Vanilla planifolia.

The corm of Bletia verecunda

bitter; the expressed juice of

be purgative.
TYPICAL GENERA.

Orchis,

Epidendrum bifidum

is

is

said to

Epidendrum, Spiranthes, Onci-

dium.

AAA

fl Wer
Seen
P Uen m

skin

in

P rofile

'

"

The

en from
"

A SCpd

ENDOGENJE FLORID JE.

377

Marantacea. Herbaceous tropical plants destitute


Leaves with divergent veins. Calyx superior, of

162.

of aroma.

sepals.

Stamens

Corolla irregular, with the segments in 2 w~horls.


petaloid, of which one of the laterals and the

3,

intermediate are barren or abortive.

Filament petaloid;

Seeds
Stigma cucullate, and incurved.
without aril albumen hard ; embryo naked.
USES.
Maranta arundinacea and some others form a large
quantity of pure starch in their tubers, and this, when extracted, forms arrow-root ; the leaves of both this and the
anther

1-celled.
;

Zingiberaceous order are plaited into baskets by the Indians.


TYPICAL GENERA. Maranta, Canna.
163.
Musaceee.
Leaves with divergent veins, sheathing at
the base, and forming a kind of spurious stem ; often very
Flowers spathaceous. Perianth 6-parted, petaloid,
large.

2 rows.
Stamens 6, some abortive ; anthers 2-celled.
Stigma usually 3-lobed. Fruit either a 3-celled capsule, or
succulent.
Embryo in the axis of mealy albumen.
in

USES.

The

large fleshy fruits filled with starch in

Musa

are the Plantains and Bananas of tropical countries, where


they furnish the inhabitants with an abundance of most nutritious food.

TYPICAL GENERA.

Musa,

Strelitzia.

Amaryttidacea. Generally bulbous, sometimes fibrousLeaves ensiform.


rooted, occasionally with a lofty stem.
and
Stamens 6 ;
corolla
coloured,
superior.
Calyx
equally
1

64.

anthers bursting inwardly.


or corneous.

Stigma 3-lobed.

Albumen

fleshy

toxicarius, and many others, have


deleterious principle in a diffused state
renders them simply emetic, as in Narcissus, several species of
which possess this quality ; or purgative, as Oporanthus luteus.

USES.

Hsemanthus

poisonous bulbs.

The

In some Alstromerias with fleshy roots a large quantity of


starch exists, which, when freed from impurities, forms a sort
of arrow-root. Agave Americana, the American Aloe, as it is
miscalled

by gardeners, abounds, when flowering, in a sweet


becomes an intoxicating liquid,

sap, which, being fermented,


called Pulque.

TYPICAL GENERA.

Amaryllis, Oporanthus, Narcissus.

DE CANDOLLE'S SYSTEM.

178

Pancratium maritimum.

1.

flower cut open, and showing that there is a bifid


transverse section of the

2.
tooth, forming a coronet or cup, between each stamen.
3.
section of the seed, showing the embryo.
ovary.

165.

Leaves alternate,
Twining shrubs.
Flowers
minute, dioecious.
petiole.
Stamens
6.
superior.
Ovary 3-celled,

Dioscoreacea.

netted, with a distinct

Calyx and corolla


with 1- or 2-seeded

Fruit leafy,
;
style deeply trifid.
near the
succulent.
small,
compressed, occasionally
Embryo
cells

hilum, in a large cavity of cartilaginous albumen.


USES. The roots of many species of Dioscorea abound in
starch,

and are the

Yams

instead of Potatoes.

used for food in tropical countries


is present a highly

Nevertheless there

deleterious principle, that, when concentrated, renders these


plants dangerous. The root of Tamus communis is very acrid ;

and even some Yams are too nauseous to be used for food,
even after careful cooking.
TYPICAL GENERA. Tamus, Dioscorea.

ENDOGEN/E FLORIDA.

179

Herbaceous plants or under-shrubs. Stem


Leaves usually equitant. Calyx
and corolla confounded, sometimes irregular. Stamens 3, from
166.

Iridacea.

often a rhizoma or cormus.

the base of the sepals ; anthers bursting externally.


Stigmas
Albumen corneous, or densely fleshy.
3, often petaloid.

USES.

which,

Crocus sativus has long orange-coloured stigmas,


Orris-root is the slightly
dried, form saffron.

when

stimulating aromatic rhizoma of Iris florentina and others


that of I. pseudacorus is acrid, purgative, and emetic.

TYPICAL GENERA.

Crocus vernus.
tion of the ovary.

1.

4.

Iris,

Crocus, Tigridia.

flower split open.

section of the seed to

2.

The stigmata. 3.
show the embryo.

transverse sec-

Stemless or short-stemmed plants, with


scurf.
rigid channelled leaves often covered with cuticular
coloured.
Petals
herbaceous.
Calyx 3-parted, superior, usually
167.

Stamens

Bromeliacea.

6,

or more.

Stigma 3-lobed, or

entire, often twisted.


N 2

DE CANDOLLE'S SYSTEM.

180

Seeds numerous; embryo taper, or minute, in the base of


mealy albumen.
The sub-acid fragrant fruit of Ananassa sativa is the
USES.

The dry filamentous stems of Tilwell-known Pine-apple.


landsia usneoides are used in tropical countries for stuffingmattresses.

TYPICAL GENERA.

Bromelia, Tillandsia.

Herbaceous plants or under-shrubs, with


Stems woody. Flowers hermaphrodite
or dioecious. Calyx and corolla inferior, 6-parted. Stamens 6.
Ovary 3-celled stigmas 3. Fruit a roundish berry. Albumen between fleshy and cartilaginous.
The diuretic demulcent called Sarsaparilla is the
USES.
root of several species
others have a large fleshy root possessing similar properties, and called Chinee radix : it appears to
be nutritious. The leaves of Smilax glycyphylla are bittersweet, and are used for tea in New Holland.
168.

Smilacex.

a tendency to climb.
;

Smilax glycyphylla. 1.
male flower seen from above. 2.
female flower.
3.
transverse section of an ovary.
4.
seed.
5.
section of a seed, showing
the embryo.

169.
Liliacetz.
Boots fibrous or fasciculate. Stem none
a bulb; or tuberous, or creeping, or arborescent.
Calyx and
corolla inferior, coloured,
Stamens
6.
Anthers
regular.
open;

ENDOGEN^l FLORIDA.

181

ing inwards.
Ovary 3-celled; stigma simple, or 3-lobed.
Fruit 3-celled.
Embryo in the axis of fleshy albumen.

USES.
nalis.

and

Asparagus

is

Squills, so well
diuretic qualities,

the

young shoots of Asparagus

known
are

offici-

their expectorant, emetic,


roots of Squilla maritima.

for

the

What

are called Alliaceous plants are found here in the form


of Garlic, Onions, Chives, Leeks, and Rocambole, all species of
Alliuin.

socotrina
is

The purgative drug Aloes is an extract from Aloe


and other species.
The Dragon-tree of Teneriffe

an arborescent form of the order, and yields an astringent


Gum Dragon.
TYPICAL GENERA. Tulipa, Fritillaria, Hyacinthus.

substance called

1. A flower magnified.
A vertical section of a seed.

Arthropodium paniculatum.
transverse section of

170.

it.

4.

MelantliacetK.

Rhizoma sometimes a

Roots

ripe capsule.

3.

sometimes fascicled.
Leaves sheathing at the

fibrous,

fleshy corm.

2.

DE CANDOLLES SYSTEM.

182
Perianth

base.

in 6

inferior,

anthers opening outwards.

style trifid or 3-parted.

bumen

Stamens

pieces, or tubular.

Ovary

3-celled,

many-seeded

Capsule divisible into 3 pieces.

Al-

dense, fleshy.

Poisonous plants.
Meadow Saffron (Colchicum auan
acrid
narcotic
and cathartic. White Helletumnale)
bore the root of Veratrum album, Cebadilla produced by
USES.

is

Asagrsea officinalis, and the roots of various kinds of Trillium


and Helonias, possess similar properties.
TYPICAL GENERA. Veratrum, Colchicum.

tZr mg ,Theova L V^
'

coa^s

r
SonenLl T
t

6S

floral

'

"F

*****.

'~

uncacea; '

Mowers

unisexual.

2 The

""

stri

envelopes are cut

away

5.

;
open to show the stamens
and the 3-parted

in flower '

nes a ft er the

ctfon of

"

PPd

section of a seed.

3.

of

its

outer

A transverse

6.

The

flower

style.

Herbaceous plants, with fascicled or fibrous

generally

brown or green, hermaphrodite

Calyx and corolla more

or

less

glumaceous.

ENDOGENJE FLORIDA.
Stamens

6,

sometimes

3.

Ovary

1-

or

183
3-celled.

Stigmas
Seeds neither

Fruit capsular, with 3 valves.


generally 3.
black nor crustaceous ; albumen firm ;

USES.

Unimportant.

Used

for

embryo within it.


making mats and similar

objects.

TYPICAL GENERA.
172.

Juncus, Luzula.

Commelynacete.

sheathing.

Sepals

Herbaceous

inferior,

3,

sometimes cohering at the base.


deformed.

from the hilum

Capsule 2- or 3-celled.
embryo pulley-shaped, in a cavity remote

albumen

TYPICAL GENERA.
73.

coloured,

Stamens hypogynous, some

fleshy.

Unknown.

USES.

Leaves usually
Petals

3-celled; stigma 1.

Ovary

Seeds often twin

plants.

herbaceous.

Commelyna, Tradescantia.

Butomacete.

Aquatic plants.

Leaves very

cellular,

often milky. Sepals 3, inferior, herbaceous. Petals 3, coloured.


Stamens definite or indefinite. Ovaries 3, 6, or more. Follicles many-seeded.
Seeds minute, attached to the whole of

the inner surface of the fruit.

USES.
Unimportant.
TYPICAL GENERA. Limnocharis, Butomus.

Palmaceee.
Stem simple, rarely forked.
174.
Leaves
terminal, very large, pinnate, or flabelliform, plaited in vernation.
Spadix enclosed in a valved spatha. Flowers small,

hermaphrodite, or polygamous. Perianth 6-parted, persistent.


Stamens inserted into the base of the perianth, definite or

Ovary 3-celled, or deeply 3-lobed, with an erect


Fruit baccate or drupaceous. Albumen cartilaginous
or fleshy ; embryo in a cavity at a distance from the hilum.

indefinite.

ovule.

The Cocoa-nut, whose whole structure appears useindependently of its agreeable fruit, is the produce of Cocos
nucifera ; the tough coarse fibre of this plant is manufactured
USES.

ful,

The Date-tree is
called Coir-rope.
the Phoenix dactylifera.
Sago, a nutritious starchy substance,
is secreted in the trunks of several species, especially of Sagus
into the elastic cables

The sugary nature of their sap, and


its great abundance, enables the natives of Palm countries to
obtain an intoxicating beverage called Palm wine from others.
Isevis

and Caryota urens.

DE CANDOLLE'S SYSTEM.
The Palms of Scripture were the

leaves of the Date-tree.

foliage

and hard,
of the order generally, being large

suited

to

is

The
well

The Canes, whose


such purposes as thatching.
into
woven
chair-bottoms, are
are
flexible stems when split
different species of Calamus.
TYPICAL GENERA.

Phoenix, Chamaerops.

Jft

I.

2.
Inflorescence of Chamaerops humilis, in its spathe.
portion of the same
3.
male flower. 4.
female flower.
fruit ripening.
5.
ripe fruit.
seed with a portion of the
section of another variety, showing the seed.
7.

with the
6.

surface cut

away,

175.

to display the

Juncaginacea.

embryo.

Herbaceous bog-plants.

Flowers

siform.

Leaves enand petals both

inconspicuous.
Sepals
Stamens 6. Ovaries 3 or
herbaceous, inferior, rarely absent.
Fruit dry ; albumen
6, cohering firmly ; ovules 1 or 2, erect.

wanting
USES.

embryo orthotropous, with a

lateral cleft.

Unknown.

TYPICAL GENERA.

Triglochin, Scheuchzeria.

ENDOGEN^E FLORIDA.

185

Alismacete.
176.
Sepals 3,
Floating or swamp plants.
Petals 3, petaloid.
Stamens definite or
herbaceous, inferior.
indefinite. Ovaries several, 1-celled. Ovules ascending. Fruit

not opening, 1- or 2-seeded.


Embryo doubled upon itself.
USES.
The leaves are acrid. The rhizoma of the Arrowhead, Sagittaria,

is

eatable.

TYPICAL GENERA.

1.

Leaf of Ouvirandra
O. Bernieriana.

Alisma, Sagittaria.

fenestralis.

carpel of

4, 5.

2.

Embryo

flower cut open.

in different positions

3. Section of
:

a ripe

the thicker part

is

the cotyledon, the smaller the plnmula.

177.

Acoracea. Rhizoma jointed. Leaves ensiform. Flow-

ers hermaphrodite, surrounded with inferior scales.


Spathe
leaf-like.
Stamens with 2-celled anthers, turned inwards.

Seeds albuminous.
on
one
side.
Embryo
USES. Acorus Calamus was the sweet rush with which the
rooms of the higher orders were strewed before the introduction of carpets, &c.
It has a fragrant rhizoma, whose
aromatic qualities have rendered it useful in medicine it is,
however, chiefly employed as an ingredient in hair-powders.
TYPICAL GENUS. Acorus.
Ovaries distinct.

with a

Fruit finally juiceless.

cleft

178.

very

Leaves
Water-plants.
Flowers inconspicuous, hermaphrodite or uniPerianth of 2 or 4 pieces, rarely wanting. Stamens

Naiadaceee or Flumales.

cellular.

sexual.

Ovaries 1 or more, superior ; ovule pendulous.


Fruit not opening, 1-celled, 1-seeded.
Albumen none ; embryo antitropous, with a lateral cleft.
definite.

DE CANDOLLE'S SYSTEM.

186
USES.

Unknown.

TYPICAL GENERA.

Zannichellia

palustris.

Potamogeton, Zanuichellia.

1.

flower.

ovary opened to exhibit the ovule.


folded up embryo.

4.

An

3.
cluster of ripe ovaries.
vertical section of a seed, showing the

2.

Herbaceous or shrubby, stemless or arborescent plants.


Leaves with parallel or branching veins ;
sometimes compound. Spadix generally enclosed in a spathe.
Flowers unisexual. Perianth wanting. Stamens definite or
179.

Araceee.

indefinite,

'

very short.

Ovary

1-celled,

very seldom 3-celled

ovules erect, or pendulous, or parietal. Fruit succulent.


bryo in the axis of albumen, with a cleft in one side.

USES.
the

Dumb

Em-

Acrid plants which are sometimes dangerous, as


cane, Dieffenbachia Seguina, which paralyses the

muscles of the mouth

if bitten.
Nevertheless, by cooking, this
that the leaves of Colocasia
diminished,
acridity
esculenta and others are used in tropical countries instead of
is

so

much

187

ENDOGENJE FLORIDA.

Some, too, secrete large quantities of starch,


Cabbages.
becomes fit for
when
which,
separated from the acrid matter,
food, as in

Arum

maculatum.

TYPICAL GENERA.

1.

Spathe of

A transverse

Arum

Arum, Dracontium, Caladium.

maculatum.

Its spadix loaded with flowers.


6.
cluster of ripe fruits.

2.

an ovary. 5.
section of the same, showing the embryo.
4.

section of

3.

An

seed.

anther.
7.

180.
Herbaceous plants, growing in marshes
Typhaceae.
or ditches. Leaves rigid, ensiform.
Flowers unisexual, upon
a naked spadix. Sepals 3, inferior, sometimes a bundle of
hairs.

Petals wanting.

Stamens 3 or 6

anthers wedge-

ovule pendulous ; stigmas 1


shaped.
Ovary
Fruit not opening.
or 2, linear.
Embryo in the centre of
single, 1-celled

albumen, with a
USES.

cleft in

one

side.

Unknown.

TYPICAL GENERA.

Typha, Sparganium.

DE CANDOLLES SYSTEM.

188
181.

Pistiacea.

cular, or lobed stems

Floating plants, with very cellular, lentiand leaves. Flowers from the margin

Stamens definite. Ovary


of the stems, inconspicuous, naked.
Fruit membranous or
with
erect
ovules.
1
-celled,
superior,
capsular, 1- or

more seeded.

Embryo

fleshy albumen, and having a lateral

either in the axis of

cleft,

or at the apex of

the nucleus.

USES.

Acrid plants of no importance.

TYPICAL GENERA.

Lemna,

SUBCLASS

182.

Cyperacete.

Pistia.

III.

GLUMACE.E.

Leaves with their sheaths

Stem

entire.

Flowers consisting of imbricated solitary bracts. Perianth none.


Stamens definite, 1,2, 3, 4, 5, 6, 7, 10, 12 ;
solid.

anthers fixed by their base.


Ovary often surrounded by
bristles ; ovule erect ; style single, trifid or bifid.
Nut crustaceous or bony.

Embryo

lenticular, within the base of the

albumen.

hyp08yi""" bris ""-

'

ENDOGENJE GLUMACE^.

Of no

USES.

189

situations with

other importance than as covering many


a coarse herbage containing but little nu-

tritive matter.

quantity of starch secreted in the tubers


of some species renders them eatable, as Cyperus esculentus
and others. The Papyrus of the ancients was made from the

stems of the Papyrus antiquorum.

few species are

slightly

aromatic.

TYPICAL GENERA.
183.

Scirpus, Schosnus, Carex.

Stems cylindrical, usually fistular.


Graminacea.
with a split sheath. Flowers in little
consisting of imbricated bracts, with distinct glumes

Leaves alternate,
locustse,

Hypogynous scales 2 or 3, sometimes


Stamens hypogynous, 1, 2, 3, 4, 6, or more anthers versatile.
Ovary simple styles 2, very rarely 1 or 3
Albumen farinastigmas feathery. Pericarp membranous.
ceous embryo on one side of the albumen, lenticular.
USES.
The most important of all orders, because the floury
albumen of certain species furnishes man with bread, and

or palese, or both.

wanting.

the nutritious herbage of others is the sustenance of herbiTo the class of Corn belong Wheat, Barley,

vorous animals.

Rye, Oats, Maize, Rice, and

warmer

countries

many other species cultivated in


to that of fodder, Crested Dogstail, various

kinds of Fescue, Foxtail,


cultivated

by

farmers.

Rye

Sugar

Grass,
is

and a number of others

obtained from the juice of

the Saccharum officinarum, whose stem is solid, contrary to


the custom of the order.
Bamboos, whose hard stems are
so valuable in hot countries, are arborescent grasses
60 to 100 feet high and more.
fragrant principle

growing
is found
in Anthoxanthum odoratum and others, especially Andropogon Schoenanthus, called Lemon-grass in the gardens,
which is used as a stomachic in India ; where also an oil,
valued as an external application in rheumatism, is obtained
from the Andropogon Calamus aromaticus, believed to have
been the ancient drug of that name.
The diseased grain

of

Rye

uterus.

Ergot, valuable for its powerful action upon the


Finally, a narcotic quality has been remarked in a

is

few species, especially Lolium temulentum.


TYPICAL GENERA. Agrostis, Bromus, Aira, Lolium.

190

DE CANDOLLE'S SYSTEM.

3. Paleae of
2. Palese and stamens, &c. of the same.
1. Locusta of Agrostis alba.
5. Lo4. Pistil, stamens, and hypogynous scales of the same.
Leersia oryzoides.
6. Paleae, &c. of the same.
7. Locusta of Stipa
custa of Polypogon monspeliensis.
9. Locusta of Cy8. Rachis, bracteae, and florets of Cynosurus cristatus.
pennata.

nodon Dactylon.

10. Paleae,

and abortive

floret of

the same.

Locusta of Corynephorus canescens. 2.


3. Locusta
Paleae, &c. of the same.
4. Locusta of Alopecurus
aquatica.
5. Locusta of Aira
pratensis.
6. Floret of the same.
caryophyllea.
7. Locusta of
Festuca duriuscula.
8.
Locusta of Glyceria fluitans.
9. Floret of the same.
10. Locusta of Eragrostis
1.

of Phalaris

poseformis.

ACROGENS.
CLASS

III.

191

ACROGENS.

Substance of the plant composed of cellular tissue chiefly

ducts only present in the highest orders.


either increasing by an extension of its point, or by
vessels or

spiral

Stem

a developement in all directions from one common point ;


not increasing in thickness when once formed.
Sexual organs
absent.
Reproduction taking place by spores, or by a mere
dissolution of the utricles of tissue.

184.

Plants, with creeping stems, the


Lycopodiacea.
annular ducts.
Organs of reproduction

axis abounding in

axillary sessile thecse, containing either minute powdery matter, or.sporules, marked at the apex with three minute ridges.
USES.
Some are powerful emetics and cathartics, especially

L. Selago and rubrum.

TYPICAL GENERA.
185.

zoma.

mous

Lycopodium, Bernhardia.

Filicales or Filices.
Leafy plants producing a rhiLeaves usually coiled up in vernation, with dichoto-

Thecse or sporangia arising

veins of equal thickness.

from the veins upon the leaves, pedicellate with an elastic


ring, or sessile and destitute of a ring.
Thecse with a vertical, usually
Division 1.
Polypodiacea.
incomplete ring ; bursting irregularly and transversely.
Thecse with a transverse, ocDivision 2.
Gleicheniacea.
casionally oblique ring, nearly sessile,

and bursting lengthwise

internally.

Division

or without

ring,

Thecse with an operculiform


reticulated, striated with rays at the

Osmundacea.

3.

any

bursting lengthwise, and usually externally.


Thecse sessile, without any ring,
Division 4.
DaneEacea.
concrete into multilocular sub-immersed masses, opening at

apex

the apex.
Division

5.

Ophioglossaceee.

Thecse single, roundish, coria-

cellular reticulation, half


ring
opaque,
Vernation straight.
The rhizomata of some are astringent ; that of NeUSES.
phrodium Filix mas has been used as an anthelmintic. In

ceous,

without

or

2-valved.

used as food by the


South Seas.
Polypodium, Pteris, Adiantum.

some countries the pith of the stem

natives, especially in the islands of the

TYPICAL GENERA.

is

DE CANDOLLE'S SYSTEM.

192

JOS

307

303. The same of Schizaca


302. Theca of Trichopteris excelsa (GleicJteniaceai).
304. Part of the frond of Aspidium Lonchitis (Polypo306. Part of frond of Daval305. Bernhardia dichotoma (Lycopodiaceai).
diacece).
308. Part of frond of Aslia pyxidata (Polypodiacea).
307. Theca of the same.
310.
309. Theca of Bernhardia dichotoma (Lycopodiacea).
pidium exaltatum.
scale and theca of the same.
Lycopodium annotinum. 311.

pectinata (Osmundacece).

1 86.
fistular

Leafless branched plants with a striated


Equisetacea.
the
articulations
;
separable, and surrounded

stem

by a toothed

Spiral vessels very few.

sheath.

Inflorescence

consisting of peltate scales. Reproductive bodies in the inside


of the lobes of the scales.
Four clavate bodies, wrapped

round a naked spore.


USES.
The hard, flinty skin renders them
purposes, for which some are used under the
rushes.

TYPICAL GENUS.
1 87.

fluid,

for polishing

name

of

Dutch

Equisetum.

An axis, consisting of parallel tubes.

Characea.

gans of reproduction

and

fit

Or-

succulent globules, containing filaments


and axillary nucules, formed of short tubes, twisted
:

spirally.

USES.

Unknown.

TYPICAL GENERA.

Chara, Nitella.

Cellular plants, having a distinct


Bryaceee, or Mmci.
covered with minute leaves.
Reproductive organs of
two kinds viz. axillary, cylindrical stalked sacs, contain-

188.

axis,

ACROGENS.

193

ing a multitude of particles emitted upon the application of


water ; and thecoo or hollow urn-like cases, covered by a
calyptra, closed by a lid, within which are rows of processes,
called the peristome ; the centre of the theca occupied by a

columella.
filaments,

USES.

Sporules,

when germinating, protruding

which afterwards ramify, and form an

confervoid

axis.

Unknown.

TYPICAL GENERA.

Hypnum, Bryum.

313. Peristome of Tortula


312. Jungermannia bicuspidata (Jungermanniacece).
mralis (Muscaceae).
315. Supposed
314. Theca of Ceratodon purpureus (Muse.)
316. Bryum roseum
representatives of sexual organs in Meesia longiseta (Muse.)
319. Apophysis and
318. Peristome of Octoblepharum albidum (Muse.)
(Muse.)
theca of Splachnum luteum.

189.

Andraacea.

bricated leaves.

Branching moss-like plants, with imwith a calyptra, splitting longi-

Thecse

tudinally into four valves.


to a central columella.

USES.

Peristome

0.

Spores attached

Unknown.

TYPICAL GENUS.

Andrsea.

190.
Jungermanniaceee.
Creeping moss-like plants, either
with imbricated leaves, or with the leaves and axis all fused
into one.
Thecse without an operculum, 4-parted, or 2-4valved.

USES.

Spores mixed with elaters.

Unknown.

TYPICAL GENUS.

Jungermannia.

DE CANDOLLE

194

SYSTEM.

Marclantiacea or Hepatica. Plants composed encellular tissue, emitting roots from their under side,
of
tirely
and consisting of an axis, bordered by a membranous expan191.

sion,

which sometimes forms a broad lobed

thallus.

Repro-

ductive organs consisting of a peltate stalked receptacle, bearof sessile naked thecse, iming theca3 on its under surface ; or

mersed, or superficial.
Unknown.
USES.

TYPICAL GENERA.

Lichenacea. or Lichenes.

192.
in the

kinds

form of a lobed

thallus.

sporules lying in

1,

shields

Marchantia, Riccia.
Perennial plants spreading
Reproductive matter of two

membranous

separated cellules of the

2,

tubes,

immersed

in

medullary layer of the

thallus.

USES.

Several are bitter,

and some have been used as

as Variolaria faginea, and Parmelia parietina. Others


are nutritious, as Iceland Moss, Cetraria islandica.
Roccella
tonics

Orchal, and Lecanora Perellus, Cudbear, used ex-

tinctoria

is

tensively

by

dyers.

TYPICAL GENERA.

Parmelia, Lecidea, Peltidea.

328. Shields of Variolaria amara.


329.
portion of the thallus of the same plant.
330.
piece of the thallus of Sticta pulmonacea with lacunae and soredia.
331.
Thallus of the same,
332. Shield of Opegrapha
333. Thalbearing shields.
scripta.
lus of the same.
334. Shields young and old of Lecanora Perellus.
335. Shields of
Bseomyces rufus. 336. Part of thallus of Peltidea canina.
337. Section of a shield
oi hticta pulmonacea.
338. Podetia of Cenomyce coccinea.
339. Section of shield
ol
Baeomyces rufus. 340. Shields of Endocarpon miniatum. 341. Thallus of the
same.
Chiefly from Greville's Flora Edinensis.

ACROGENS.

195

Algacea or Alga. Leafless plants, with no distinct


growing in water, consisting either of simple vesicles,

193.
axis

or of articulated filaments, or of lobed fronds.


Reproductive
matter either wanting or in the joints of the filaments, or
in thecse of various forms.
in

two opposite
USES.

Spores in germination elongating

directions.
is obtained from
Chondrus crispus, sometimes called
Helminthochorton has been em-

gelatinous matter

nutritious

certain Gracilarias, and


Irish Moss.
Gigartina

ployed as an anthelmintic. They are generally collected


under the name of Wrack for burning for Kelp, formerly the
source of Carbonate of Soda.
The substance sold in the

name of Laver is the Porphyra


and
the Ulva latissima.
vulgaris,
TYPICAL GENERA. Fucus, Conferva.

shops under the

laciniata,

and

321. Vaucheria geminata,


Reproductive organs of 320. Lawrencia pinnatifida.
324. Aspe323. Bonnemaisonia asparagoides.
322. Dictyosiphon foeniculaceus.
326. Lichina confinis.
327. Fucus
rococcus echinatus.
325. Odonthalia dentata.
All from Greville's Algae Britannic*.
vesiculosus.

Fungacea or Fungi. Plants consisting of cellules,


which
filaments are occasionally intermixed, increasamong
194.

ing in size

by addition

going no change after

to their inside

their outside under-

its first

formation, frequently ephemeral.


the tissue, or enclosed in
either
loose
among
Spores lying

sporidia.

o2

DE CANDOLLES SYSTEM.

196

USES.
Agaricus campestris, the common Mushroom, and
some other species of the same gemis, Tuher cibarium, the
Others
Truffle, and many others, are eatable and nutritious.

Amadou is made from Boletus


dangerous poisons.
Great numbers are mischievous parasites, infesting
both live and dead organized matter, and even attacking
are

igiiiarius.

living insects.

the

name

Vast damage

is

committed by them under

of Mildew, Ilust, Brand, Smut, and Dry-rot.

TYPICAL GENERA.

Agaricus, Geastrum, Mucor,

Hypoxy-

lon.

342. Arcyria flava.


343. Geastrum multifidum.
344. Mucor caninus.
345.
Basidia and cystidia of an
346. Agaricus cepjcstipcs.
Agaric.
347. Vcrmicularia
tnchella.
348. Vertical section of
349.
sinuHypoxylon

osum.

From

punctatum.

Greville's

Cryptogamic Flora.

Angioridium

197

THE ALLIANCES OF PLANTS.

V.

THE
ment

following pages explain the author's own views of arrangeand serve as a key to the Natural System of

in 1836,

Botany (Edition 2, London 1836). Although his opinion is much


modified by subsequent consideration, yet he knows from experience that these short characters are of considerable value to
students.

CLASSES.

The whole
which

may

vegetable kingdom

is

divisible into five principal classes,

be characterised as follows

/^Exogens with their seeds


\
in an ovary
naked
'having spiral/ Exogens with
.

vessels

Propagated

by

sexes

EXOGEIXLE.

II.

GYMNOSPERMJE.

III.

ENDOGEN^l.

.....
....

without spiral vessels, or with scarcely

any
Propagated without sexes

They

seeds

VEndogens

I.

are further

known by a

all their principal organs,

thus

IV. RHIZANTH^.
V. ACROGEN^E.

separate consideration of the nature of

ALLIANCES OF PLANTS.

198

CLASS
The

I.

EXOGEN^E.

Subclasses are
their calyx and corolla perfect ; or at
with the calyx highly developed, if the petals are absent

COMPLETE PLANTS ; with both


least

these divide into

POLYPETAL.E, with the petals distinct.


MONOPETALJE, with the petals united into a tube.
INCOMPLETE PLANTS; in which there is no corolla; their calyx
1.

3.

2.

generally either but

little

is

developed or altogether absent.

No division of Exogens has been discovered more in accordance with natural affinities,
than that which depends upon the different degree of developement of the flower ; it is
true, indeed, that its characters are not always constant, and that practical difficulties
arise from the circumstance of some genera belonging to polypetalous orders having no
so on.
petals, while a portion of some monopetalous orders are actually polypetalous, and
Nevertheless the arrangement founded upon the distinctions above recorded appears to be
natural, if the latter are rightly considered.
As understood by me, all those orders in

which the floral envelopes are herbaceous, and


imperfectly developed, belong to Incompletae, whether there are two rows or not, as Me:
nor
as
nispermaceae
ought others,
Euphorbiaceae, to be removed from Polypetalae ; because, although the mass of such orders is polypetalous, certain European genera, with
which we are best acquainted, have no petals. With regard to those polypetalous orders,
in some genera of which the petals cohere by their edges, so as to resemble a monopetalous corolla, the only means of recognising them is by observing that their petals are
scarcely joined at the base ; there is this, however, which assists in removing the diffi:
in true monopetalous orders the
style is scarcely ever divided, except just at the
point, and their fruit is therefore, in all cases, syncarpous ; while, in those polypetalous
which
take on a monopetalous appearance, the fruit is in reality apocarpous, as is
genera,
the case with Anonaceae, Crassulaceae,
The two
Meliaceae, Rutacese, &c.

culty

Leguminosse,

latter,

although syncarpous

when young, yet become

truly apocarpous as their fniit ripens.

SUBCLASS I. POLYPETAL.E.
These comprehend the
following groups
:

1.

Albuminosa:.

Embryo very

considerably shorter

and smaller than the

albumen.
2. EpigynoscB.
disk.
3. ParietostB.

Ovary

Calycosa.

5.

being exterior.
None of the
SyncarposcB.
l-^J*J-t_J_l
1
and
with the carpels

6.

7.

Calyx incompletely whorled; two of the sepals


characters
T

of the other groups,7

OJ."

compactly united.
GynobaseoscB.
Carpels not exceeding five, diverging at the
base, arranged in a single row around an elevated axis, or
Stamens usually separate from the calyx.
gynobase.
None of the characters of the other groups,
Apocarposa.
but with the
their faces
carpels distinct ; or

by

separable

or

third column

is

solitary.

-In the
succeeding pages the first column contains a
der; the second the name of the Order; the third
with some officinal
example in italics within brackets, when any
J

epigynous

Placentae parietal.

4.

II
C gn

usually having an

inferior,

blank, nothing

is

known

brief character of the


sensible properties,

its
is to

of the sensible
property.

be found.

When

ALLIANCES OF PLANTS.

GROUP
Alliance

1.

i.

Herbaceous

Ranales.

199

plants, either apocarpous,

or with parietal placentae.


Floral envelopes in threes
1. Ranunculacese
or fives. Sap transparent.

Sap usually milky

Embryo

enclosed in a vitel-

poisonous

(Black Hellebore,
Aconite).
Cathartic.

Podophylleae
2. Papaveraceac

Floral envelopes in twos or


fours.

Acrid,

Fumariese

Narcotic (JPoppy).
and
Diaphoretic

3.

Nymphseaceas

Slightly astringent.

4.

Nelumbiaceae

aperient.
lus.

Ovaries

Floaters.

Hydropeltideae.

concealed

fleshy receptacle.

in

Wholesome.

Float-

ers.

Stamens perigynous
Alliance 2.

5. Cephalotaceae.

Anonales. Apocarpous

Flowers unisexual, threelobed.


Stamens colum-

6.

woody

plants.
Acrid, aromatic

Myristicaceae

(Nutmeg}.

nar.

Leaves with

with-

7.

Magnoliaceae

Bitter, tonic.

Leaves with stipules and

8.

Winteraceae

Aromatic,

stipules,

out dots.
lant

transparent dots.

Flowers

9.

hermaphrodite,

Bark).
Aromatic (Piper

Anonacese

Stamens

three-parted.

stimu-

( Winter's

(Ethiopicum).

usually distinct.

Leaves without stipules. 10. Dilleniaceae


Flowers pentapetalous.

Alliance 3.

Astringent.

Umbettales. Flowers epigynous, arranged in


bels.
Stems usually hollow.

Carpels two

11. Umbelliferae

Apiaceae.

or

um-

Herb poisonous
(Hemlock) some;

times stimulant

and eatable (Parsley,

Parsnip)

fruit

aromatic

(Anise).
Carpels more than two

12. Araliaceae

Slightly stimulant
(

Ginseny).

ALLIANCES OF PLANTS.

200

Flowers epigynous, arranged in

Grossales.

Alliance 4.

Stems

cemes.

Tonic, or harmless

13. Grossulacese

Placentae parietal

ra-

solid.

(BlackCurrants).
Placentae

central,

many-

14. Escalloniaceae.

seeded.

Placentae central few-seeded


,

Alliance

Br uniaceae.

Berberales. Apocarpous, with the valves of the

5.

anthers curved backwards.


Acid, astringent

16. Berberacese

(Berberry').

Nandinese.

Alliance

6.

Pittosporales.

Dye

yellow.

Syncarpous,

stamens, and the placentae

with

hypogynous

in the centre of the

fruit.

Seeds

Fruit two-celled.
few.

Fruit with more than two

17. Vitaceae

Acidity and sugar


( Vine),

18. Pittosporaceae.

Seeds numerous.

cells.

Stamens
Petals split.

all perfect.

Flowers un-

symmetrical.
Fruit with more than two
cells.
Seeds numerous.
Stamens half sterile.

Stigma

19. Olacaceae.

20. Francoaceae.

21. Sarraceniaceae.

leafy, peltate

group in appearance natural, and agreeing with its technical character in all respects,
with the exception of Nelumbium, which has no albumen ; and the genus Berberis, in
which the embryo is much larger in proportion to the albumen than in any other instance.

Some connecting links are obviously wanting in this group ; and, until it is well
considered, it will appear less natural than it really is, especially if we compare such
Nevertheless, it is to
plants as the Vine with the Crowfoot, or either with Sarracenia.
be observed, that it very nearly agrees with De Candolle's Thalamiflorous subclass, and
that the mutual affinities of the alliances may be demonstrated.
Take Anonales and
Ranales for the centre of the ciisumference of a circle composed of the six foregoing
alliances

Then Anonales

pass into Pittosporales through Cheiranthera ;


Grossales
Ribes ;
Berberales
Berberis ;
Berberales
Umbellales
Nandinea? ;

Pittosporales
Grossales

Umbellales
Ranales

and the

Ranales
Anonales

Thalictrum

Magnolia

relative position of the alliances will

Anonales

....

be thus:
Ranales
Umbellales

....

Berberales.

Pittosporales

Grossales

ALLIANCES OF PLANTS.

201

There is no difficulty in pointing out the various gradations that connect the genera
The most paradoxical
belonging to the orders comprehended in the Albuminous group.
but even
part of the combination is the union of baccate-fruited with dry-fruited plants
Vitacese pass into Umbelliferae through Leea ; and the petals of the genus Vitis itself are
inflected at the points, in the way of Umbelliferae.
:

GROUP
Alliance

II.

EPIGYNOSvE.

.Estivation of corolla not valvate.

1.

Onagrales.
Placenta} central.

Every part of the flower


some regular multiple of two.
None.

22. Onagraceae

Harmless.

Circaeeae.

None.

Halorageae

Alliance

^Estivation of corolla not valvate.


Myrtales.
Placenta} central. Number of parts of the

2.

flower uncertain.
Carpels single. Petals broad

23. Combretacese

Astringent (Myro-

24. Alangiacese

Hydragogues.

25. Rhizophoraceae

Astringent

balan).

Carpels single.

Petals very

narrow.
Stipules between the leaves

(Man-

grove).

Stamens bent downwards.

26. Memecylaceae.

Leaves one-ribbed.
Stamens bent downwards.
Leaves three- or more-

27. Melastomaceae

Slightly astringent.

28. Myrtaceae

Aromatic stimulant

ribbed.

Leaves dotted, with an intrarnarginal vein.

Cajeputi, Cloves) ;

bark astringent.

Flow-

Leaves alternate.

29. Lecythidaceae

Leaves not dotted.

mens

Fruit eatable (Brazil nut).

ers irregular.

Sta-

30. Philadelphaceae.

straight.

Alliance

3.

Cornales.

Leaves with stipules


Leaves without stipules

.
.

Parasitical plants, bearing


their stamens on their
petals.

^Estivation of corolla valvate.


SI. Hamamelaceae.
32. Cornaceae

Tonic.

33. Loranthaceae

Astringent.

ALLIANCES OF PLANTS.

202

Placentae parietal.

GucurUtales.

Alliance 4.

Flowers unisexual

34. Cucurbitaceae

Purgative (ColocyntK); or eatable

of

35. Loasaceae

Stinging.

nume-

36. Cactaceae

(Melon, Gourd).
a

with

Flowers

ring

abortive stamens.
Petals

extremely

Subacid; wholesome.

rous.

Sepals

and

37. Homaliaceae.

alike.

petals

Glands between the stamens.

Petals extremely narrow and nume-

Ficoidales.

Alliance 5.

rous.
38. Mesembryaceae

Alliance 6.

Wholesome.

Flowers unisexual.

Begoniales.

Placentse cen-

tral.

39. Begoniacese

Slightly astringent.

connected by a general natural relationship ; and yet it is


with
extremely difficult to fix the limits of their alliances. They appear to be connected
Melastoma
and Lythraceae, and with the Albuminous group
the Syncarpous group through
by the genus Eupomatia, and even by Caetacese, which evidently touch upon Grossulaceae.
They also pass into Monopetalae by Melastomacese, which join them with GenI entertain no doubt about this being nearly the true position of Begoniaceae.
tianaceae.

These plants seem to be

all

GROUP
Alliance 1.

Cruciales.

Stamens tetradynamous

III.

PAKIETOSJE.

Embryo

Albumen

curved.

40. Cruciferae or

Pungent, stimulant
(Mustard).
Stimulant, sometimes poisonous.

Brassicaceae.

Stamens
Fruit

indefinite

composed

41. Capparidaceae

of three

42. Resedaceae

absent.

None.

carpels.

Alliance

2.

Violales.

Stamens few, with no coronet

to the

flower.

Leaves with stipules

43. Violaceae

Roots emetic ( White


Ipecacuanha).

Leaves dotted
-\ .
44.
Fruit siliquose
.
.
45.
Lea vescircinate when young 46.
Calyx ribbed
.47.
.

Samydacese.
Moringaceae

Pungent, aromatic.

Droseraceae

Subacrid.

Frankeniaceae.

ALLIANCES OF PLANTS.
Alliance

Passionales,

3.

sterile

203

Flowers with a ring or coronet of

stamens.

Petioles generally glandular.

Leaves with stipules


Flowers unisexual .

48. Passifloraceae
49. Papayacese

Subacid.

spread over all


the lining of the fruit.
absent.
Stipules
Ovary

50. Flacourtiaceae

Suspicious.

Placentae

Vermifugal.

52. Malesherbiaceae.

stalked.

Stipules
sessile

absent.

Ovary

53. Turneraceae.

(Coronet 0).

Alliance 4.

Bixales.

Leaves dotted.

Polyandrous.
54. Bixaceae

Purgative and sto-

machic (Arnotlo).
connected with the Epigynous group by Passiflora, and with the Calycose by
Turnera, which passes into Cistaceae. Otherwise its external relationships are not well
marked. The orders themselves are intimately related.

This

is

GROUP
Alliance

Guttales.

1.

IV.

CALYCOSE.

Polyandrous. Albumen absent.


number to the sepals.

Petals

equal in
Leaves simple.

Seeds few.

55. Guttiferae or Clusiaceae.

Fruit sometimes
eatable (Mangostan) ; purgative,
acrid (Gamboge).

Leaves compound.

Seeds

56. Rhizobolaceae

Flow-

57. Marcgraaviaceae.

Seeds eatable (Sa-

few.

pocaya nuts).

Leaves alternate.

ersunsymmetrical. Seeds
numerous.
Seeds nuStyles several.
merous.

Alliance 2.

58. Hypericaceae

Slightly purgative

and

febrifugal.

Polyandrous. Albumen absent.


unequal to the sepals in number.
59. Ternstrbmiaceae

Subnarcotic and astringent

Alliance 3.

Petals

Theales.

Acerales.

Stamens

definite.

Tea).

Flowers unsymme-

trical.

Petals without appendages.


Fruit indehiscent, winged, consisting of two carpels.

60. Aceraceae

Saccharine {Sugar
maple').

ALLIANCES OF PLANTS.

204
Petals

having

scales

in

Leaves and branch-

61. Sapindaceac

es poisonous,

Fruit indehiscent,
consisting of three car-

front.

pels.

disk.

(Litchi).

Petals without appendages.


Fruit dehiscent.

Flowers papilionaceous
Flowers spurred

Alliance

Stamens

fruit eatable

4.

num-

astringent,

febrifugal

Flowers regular.

(Horse-

chestnut).
Bitter, emetic, &c.

63. Polygalaceas
64. Vochyaceae

Cistales.

equal to the

Bark

62. ^Esculaceae

Astringent (Ratanhia root}.

Albumen

present.

65. Elatinaceae.

ber of sepals.

Decandrous,

without

sti-

66. Linaceae

Mucilaginous, tough
(Flax).

pules.

Decandrous, with stipules


Polyandrous, with an in-

67. Hugoniacese.
68. Chlenaceae.

volucre.

Balsamic (Labda-

Polyandrous. Style simple.


Radicle remote from the
hilum.

69. Cistaceae

Polyandrous. Styles many.


Seeds hairy.

70. Reaumuriaceae

nurri).

Saline.

The characters of this group require careful consideration. Many gynobaseous plants
have a calyx imbricated in a similar way, but they are removed by their gynobasic
structure.
The imbricated character of the calyx depends upon this ; that the whorl of
floral leaves is broken, so that about two of the
sepals are out of the place of the others,
and are, consequently, altogether external.

The Calycose passes into the Parietose group


Hugoniaceae.

GROUP
Alliance

1.

Malvales.

by Turnera, and

into

the Syncarpous

by

V. SYNCARPOS^E.

.^Estivation of calyx valvate

carpels

four or more.

Stamens
monadelphous.
Anthers two-celled.
Stamens
monadelphous.
Anthers one-celled.
Anthers bursting by pores.

71. Sterculiaceae

Mucilaginous.

72. Malvaceae

Mucilaginous
(Marsh mallow).

73. Elaeocarpaceae.

Petals lacerated.

Stamens

monadelphous.

74. Dipteraceae

Resinous (Cam-

irregular and enlarged in the fruit.

Calyx

Stamens distinct, separate


from calyx.
Stamens distinct,
growing
on a tubular calyx.

phor).
75. Tiliaceaj
76. Lythraceae

Mucilaginous.
Astringent, acrid.

205

ALLIANCES OF PLANTS.
Alliance

2.

Meliales.

^Estivation of calyx imbricated

pels four or more.

Stamens combined
tube.

into a

77. Meliaceae

Seeds wingless.

Stamens somewhat mona-

78.

delphous. Seeds winged.

Stamens

monadelphous,
with a dilated connective.
Leaves dotted. Fruit sue-

79.
80.

culent.

Stamens
calyx.

growing to the
Disk very large.

81.

car-

ALLIANCES OF PLANTS.

206
Alliance 5.

Silenales.

men

Embryo
or

round

rolled

herbs with leaves

mealy albuhaving tumid

joints.

two

93. Portulacacceae

Insipid, eatable

Sepals four or five, united


into a tube.

94. Silenacea?

Inert.

Sepals four or five, distinct


Dehiscence of fruit loculicidal.
Seeds hairy.
Leaves with stipules

95. Alsinaceae

Inert.

Sepals

(Purslane).

9G. Tamaricaceae

Slightly astringent.

97. Illecebracese

Ditto.

All these orders correspond in so intimate a manner as to leave little doubt of their
Malvales and Meliales are the highest form of the group, Sibeing correctly associated.
lenalcs the lowest ; while Rhamnales on the one hand, and Euphorbiales on the other,
form the connection. The Syncarpous group passes into Epigynosse by Lythracese, and
into Gynobaseosae

by Aurantiaceae.

GROUP
Alliance

Rutales.

1.

VI.

GYNOBASEOS.E.

Style single (or at least

the leaves

dotted).

Gynobase

fleshy.

Carpels

98. Ochnaceae

Tonic, stomachic.

99. Simarubaceae

Bitter

distinct.

Leaves alternate. Stamens


arising from scales.
Stipules 0.

Fruit capsular. 100. Kutaceae

(Quassia).

Bitter, anthelmintic

(Rue); antispasmodic
(Bucku) ; febrifugal
(Angostura Bark).
Stipules present, leaves op-

101. Zygophyllaceae

Sudorific, alterative

Aromatic, pungent.

posite.

Flowers unisexual

.102. Xanthoxylacese

Alliance Z.Geraniales.
point.
Fruit

beaked,

separating

Styles distinct ; at least near the


Carpels combined.

103. Geraniaceae

into five cocci.

Fruit not beaked.

Flow-

ers irregular.

Fruit not beaked.


ers regular.

104. Balsaminacese
Tropaeoleae

Flow-

(Guaiacum).

105. Oxalidaceae

Astringent.
Diuretic.

Pungent (Nasturtium).
Acid.

ALLIANCES OF PLANTS.
Alliance 3.

207

Styles several, and carpels quite dis-

Coriales.
tinct.

Ovules pendulous.
bryo straight.
Ovules ascending.
bryo bent double.

Alliance 4.

Em-

106. Coriariaceae

Em-

107. Surianaceae.

Dyes

FlorJceales.

deep

Fruit poisonous.

Style

Fruit

simple.

divided

into

lobes.

108. Limnanthaceae
This

black.

apparently a natural group

Pungent.

but the student will be likely to confound it with


other groups, unless he pays great attention to its distinctions.
In addition to the remore
or
less
between
the carpels, so as to make them diverge from each
ceptacle rising up
other at the base, it is to be remembered that they fonn only one single whorl, and do
not exceed five in number. If this is neglected, they may be confused with some RoThe group is very incomplete, and may be expected to be much
saceae, Malvaceae, &c.
is

and increased before its orders are finally settled.


Rutales connect this with the Syncarpous group through Luvunga, a genus belonging
to Aurantiacese.
Flb'rkeales distinctly pass into Resales through the genus Florkwi.
Geraniales join this to the Parietous group through Violales, and it is probable that Ru-

altered

tales also lead to the Calycose group.

GROUP
Alliance

Rosales.

Flowers quite regular

VII.

APOCARPOS.E.

Albumen wholly
109. Rosaceee

absent.

Astringent.
Fruit eatable {Ap-

Pomese

ples).

Bark tonic; Prussic


acid
(Lau-

Amygdaleae

rel)

fruit eatable

(Peach).
Astringent (Bur-

Sanguisorbese

net).

Legume-bearing, with the


radicle next the hilum.

110. Leguminosse

or

Leaves and

eatable (Pulse).

Csesalpinieae

Purgative (Senna).
Astringent (Catechu) ; gummy

Mimosese

(Gum
Legume-bearing, with the
radicle remote from the
hilum.
Style from the base of the

2.

Chrysobalanaceae

Fruit eatable.

carpels.

Petals very

numerous

Arabic).

111. Connaracese.

fruit

Fabacese.

113. Calycanthaceae

Fragrant.

ALLIANCES OF PLANTS.

208

Alliance l.Saxales.

Carpels two,

diverging.

Seeds very

numerous with albumen.


Anthers opening by pores.

1 1

4. Baueraceae.

Polyandrous.

115. Cunoniaceae

Leaves opposite. Stipules


between the petioles.
.
Leaves alternate

116. Saxifragaceae

Astringent.

Seeds very numer-

Carpels several.
albumen.
with
ous

Crassales.

Alliance 3.

Succulent plants

117. Crassulaceae

Astringent.

Refrigerant, abster-

gent (Houseleek).

Balsamales.

Alliance 4.
Leaves dotted.

Carpels so-

Abounding
1

8.

in balsamic juice.

Amyridaceae

Fragrant,

resinous

(Gum

Elemi}.
Resinous, poisonous
( Cashew}.

litary.

Leaves not dotted

119. Anacardiacese

This group passes into Albuminosae by Rosacese and Ranunculaceae, and also by Calyand Magnoliaceae ; and into Gynobaseosse by Florkea. It is probable that
the divisions into alliances require much re-examination ; but there can be no doubt about
the close relationship of all the orders comprehended in the group. An unpublished
genus of Cunoniaceae connects this group with Cinchonaceae in Epigynous Monopetalae.

canthaceae

It is obvious from the notes appended to each of the


foregoing groups, that their
mutual relations may be expressed as follows
1. Albuminosae pass into Epigynosae through Eupomatia.
:

2.

Parietosae

Epigynosae

3. Parietosae

6.

Passiflora.

Turnera.

Calycosae

4. Calycosae
5. Syncarposae

Syncarposae
Gynobaseosae
Apocarposae

Hugoniaceae.

Luvunga.

Florkea.
[thaww.
Ranunculaceae and CalycanTheir true relations will therefore be better expressed as follows :
Albuminosae .... Apocarposae.
7.

Gynobaseosse
Apocarposae

Albuminosae

Epigynosae
Parietosae

This subclass

With

is

Gynobaseosrc
.

Calycosae

otherwise allied as follows

Syncarposae.

Incompletae through Rhamnales

With Monopetalae through

to Daphnales.

Euphorbiaceag
Loranthaceae

Empetracse.

Guttiferae

Lauraceae.
Ebenaceae.

Umbelliferae

Galiaceae

Rhamnaceae

Myrsinaceae.

Myristicaceae

Rutaceae
Cunoniaceae
Melastomaceae
>V ith Endogenae
Ranunculacea;
through

Nymphaeaceae

ProteaceaB.

and

Caprifoliaceje,

Ericaceae.

Cinchonaceae.
Gentianacea?
Alismaceae.

Hydrocharacece.

209

ALLIANCES OF PLANTS.
SUBCLASS

INCOMPLETE.

II.

These comprehend the following groups


Rectembryosce.
Calyx very imperfect. Embryo straight.
Acklamydosce.
Calyx and corolla altogether absent.
TMferoscB. Calyx tubular, often resembling a corolla (and with
none of the characters of the other groups).
ColumnosfB.
Stamens monadelphous, and ovary many-(six-)celled ;
or, at all events, the latter character combined with an epigynous
:

2.
3.

4.

flower.
5.

Curvembryosce.
of a horseshoe

or spiral (calyx rarely tubular).

GROUP
Alliance 1.

Female flowers surrounded

RECTEMBRYOSyE.

I.

Flowers

Amentales.
1

cupule.
Female flowers arranged in
scaly catkins.

Urticales.

Carpels several.

in catkins.

20. Corylaceae or

by a

Alliance 2.

or having the form

Embryo curved round albumen ;


;

Cu- Bark astringent


(Oak).

puliferae.

121. Betulacese

Ditto,

Carpel solitary,

or

several.

Stems

continuous, without sheaths.


Leaves opposite. Calyx su-

122. Garryaceae.

perior.

Leaves opposite. Calyx in-

123. Hensloviaceae.

ferior.

Leaves rough.

Anthers

124. Urticaceae

Narcotic, tough

(Hemp).

bursting longitudinally.

Morese

Fruit eatable (Mul-

Artocarpeae

Milky, juice poisonous ( Upas) ; fruit

berry).

eatable (Fig).
Ceratophylleae.

Anthers bursting trans-

125. Stilaginaceae.

versely.

Insipid plants with hypo-

gynous flowers.
Aromatic plants with hypogynous flowers.
Balsamic plants with epi-

gynous

Slightly acrid.

127. Myricaceae

Aromatic, tonic.

128. Juglandaceae

Fruit eatable, laxative (Walnut).

flowers.

N.B. The
to

126. Empetracese

stigma of

show that the

Euphorbiaceae.

Empetrum and

its

true affinity of that plant

hypogynous
is

scales seem,

with Myrica.

It

is

among

other things,

a sort of transition

to

210

ALLIANCES OF PLANTS.

Alliance 3.

Casuarales.

Carpels

Stems jointed

solitary.

and furnished with sheaths.


129. Casuaracese.

Alliance 4.

Leaves rough.

Carpels two.

Ulmales.

130. Ulmaceae

Bitter,

astringent

(Elm).
Alliance

5.

Flowers epigynous
Flowers hypogynous
Of

Seeds numerous.

Datiscales.

131. Datiscaceae
132. Lacistemacese.

Leaves alternate.
Bitter.

the orders in this natural group, Garryacese point to Gnetacese through Chloranand so establish a connection with Gymnospermous Exogens. Their approximaCurvemhryosas by Urticaceae is pointed out under that group. Their relation to

thaceae,
tion to

Achlamydosse

is

demonstrated

by

Ceratophyllese,

Lacistemacese,

Podostemaceae, and

Callitrichaceae.

GROUP
Alliance

1.

II.

ACHLAMYDOSJ3.

Flowers

Piperales.

in spikes.

Leaves opposite, with inter-

133. Chloranthaceae

petiolar stipules.
Leaves alternate. Carpels
several.

134. Saururaceae.

Leaves alternate.

135. Piperaceae

Apocarpous.
Aromatic, stimulant.

Carpels

solitary,

Stimulant, pungent,
aromatic (Pep-

per).

Alliance 2.Salicales.

Flowers

Polyspermous, with comose

in catkins.

136. Salicaceae

Apocarpous.
Bark

* eeds -

Monospermous

febrifugal

(Willow).
.

137. Platanaceae.
138. Balsamaceae.

Polyspermous, with naked


seeds.

Alliance

3.Monimiales.

Flowers

in

an involucre.

Anthers bursting lengthwise .139.


Monimiaceae
Aromatic.
Anthers bursting by re140. Atherospermaceaa Ditto,
curved valves.

Alliance 4,Podostemales.

Carpels two, united.

definite.

141. Podostemaceae.

Seeds

in-

ALLIANCES OF PLANTS.
Alliance

5.

Cattitrichales.

211

Carpels several.

142. Callitrichaceae.
Probably the two last alliances ought to be combined. But it is evident that the
whole group is so
incomplete, that no distribution of the orders is likely to be worth
e P resent
Achlamydosae join Rectembryosse by Garryaceae, Podostemese,
"f o Y?
and
Callitrichaceae ; and Tubiferosfe
by Monimiales.

'

GROUP
Alliance

1.

III.

TUBIFEROS.E.

Flowers epigynous.

Santalales.

143. Santalaceae

Sedative

(Sandal

Wood).
Alliance

Stamens

2.

Daphnales.

Calyx with an imbricated

aestiva-

tion.

Carpels solitary.

distinct.

Leaves

144. Elaeagnaceae

Harmless.

distinct.

Leaves

145. Thymelacese

Caustic bark.

146. Hernandiaceae

Purgative.

147. Aquilariaceae

Fragrant, resinous.

scurfy.

Stamens

smooth.
Flowers unisexual.

Coty-

ledons lobed.

Stamens monadelphous
Alliance 3.

Proteales.

^Estivation of calyx valvate.


148. Proteaceae

Alliance 4.

Laureates.

None.

Valves of the anthers curved back-

ward.
Leafy, arborescent, aromatic

plants,

with

cotyledons.
Leafy arborescent

Aromatic, stomachic

149. Lauraceae

(Cinnamon).

fleshy
plants,

150. Illigeraceae.

with leafy crumpled cotyledons.


Leafless, herbaceous, insipid

151. Gassy thaceae.

plants.

Alliance 5.

Penaales.

Carpels several.
152. Penaeaceae

nauseous,

Sweetish,

gummy,

resinous

(Sarcocol).
Their tubular calyxes distinguish them at once from all the other groups, except
Columnosae
and the latter are in general clearly characterised by their stamens united
into a column.
Tubiferosae touch Achlamydosae by Lauraceae, and Columnosae by Aris;

tolochiaceae.

They

are also strongly related to Curvembryosae

by

Elasagnaceai.

p-2

ALLIANCES OF PLANTS.

212

GROUP
Alliance

1.

IV.

COLUMNOSJ2.

Flowers hypogy nous.

Nepenthales.

153. Nepenthaceae.

Alliance 2.

AristolocMales.

Flowers epigynous.

154. Aristolochiaceae

GROUP
Alliance

1.

V.

Chenopodales.
the hilum.

Tonic, stimulating.

CURVEMBRYOSJE.

Albumen

Radicle next

present.

Flowers dry, with nume-

155. Amarantaceaj

rous bracts.
Flowers herbaceous.

Car-

156. Chenopodiacese

Ditto (Spinach).

Carpels

157. Phytolaccaceae

Emetic.

pels solitary.
Flowers coloured.

Wholesome,

insipid.

several.

Alliance

2.

Albumen present.
Polygonales.
from the hilum.
158. Polygonaceae

Eadicle

away

Acid (Sorrel) ; purgative and tonic


(Rhubarb).

Alliance

3.

Petivales.

Albumen

absent.

Cotyledons

spiral.

159. Petiveriaceae.

Alliance 4.

Sclerales.

Border of the calyx herba-

Tube of

the calyx hardened.

160. Scleranthaceae.

ceous.

Border of the
calyx peta-

161. Nyctaginaceae

Roots purgative.

loid.

Alliance

5.

Cocculales.

Albumen

upon a ternary

present.

Flowers formed

plan, dichlamydeous.

162. Menispermaceae

Root

tonic

bitter,

(Calumbd)
narcotic

seeds

(Coccu-

lus).

vemn
ver, much

relation to them,

and the resemblance in

their calyx is

; they have not, howovercome by the struc-

213

ALLIANCES OF PLANTS.
ture of the seed.

Nyctaginaceae require a

much more

have received.

careful examination than they yet

Menispermaceae have, strictly speaking, both calyx and corolla ; but


their organs are so small and so much
alike, that I place the order here ; it has but b'ttle
apparent relation even to Schizandreae
the circumstance of the

among Anonales, beyond

parts of its flower being ternary, while it seems closely allied to Aristolochiaceae.
Menispermaceae must be considered one of the natural orders
Exogens which tend

towards Endogens.

The passage

podiaceae on the one hand,

among

of Curvembryosae into
Rectembryosae through Chenoother, is obvious.

and Urticaceae on the

The mutual

relations of these groups


may be expressed as follows
Rectembryosae pass into Achlamydosae through Garrvaceae &c.
2. Achlamydosae
Tubiferosae
Monimiaceae.
3. Tubiferosae
Columnosae
Aristolochiaceae.
4. Columnosae
Curvembryosse
Menispermaceae.
5. Curvembryosae
Rectembryosae
Chenopodiacese.
Their true relations will therefore be thus
:

1.

Rectembryosae, Curvembryosae,
Achlamydosae,
Columnosae,
Tubiferosae.

The

subclass of Incompletae
the following manner, viz.

With

Polypetalae

may

be considered

through

allied

with other parts of the system in


to

Daphnalea
Proteaceae
Lauraceae

Loranthaceae.
Myristicaceae.
Euphorbiaceae.

Empetraceae
? Nyctaginaceae

With Monopetalae
With Gymnospermae
With Endogenae

Solanaceae.
Gnetaceae.

Chloranthaceae

SUBCLASS

Rhamnales.

Menispermaceae

Smilaceae.

Aristolochiaceae

Araceae.

III.

MONOPETAL^:.

These comprehend the following groups


Polycarposa. Flowers hypogynous (rarely epigynous).
:

1.

posed of many carpels.


2. Epigynosce.
Flowers epigynous.

Ovary com-

Ovary composed of two

or

many

carpels.
3.

4.

5.

Ovary consisting of but one perfect carpel.


Nucamentosa. Ovary composed of two carpels, which are separate,
nut-like, and often divided into two parts.
Dicarposce.
Ovary composed of two carpels. Fruit capsular.
Aggregoste.

GROUP
Alliance

1.

Brexiales.

I.

POLYCARPOSJS.

Albumen

absent.

Carpels

five.

163. Brexiaceae.

Alliance 2.

Seeds winged.

Brown,

Anthers opening by pores.


from four to five, or more.

Ericales.

Herbs

leafless, parasites

164. Pyrolaceae
165. Monotropaceae.

Carpels

Diuretic, tonic.

ALLIANCES OF PLANTS.
Anthers two-celled.

Seeds

AnThersTwo-celled.

Ovary

6 C. Ericaceae

167. Vacciniaceae

Ditto, ditto.

inferior.

Anthers one-celled

168. Epacridaceae.

Anthers
Alliance S.P-rimulales.
five.
four
Carpels
mens

169. Primulacea.

Sta-

Herbaceous plants.

bursting longitudinally.

narcotic
j
Slightly
~o

(Cowslip).

opposite petals.

Woody

Stamens

plants.

170. Myrsmacese.

opposite petals.

sweet, eatable; bark febri-

and

171. Sapotaceae

Fruit

with twice
stamens as pe-

172. Ebenacese

Astringent.

Calyx
plants.
corolla double.

Milky

fugal.

Watery
as

plants,

many

Fruit

eatable.

tals.

Resinous, astringent, aromatic

Styraceae

(Storax,

Ben-

zoin).

Watery

plants,

same number

with

the

Astringent, tonic

173. Aquifoliacese

(Holly).

of stamens

as sepals.

Alliance 4.

Nolanales.

Fruit divided into deep lobes.

Car-

pels five, or more.

174. Nolanaceae.

Alliance
Leafless

5.

plants.

Volvales.

Embryo

Carpels from two to four.


175. Cuscutaceac, m.

spiral.

176. Convolvulaceae

Erect plants, with an imbricated corolla and three

177. Polemoniaceae.

Twining

plants,
plaited corolla.

with

Roots purgative
(Jalap, Scammany).

carpels.

Styles numerous.

Seeds

178. Hydroleaceee

Bitter,

indefinite.

Ebenaceae
Nolanaceae adjust these to Dicarposae, and Primulaceae to Epigynosse.
touch upon Guttiferae, and Myrsinaceae upon Rhamnaceae through the genus Choripetalum.
Ericaceae moreover have an evident affinity with Rutaceae, first through Ledum, which
may be compared with Phebalium, and secondly through Andromeda, which simulates
Corraea.

215

ALLIANCES OF PLANTS.
GROUP
Alliance 1.

.182.

Alliance 2.

Stamens

179. Lobeliaceae
180. Campanulacese
181. ? Belvisiacese.

distinct.

indefinite.

Acrid, poisonous.

Inert.

Columelliacess.

Seeds

183. Stylidiaceae.
184. Goodeniaceae.

Seeds

185. Scaevolaceae.

Seeds

Stigma with an indusium.

Goodeniales.

Flowers gynandrous

EPIGYNOSJE.

Stipules absent.

Campanales.

Anthers united
Anthers distinct
Polyandrous
Diandrous

II.

indefinite.

Stamens

distinct.

definite.

Alliance 3.

Stipules between the leaves.

Cinchonales,

186. Cinchonaceae

Bark febrifugal (Jesuits'

bark}

root emetic

(Ipecacuanha).

Alliance 4.

Stipules

Caprioles,

Seeds

none.

definite

in

number.
188. Caprifoliaceae

Alliance 5.

Stellales.

no

Fruit double.

Bark

astringent.

Leaves whorled, with

stipules.

189. Stellatae or

Ga-

Astringent,

liaceae.

dying

(Madder).

It is evident that, in this group, Galiacese have a close relationship with Apiaceae ; and
is participated in by Caprifoliaceae, through the genera Viburnum
and Sambucus. Some Primulacese seem to approach Cinchonaceae ; and the Goodenial

that this approximation


alliance,

by means

of Scaevolaceae, passes directly into Brunoniaceae

GROUP
Alliance 1.

Albumen

Asterales.

present

in

the

III.

among Aggregosae.

AGGREGOS^:.

Anthers syngenesious.
190. Calyceraceae.

seeds.

Corolla bilabiate

Corollas all ligulate. Milky.

Involucre

hemispherical.

Florets of ray ligulate.


Involucre rigid or spiny, coFlorets of ray
nical.

tubular and inflated.

191. Mutisiaceae.
192. Cichoracese
193. Asteraceae

194. Cynaraceac

Narcotic (Lettuce).
Bitter, tonic (Cltamomile), diuretic.

Bitter (Thistle).

ALLIANCES OF PLANTS.

216

Anthers

Alliance 2.Dipsales.

Flowers epigynous.

distinct.

.
.195. Dipsacess
Carpel solitary
196. Valerianaceae
Carpels triple; two of them

None.

Bitter, antispasmo-

abortive.

die,

vermifugal

( Valerian).

Alliance 3.

Brunoniahs.

Stigma with an

Style single.

in-

dusium.
197. Brunoniaceae.

Alliance 4.
Fruit

Plantales.

spuriously

Style single.

Stigma naked.

198. Plantaginacese

Bitter, astringent

199. Globulariaceae

Bitter,

double-

celled.

(Plantain}.

Flowers unsymmetrical

tonic,

pur-

gative.

N.B. The situation of the dissepiment in Plantaginaceae


part to be spurious, and that the fruit is in reality quite simple.

Alliance 5.

Plwmbales.

Styles

five.

sufficiently

shows that

Flowers formed upon

a quinary plan.
200. Plumbaginaceae.

Some

tonic, astrin-

gent

others

acrid, caustic.

GROUP
Alliance

1.

IV.

Phaceliales.

NUCAMENTOS.E.
Fruit capsular.

Inflorescence gy-

rate.

201. Hydrophyllaceae.

Alliance

2.

EcMales.
gyrate.

Fruit

deeply lobed

Fruit nucamentaceous.

Inflorescence

Flowers symmetrical.
202. Boraginaceee

Mucilaginous (Borage) ; roots dyeing (Alkanef).

.
byncarpous, style bifid
Syncarpous, style dichoto-

mous

203. Ehretiacese.
204. Cordiacese

Emollient (Selesten

Plum).

ALLIANCES OF PLANTS.
Alliance 3.

Labiates.

Fruit nucamentaceous.

divided

into

four

Inflorescence

Flowers unsymmetrical.

gyrate.
Fruit

217

Tonic, stomachic

205. Lamiaceae or

lobes."

Labiatae.

(Thyme, Mint,
&c.)

Fruit consisting of about 4


cells.
Radicle inferior.
Fruit consisting of about
four

cells.

206. Verbenacese

Slightly bitter.

207. Myoporaceae

Tanning.

Radicle supe-

rior.

Fruit two-celled.

Ovules

208. Selaginaceae.

pendulous. Anthers onecelled.

Fruit two-celled.
erect.

Ovules

209. Stilbaceae.

Antherstwo-celled.

GROUP
Alliance

1.

Bignoniales.

V.

DICARPOS.E.

Neither albumen nor hooks to the

seeds.

.210. Bignoniaceae.
.
Seeds winged
211. Pedaliaceae
Fruit hard and like a nut .
212. Cyrtandraceae.
Placentae 4. Seeds wingless
.

Alliance 2.

Seeds numerous, with albumen.

ScropJiulales.

Leafy plants with a superior

Emollient.

213. Scrophulariaceae.

ovary.

(Digi-

talis).

Leafless plants with a mi-

214. Orobanchaceae.

nute embryo.
Leafless plants with a one-

215. Gesneraceae

celled

Suspicious

Harmless,

ovary, partly in-

ferior.

Alliance 3.

AcantJiales.

Seeds without albumen, with hooks

to the seeds.

Calyx remarkably imbricated.

216. Acanthaceae.

Alliance 4.

Lentibales.

free central placenta.

217. Lentibulaceae.

ALLIANCES OF PLANTS.

218

Flowers

Alliance 5.Gentianales.

standing right and


rescence.
Corolla
fruit

on the

withering

in aestivation

Carpels
symmetrical.
of the axis of inflo-

left

218. Gentianaceae

Bitter (Gentian).

Anthelmmtic.

Milk and

im-

bricated.

Estivation of corolla val-

219. Spigeliacese

vate.

Estivation contorted. Sta-

mens

220. Apocynacese

distinct.

fruit

poisonous

(Nux

vomica)

bark

febrifugal some-

times.

Anthers grown

to the stig-

221. Asclepiadaceae

Emetic,

Acrid.

Oil eatable (Olive).

ma.
Alliance

6.

Oleales.

Estivation of corolla val-

Diandrous.
222. Oleaceae

vate.

Estivation of corolla im-

223. Jasminaceae.

bricate.

Alliance

7.

Flowers unsymmetrical, with seveLoganiales.


ral stamens.

Leaves furnished with

sti-

224. Loganiacese.

pules.

Flowers somewhat pentan-

225. Potaliacese

Emetic.

Acrid.

drous.

Alliance 8.

Embryo

curved.

Solanales.

Flowers

standing

fore

rescence.

Cotyle-

and

symmetrical.
of the axis

aft

226. Solanaceae

Carpels
of inflo-

Poisonous.

dons cylindrical.

cotic

Nar-

(Belladon-

na, Stramonium,
Tobacco).

Embryo

straight.

Cotyle-

227. Cestraceae.

dons leafy.
It appears that the connection

nature

between the foregoing groups

is

of a most decisive

for,
1.

Polycarposae pass into Epigynosae

2-

Aggregosse

3.

Epigynosae
Aggregosae

Nucamentosoc

Dipsaceae.

4.

Nucamentosce

Dicarposae

Scrophulariacese.

5.

Dicarposae

Polycarpos;c

Boraginaceae.

through Primulales.
Scaevolacese.

ALLIANCES OF PLANTS.
The

relations of the groups

may

therefore be expressed thus

Polycarposae,
Epigynosae,

219
:

Dicarposae,

Nucamentosae,

Aggregosae.
regard to the connection of Monopetalous Exogens with other parts of the
system, they appear to have only the following strongly-marked affinities
With Polypetalae through
to
Gentianaceae
Melastomaceaj.
Ebenaceae
Clusiaceae.

With

Galiaceae

Apiaceae.

Caprifoliacese

Myrsinaceae

Rhamnaceae.

Ericaceae

Rutaceae.
Cunoniaceae.

Cinchonaceae

With

Incompletae

It also results

Solanaceae

Nyctaginaceae.

from the previous investigations, that true Exogens are only connected

immediately with other classes by the following points


With Endogens
Ranunculaceae
through
Nymphaeaceae
Menispermaceae
:

Alismaceae.

Hydrocharaceae.
Smilaceae.

Aristolochiaceae

With Gymnospermae

CLASS
Stem

with

articulations.

Araceae.
Gnetaceae.

Chloranthaceae

II.

GYMNOSPERMAE.

228. Gnetacese.

Fruit in spikes.

Stem bearing many buds.

229. Taxaceae

Leaves deleterious

Fruit single.

(Yew).

Stem terminated by a
gle bud.

sin-

Wood

230. Cycadacese

Leaves gyrate

contains

starch.

before developement.

Stem bearing many buds.

231. Pinaceae or Co-

Fruit in cones.

Terebintaceous

niferse.

( Turpentine,
Pitch, &c.)

These plants are connected by close affinity ; but some links


They are in alliance with other parts of the system, thus
to
With Exogens
Gnetaceae
through
With Endogens
Cycadaceae
With Acrogens
Pinaceae

in the chain are wanting

III.

Lycopodiaceae.
Filicales.

Cycadaceae

CLASS

Chloranthaceae.
Palmaceae.

ENDOGENS.

These comprehend the following groups


Anthers distinct. Flowers complete. Ovaiy inferior.
Epigynosce.
Flowers complete.
Gynandrosce. Stamens united with the styles.
:

1.

2.

Ovary
3.

inferior.

Hypogynosce.

Flowers coloured, with

its

parts

in

superior.
4. Retosce.
Leaves netted,

threes.

Ovary

with a taper footstalk articulated with the


Floral envelopes complete.
5. Spadicosce.
Flowers herbaceous, or imperfect. Perianth often absent.
stem.

Embryo with a
6.

Glumosce,

lateral

slit.

Bracts scalelike in the room of a perianth.

ALLIANCES OF PLANTS.

220

GROUP
Alliance 1

EPIGYNOS^E.

Leaves with the veins diverging from

Amomales.

I.

the midrib to the margin.


Monandrous. Anther two-

232. Zingiberaceae

Aromatic, stimu-

233. Marantaceee

Amylaceous,

latin

celled.

Monandrous.

Anther one-

.234.

Musaceae

insi-

pid (Arrowroot).
Fruit nutritious

celled.

Several anthers

(Ginger}.

(Banana).
Narcissales.

Alliance 2.

hexandrous

Hexapetaloideous

plants.

Texture
Flowers large.
smooth.
Plant
Leaves equitant.
woolly.
Leaves equitant.

Fruit

Acrid.

235. Amaryllidaceee

Poisonous.

236. Heemodoraceae.
237. Burmanniaceae.

winged.
Fruit

-celled.

Placentae

238. Taccaceae.

Ixiales.

Triandrous.

parietal.

Alliance

3.

239. Iridacese

Alliance 4.

Bromeliales.

Purgative.

Tripetaloideous scurfy plants (with

albumen).
240. Bromeliacese

Sap sugary (Pineapple).

Alliance 5.

Tripetaloideous smooth plants.

Hydrates.

mens more than

six.

(Albumen

Sta-

absent.)

241. Hydrocharacese.
Both Hydrocharaceae and Bromeliaceae pass into Spadicosae by Pandanaceae. Iridaceae,
particularly the genus Gladiolus, offer a very near approach in structure to Gynandrosae.

GROUP
Ovary

one-celled

II.

GYNANDROS.E.

242. Orchidaceae

Aromatic, viscid,
nutritious (Salep, Vanilla).

The

flowers of a Gladiolus would become those of an Orchis in


calyx and corolla and
if the latter were consolidated with the
style ; here there is a transition to Epigynosse.
Apostasiaceae have the nearly regular flowers of Liliacete, and through them

stamens,

connect this group with Hypogynosae.

221

ALLIANCES OF PLANTS.
GROUP
Alliance

1.

HYPOGYNOS^E.

III.

Palmales.

Hexapetaloideous
Vague embryo.
243. Palmacese

with

plants,

Amylaceous. Saccharine (Cocoa-

nut, Sago).

Alliance 2.

Hexapetaloideous plants, with an embryo in the axis of the albumen.

Liliales.

Petals rolled inwards after

244. Pontederaceae.

flowering.

Hexandrous. Anthers turned outwards. Styles dis-

245. Melanthacese

Cathartic
cotic

tinct.

Flowers irregular, with appendages on the outside.


Hexandrous. Anthers turned inwards. Styles con-

246. Gilliesiaceee.

247. Liliaceae

nar-

diuretic

White Helle-

bore, Colchicum).

Unimportant.

Bitter, stimulant

Asphodelese

solidated.

Alliance 3.

(Squill, Onion, &c.)

Tripetaloideous plants, with the

Commelynales.
carpels three

and consolidated.

248. Commelynacese.

Alliance 4.

Alismales.
pels

Tripetaloideous plants, with the car-

more or

Placentae spread over the


dissepiments.
the
Placentas
occupying
margin only of the dis-

less distinct.

249. Butomacese

Acrid,

250. Alismaceae

Acrid,

sepiments, or their equivalent.

Alliance 5.

Juncales.

.
Flowers regular
Flowers irregular, with a
two-leaved calyx.
.

Here we have a marked

Flowers somewhat glumaceous.


251. Juncaceae
252. Philydraceae.

transition to

Unimportant.

Exogens on the part of Alisma, which is hardly


by its embryo. Liliaceae connect the group
Juncaceae with Glumosae through Restiaceae.

distinguishable from Ranunculaceae, except

with GynandrosEe through Apostasiaccne,

ALLIANCES OF PLANTS.

222

GROUP

IV.

RETOS^E.

253. Dioscoreaceae

Flowers unisexual. Ovary

Nauseous

some-

times eatable

inferior

(Yam).
254. Smilaceae

Flowers hermaphrodite.

Ovary

Diuretic; demulcent (Sarsapa-

superior.

rilla).

255. Roxburghiaceae.

Flowers binary, highly developed.

GROUP
Alliance 1.

V.

SPADICOS^E.

Flowers on a spadix.

Pandales.

Fruit drupa-

ceous.

Flowers

spiral.

ternately

256. Cyclanthaceee.

Spires al-

male and

fe-

male.
Flowers achlamydeous and

257. Pandanacese

Fruit eatable,

apocarpous.

Alliance 2.

Flowers on a spadix.

Arales.

Fruit

either

berried or capsular.
Flowers unisexual
Flowers hermaphrodite
.

Alliance 3.

.258.
.

Araceae .
259. Acoracese

Alliance 4.

Aromatic.

Flowers on a spadix.
Anthers clavate.

Flumales.

Ovules pendu-

Floaters.

Acrid.

Typhales.

260. Typhaceae

Poisonous.

Of no

Flowers in spikes, or
261. Naiadacese

262. Juncaginaceae.
263. Pistiacese

Sepals three.

importance.

solitary.

Unimportant.

lous.

Ovules erect
with none, or

Terrestrial.

Floaters,
scarcely

any,

axis

Acrid (Duck-weed).

of

growth.

we find a transition to Rhizanthae in the case of the


genus Lemna,
vascular system, and is the lowest known form of Endows.
Tyconnect
this
phaceae
group with Glumosae, and Pandales with Epigynosse.
It is here that

which

is destitute of

ALLIANCES OF PLANTS.
GROUP
Stems

fistular

VI.

223

GLUMOS^E.

264. Graminaceae

Fruit floury (Corn).

Herbage sweet
(Sugar-cane,
Grass, &c.)

Stems

solid.

Carpels single

265. Cyperaceae

Un-

Diaphoretic.

important.

Flowers naked.

Carpels

266. Desvauxiacese.

several.

Flowers with a calyx. Seeds

267. Restiacese.

few.

Flowers with a
Seeds numerous.

corolla.

268. Xyridaceae.

United to Spadicosae

The

by- Cyperaceae, and to Hypogynosae by Restiaceae.


Endogens with other parts of the system seems to be,
With Gymnospermas through Palmaceae
to
Cycadacese.
With Exogens
Alismaceae
Ranunculaceae.

relation of

With Rhizanthae
With Acrogens

Hydrocharaceae
Retosae
Araceae
through Araceae

Placentae

several,

parietal.
Sepals four.

Aristolochiaceae.

Cynomoriaceae.

Pistiaceae ?

CLASS
Sepals

Nymphaeaceae.
Menispermaceas.

Placentae pa-

IV.

Marsileaceae ?

RHIZANTH^E.

269. Rafflesiaceae

Astringent.

270. Cytinaceae

Astringent.

271. Cynomoriaceae

Astringent (F^m-

rietal.

Placentas central

gus melitensis).
These singular fungoid plants are neither Exogens nor Endogens, because they have no
vascular system, and their sexual apparatus is imperfect ; they are not Acrogens, because
they have flowers and sexes. They are connected
With Endogens through
With Acrogens

CLASS
Filicales.

Alliance 1.

V.

Araceae.

Fungaceae.

ACROGEN^E.

Stems

fistular,

tive organs borne

vascular.

upon the

272. Polypodiaceae
Ring of the thecae vertical
Gleicheniaceae
Ring of the thecae transverse 273.
Ring wanting. Thecae one- 274. Osmundaceae
.

celled, ribbed.

Ring wanting.
if

Reproduc-

leaves.
v

Astringent.

275. Danaeaceae

Thecae as

many-celled.
Thecae one-

Ring wanting.

ral.

Pecto-

Some eatable.

276. Ophioglossaceae

celled, veinless.

Thecae in cones

277. Equisetaceae

Epidermis

siliceous.

ALLIANCES OF PLANTS.

224
Alliance 2.

Lycopodales.
tive organs

Thecae naked
Thecae enclosed in involucres of the same form.
Thecae enclosed in. involu.

Stems

solid, vascular.

Reproduc-

growing on the stem.

278. Lycopodiacese
279. Marsileaceae

Emetic.

280. Salviniaceae

None.

None.

cres of two different forms.

Alliance 3.

M&scales.

Ger-

Without a vascular system.

minating processes uniting into a heterogeneous


body.
Thecae valveless, with

an

operculum.
Thecae opening into valves,
with an operculum.
Thecae opening into valves,
without an operculum.
Thecae valveless, without

Sporules in distinct thecse.


281. Bryaceae

Slightly astringent.

282. Andraeaceae.
283. Jungermanniaceae.
284. Hepaticaceae.

an operculum.
Alliance 4.

Charales.

Without a vascular system.

Germi-

nating processes uniting into a heterogeneous

body.

Reproductive organs axillary globules.


285. Characeae

Alliance 5.

Fungales.

Fo3tid.

Without a vascular system.

Ger-

minating processes either wholly distinct or


confluent in a homogeneous
body.

Born from a matrix which


veils

286. Fungaceae

them when young.

Stimulant ; nutritive.
Often poisonous (Ergot,

Mushroom,
Born

without

Living in

a matrix.

air.

Cellular,

rarely filamentous,

reproductive

bursting

Truffle}.

287. Lichenaceae

Orchal) ; nu(Iceland

tritive

with

Moss).

nucleus

through

Dye

their

surface.

Born without a matrix.


Living in water.

mentous
either

glued

Fila-

the filaments

solitary or several

together,

sporidia

having

and viviparous.

288. Algaceae

Nutritive.

225

ALLIANCES OF PLANTS.
This group touches Rhizanthese

Gymnospermae

through Fungaceae.
Lycopodales.
Filicales.

Characeae.

If the affinities that have thus been explained are correctly stated, a mutual connection
of the five great classes in the vegetable kingdom may be expressed by a circle, in the
middle of whose circumference stand Exogens and Endogens, side by side ; the common
point of all the classes is formed by Acrogens ; which are connected on the one hand with
Exogens by Gymnosperms, and on the other with Endogens by Rhizanths.
The following scheme will place this idea in a more distinct point of view
:

Aracets to Cynomoriaoea.

C'/Jvrant/taceee to Gnetacete.

226

VI.

SKETCH OF A NEW DISTRIBUTION OF THE


VEGETABLE KINGDOM.

SOME remarks have already been made upon what apand, howpear to be the true principles of classification (635)
1.

ever imperfect the attempt may be, this seems a proper place
to sketch out the way in which it may possibly be executed.

In Exogens there are two totally different modes in


2.
which the influence of the pollen is communicated to the seed.

The

larger part of this primary group consists of plants provided with the apparatus called style and stigma, through
which the pollen-tubes are introduced into the ovary in the

act

of fertilization.

But others are

so

constructed that the

upon the ovules, without the introduction of any intermediate apparatus ; a


peculiarity analogous
to what occurs among reptiles in the Animal
Kingdom and,
as was to have been anticipated, the
in
which this sinplants
gular habit occurs prove, upon being collected together, to
pollen falls immediately

form a group having no direct


which
affinity with those among
they had been previously associated. Hence Exogens have
been broken up into 1.
Exogens proper, or those having an
ovary, style, and stigma

and

2.
Gymnogens, which have neither.
Endogens, in like manner, two modes of propagation have been discovered, essentially different from each
other.
In the major part of them the result of the fertiliza3.

Among

tion of their seed is the


production of an embryo, having one
point upon its surface predestined to become a stem, and an-

other to become a root ; besides which their


elementary organization includes vascular tissue in abundance.
But
others,

although in a high state of developement, are wholly or nearly


destitute of vascular tissue; and when their
seed is
fertilized,

instead

of an
embryo being formed,

the issue

is

a mass of

NEW

DISTRIBUTION OF THE VEGETABLE KINGDOM.

227

sporules, or reproductive bodies, analogous to those which Aerogens have instead of seeds. The old class of Endogens required therefore to be replaced by 3. Endogens proper, whose

organs of propagation are seeds, and 4. Sporogens, commonly


called Rhizanths, whose reproductive bodies are
spores.
4.
Among Acrogens also two modes of growth occur, so
essentially different from each other that they evidently represent different kinds of vegetation.
In some of them there is
a distinct axis of growth, or stem and root,

symmetrically
they are irregular cellular expansions, destitute of true leaves ; in the former we find a
trace of something equivalent to the sexes of
Exogens and
Endogens, in the latter all indications of the kind disappear.
Thus are formed two groups, which may be called 5. Cormogens,
where there is a stem and leaves, and 6. Thallogens, where
clothed with leaves

in others

no separation of those parts.


extent dismemberments of the three classes of
Jussieu may be further carried, there is no evidence to show

there
5.

is

To what

however, probable that they are capable of much


further increase
for, with a few exceptions, the affinities of
it

is

not,

the six primary groups now indicated are too continuous and
complete to allow us to suppose that any great physiological
or fundamental differences of organization exist among them.

Some
6.

exceptions, however, do exist.

Among

Angiospermous

AristolocMacea,
Piperaceae,

Exogens

the

Nepenthacea, Lardizabalacea,

and some others

Natural

orders

Menispermacea,

allied to the latter, stand isolated,

were, in whatever part of the group they are stationed,


having no obvious affinity with any other orders; for we can
only regard the approximation of Menispermaceae to Anonacea,

as

it

Now
&c. as the result of altogether artificial considerations.
these orders appear to agree in one remarkable circumstance.
Instead of their wood being formed by zone deposited over
zone, season after season, as

is

the case in the great mass of

Exogens, they never have more than one zone of woody matWhether their
ter, to whatever age they may have arrived.
wood itself is formed exactly in the same way as that of other

Exogens, namely, by a gradual external addition of stratum


upon stratum, is doubtful ; it is probable that they have a

mode of growth

of their own, analogous to

that

of Aristolo-

NEW

228

DISTRIBUTION OF

which the wood when young is augmented by the sucbetween


cessive introduction of wedge upon wedge of wood
a
around
medullary
wedges originally placed concentrically
Such plants as these agree with Exogens in their Diaxis.
chia, in

in general appearance, but their


cotyledonous embryo, and
of growth is an approach to that of some Endogens to

mode

be presently noticed, and it therefore appears they ought to be


which from the homogeregarded as a fundamental group,
of
called
be
wood
of
the
Homogens, for the sake
may
neity
their structure with the concentrically zoned growth
contrasting
of other Exogens, to which the collective

might be applied.
three classes,

1.

name of Cyclogens
In this manner Exogens are composed of

Exogens proper,

2.

Gymnogens, and

3.

Homo-

Among Endogens we

find a group of exactly the same


and differing from the mass of the order in
The peculiar habit of Smilax and
nearly the same manner.
some other Endogens, which no one would suppose from their
7.

nature as the

last,

general appearance to belong to that class, led me formerly to


propose the separation of them into a group which was called

the Retose.
But as no better character could be found for it
than the reticulated leaves, nobody adopted it, and it has been
regarded as an unnecessary separation of plants essentially the

same ; an opinion to which, in the absence of evidence, there


has been nothing to oppose beyond the conviction that the
Retose group is in nature well founded, although its true characters may have been undiscovered.
It now, however, appears that Smilax and its allies have the wood of their axis
arranged upon a plan wholly or in part similar to that of Homoand consequently they will constitute, not a subdivision of
;

gens

Endogens as was formerly supposed, but a new class or priIf the annual branches of a Smilax are exa-

mary group.

mined, there is nothing in their internal structure at variance


with that of a stem of Asparagus ;
they are exactly Endogenous ; but in the rhizoma of the whole
genus (take the Sarsaparilla of the shops for instance) the wood is disposed in a
compact circle, below a cortical integument, and surrounding
a true pith
so that the rhizoma or
of the stem

;
permanent part
that of a
Homogen. In Dioscorea alata the stem is formed
of eight fibrovascular
with their backs
wedges placed in

is

pairs,

THE VEGETABLE KINGDOM.

229

the bark, surrounding a central pith and having


wide medullary plates between them in fact, when the stems
of this plant are in a state of decay, the eight fibrovascular

touching

wedges may be pulled asunder, like those of a Menispermaceous plant.


In Testudinaria elephantipes the structure of the
stem is of nearly the same kind ; several bundles of fibrovascular tissue form a circle surrounding a pith, and pierced
with broad medullary processes. Lapageria and Philesia have
each a zone of wood below their bark, and a central pith

which the common fibrovascular bundles of Endogens are


disposed ; a tendency to which is also observable in Smilax.
8.
It seems therefore clear that what I have elsewhere

in

composed of plants whose mode of


from that of Endogens in general
growth
and that the species composing it stand in the same relation to
the mass of Endogens, as Homogens to the mass of Exogens.
For these reasons it appears that Endogens contain three distinct types of organization, namely, Spermogens and Sporogens,
or Rhizanths, the former of which consists 1 of true Endogens
with striated inarticulated leaves, and 2. of false Endogens with
called the Retose
is

group

is

essentially different

reticulated disarticulating leaves, or Dictyogens.


From these considerations we learn that of the
9.

three

primary divisions of the Vegetable Kingdom, recognized by


Jussieu, two require to be broken up into three each, and the
other into two ; making eight in all ; the mutual relations
of which with each other and the Animal
expressed thus

Kingdom may be

Exogens.

Homogens.
Gymnogens.

Dictyogens.

Endogens.
Sporogens.

Cormogens.
Thallogens.

(Animal Acrita Kingdom.)

The

following analytical arrangement will


more plainly into view.

tinctions

bring

these

dis-

DISTRIBUTION OF

NEW

230

STATE

I.

SEXUAL OR FLOWERING PLANTS.

Division 1. Exogens.

Cyclogens.

j Class

I.

1 Class

II.

Class

III.

( Class

Division 2. Endogens.

Class

Class

<^

Exogens.

Gymnogens.
Homogens.

IV. Dictyogens.
V. Endogens.
VI. Sporogens
{Rhizanths).

STATE

II.

ESEXUAL OR FLOWERLESS PLANTS.


(

The

following

is

Class

VII. Cormogens.

\ Class VIII. Thallogens.


view
of
the orders that have to be
tabular
a

limited.
arranged in the classes thus
class divides into two series
each
that
It is assumed

the
the

one having albumen as a necessary part of the

structure,

other either wholly or almost destitute of that

substance (see

652).

have also adopted the principles before spoken of as ap-

pearing well suited to the construction of subordinate


(see 655) ; and, with reference to this, the following
are employed in the sense now assigned to them.
1

Consolidated

when

the floral envelopes are united

series

terms

both

with each other and the stamens, and with the ovary.
2. Separated; when the floral envelopes and stamens are
united with each other, but the ovary is consolidated and free.
3. Adherent ; when the petals and sepals adhere to each other
and the stamens and ovary, but have their parts disunited.
4. Disunited ; when the sepals and
petals adhere to each
other and the stamens but have their parts disunited, and do
;

not adhere to the consolidated ovary.


5. Dissolved; when the
sepals and petals are distinct from
the stamens, and also from the
ovary, whose carpels are disunited either wholly or by the styles.

These five gradations seem to


comprehend all the material
degrees of union, from complete consolidation, as in Composite,
to complete separation, as in Rammculacese.

THE VEGETABLE KINGDOM.


CLASS

SERIES

I.

I.

EXOGENS.

CONSOLIDATED.

Exalbuminous.
1.

Albuminous.

Asteraceae.

1.

Valerianaceae.

Calyceraceae.

Dipsaceae.

2.

Campanulaceae.
Lobeliaceae.
Stylidiaceae.

Goodeniaceae.
Scaevoleae.

3.

Cinchonaceae.
Galiaceae.
Caprifoliaceae.

Columelliaceae.

SERIES

II.

SEPARATED.
Albuminous.

Exalbuminous.
Brunoniacese.

1.

Globulariacese.

Convolvulacese.

Plantaginaceae.

Nolanaceae.

Plumbaginaceaj.

Salvadoraceae.
Boraginaceae.

Lamiaceaa.

Verbenaceae.
Lentibulaceae.

Cyrtandraceae.

2.

Hydrophyllaceae.

Bignoniaceae.

Primnlaceae.

Pedaliacese.

Acanthacese.

Myrsinaceae.
Ebenaceae.

Myoporacese.
Jasminaceae.

Papayaceae.

Sapotaceae.

231

NEW

232

DISTRIBUTION OF
3. Cestraceae.

3. Ehretiaceffi.

Solanacese.

Cordiacea;.

Scrophulariaceae.

Gesneraceae.
Stilbaceae.

Selaginaceae.

Oleacese.

4. Retziaceae.

Loganiaceae.

Apocynaceae.
Aselepiadaceae.
Spigeliaceae.

Gentianaceaa.

Orobanchaceae.

5.

Polemoniaceee.
Diapensiaceae.

Hydroleacese.

SERIES

III.

ADHERENT.
Albuminous.

Excdbuminous.
1.

Combretaceae.
Corylaceas.

1.

Aquifoliaceae.

Cornaceae.

Garryaceaa.
Araliaceae.

Apiaceae.
Alangiaceee.

Hamamelacea;.
Helvingiaceae.

2. Chailletiaceae.

Penaaaceae.

Lauracese.

Hernandiaceae.
Proteaceaa.

Thymelaceae.
Elseagnaceas.

2. Santalaceaa.

Loranthacese.

THE VEGETABLE KINGDOM.


3.

Rosaceae.

3.

233

Grossulaceae.

Calycanthaceae.

Homaliaceae.

Chrysobalanaceae.
Fabacese.

Malesherbiacese.

Connaraceae.

Loasacese.

Turneraceae.

Amyridaceae.
Anacardiaceae.

4.

Onagraceae.

4.

Ficoidaceae.

Scleranthaceee.

Lythraceae.
Melastomaceae.

Nyctaginaceaa.

Begoniaceae.

Memecylaceae.
Lecythidaceae.

Myrtaceae.

5.

Cactaceae.

5.

Rhamnaceae.

Passifloraceae.

Celastraceee.

Cucurbitaceae.

Bruniaceae.

Datiscaceae.

6.

Philadelphaceae.

Hydrangeaceaa.
Saxifragaceae.

Cunoniaceae.
Baueraceae.
Escalloniaceae.

Vaccinaceae.

SERIES IV. DISUNITED.

Albuminous.

Exalbuminous.
\.

Resedaceae.

1.

Ericaceae.

Capparidacese.

Epacridaceae.

Brassicaceae.

Empetraceae.

Moringaceae.

2.

Spondiaceae.
Brexiaceae.

Aurantiaceae.

2,

Pittosporaceae.

Fouquieraceas
Vitaceae.

NEW

234

DISTRIBUTION OF
Humiriaceae.

Meliacese.

Tremandraceae.

Cedrelaceee.

Burseraceae.

3.

3.

Clusiaceae.

Tiliaceae.

Marcgraaviaceee.

Elseocarpaceae.

Rhizobolacese.

Trigoniaceae.

Chlenacese.

Dipteracese.

Ternstromiaceae.

4.

Vochyaceae.
Krameriaceae.

4.

Papaveracese.

Nymphaeaceae

Sarracenniacese.

Sapindacese.
5.

Flacourtiaceee.

Bixacese.
Olacaceee.

Lacistemacese.

Samydacese.
Violacese.
Cistaceee.

SERIES V. DISSOLVED.

Albuminous.

Exalbuminous.
Rutacese.

}.

1.

Portulacaceae.

Staphyleaceae.
Aceraceae.

Caryophyllaceae.

Malpighiaceae.

Amarantaceae.

Petiveriaceae.

Chenopodiaceae.

Coriariaceae.

Phytolaccaceae.

2. Myricaceae.

Frankeniaceae.

Zygophyllaceae.
Simarubacese.

Droseraceae.

Illecebraceae.

2. Urticaceae.

Platanaceee.

Stilaginaceae.

Casuaraceae.

Monimiaceae.

Betulaceae

Ulmaceae.
Salicaceae.

Tamaricaceae.

Atherospermaceae
Myristicaceae.

THE VEGETABLE KINGDOM.


3.

Malvaceae.

Anonaceae.

3.

Nitrariaceae.

Schizandreaa.

Reaumuriacese.

Berberaceaa.

Hypericaceae.

Magnoliaceaa (Wintereee).

Elatinaceaa.

Dilleniacese.

Ranunculacese.
Podophylleae.

Hydropeltideaa.
Cephalotaceae.
Crassulaceee.

4.

235

Balsaminaceee.

4.

Geraniaceae.

Ledocarpeae.
Vivianiaceaa.

Surianaceae.

Oxalidaceaa.

Nelumbiaceae.

Linaceas.

5. Stackhousiaceaa.

Erythroxylaceaa.

Hugoniaceae.
Sterculiaceae.

Euphorbiaceae.
Scepaceae.

Putrangiveae.

CLASS

II.

GYMNOGENS.

Cupressaceae.
Pinaceae.

Taxaceae.
Gnetaceae.

Cycadaceae.

CLASS

III.

SERIES

Exalbuminous.

I.

HOMOGENS.

ADHERENT.
Albuminous.
Aristolochiacese.

NEW

236

DISTRIBUTION OF

SERIES

II.

DISUNITED.

Albuminous.

Exalbuminous.

Nepenthacese.

SERIES

III.

DISSOLVED.

Albuminous.

Exalbuminous.
1.

1.

Podostemaceae.

Callitrichaceee.

Saururacese.

Ceratophyllacese.

Chloranthaceee.
Piperaceae.

2.

Lardizabalaceae.

Menispermacese.

CLASS

IV.

DICTYOGENS.

Dioscoreaceae.

Smilacese.

Roxburgh] aceee.

CLASS

SERIES

V.

I.

ENDOGENS.

CONSOLIDATED.
Albuminous.

Exalbuminous.
1.

Apostasiacese.

1.

Musaceae.
Marantaceae.

Orchidaceae.

Hydrocharaceae.

Zingiberaceae.
2.

Iridacesa.

Bromeliaceae.

Burmanniaceae.
Vellozieaa.

Ha!modoracea3.
Amaryllidacese.
Taccaceae.

THE VEGETABLE KINGDOM.


SERIES

II.

237

DISUNITED.

Exalbuminous.

Albuminous.
1.

Aspidistreae.
Liliaceae.

Pontederaceae.
Xiphidiaceae.
Gilliesiaceae.

Philydraceae.

Juncaceae.
2. Acoraceee.

Araceae.
3.

Commelynaceae.
Xyridaceae.
Eriocaulacese.

Desvauxiacea3.

SERIES

III.

DISSOLVED.

Albuminous.

Excdbuminous.
Naiadaceae.

1.

Melanthaceae.

Juncaginaceaa.

Flagellariaceaa

Alismaceae.

Palmae.

Butomaceaa.

2.

Pandanaceae.
Cyclanthaceaa.

Typhacese.
Pistiaceae.
3. Restiaceae.

Cyperaceae.
Graminacea?.

CLASS

VI.

SPOROGENS.

(RHEANTHS.)

Rafflesiaceae.

Cytinaceae.

Balanophoraceae.

CLASS

VII.

CORMOGENS.

Filicales.

Lycopodiaceae.
Isoetaceae.

238

NEW

DISTRIBUTION OF THE VEGETABLE KINGDOM.


Salviniaceae.

Marsileaceae.

Equisetaceae.

Characese.

Bryaceae.

Andraeaceae.

Jungermanniaceae.
Marchantiaceae.

CLASS

VIII.

TIIALLOGENS.

Lichen aceae.
Algaceae.

Fungaceae.

239

MEDICAL BOTANY.

III.

THE

is

following

which are known

list

of the principal medicinal plants

Europe. The numbers


Flora Medica, (London, 1838, Longman
and Co.) an 8vo. of 656 pages, in which full descriptions of
It will be useful for London
the plants will be met with.
in a living state in

refer to the Author's

students to
Chelsea,

know

marked

that the plants in the Apothecaries Garden,


with red figures on a black ground, are num-

bered to correspond with this

list.

RANUNCULACE.E.
CLEMATIS.
1

Erecta.

Acrid.

Used

in cachectic diseases.

Powdered leaves

escharotic.
2.

Flammula.

3.

Vitalba.

Leaves used as

vesicatories.

Fruit and leaves acrid and vesicant.

Leaves employed

as rubefacients in rheumatism.

ANEMONE.
Powder

6. Pulsatilla.

and vomiting.
9. Hortensis.
1

0. Coronaria.

11. Nemorosa.
12. Hepatica.

of the root causes itching of the eyes, colic,

Extract used in

ttcnia.

Acrid in a high degree.


Acrid in a high degree.

Acrid in a

less degree.

Bland.

HYDBASTIS.
14. Canadensis.

Rhizoma

narcotic, bitter,

tonic.

dye.

KNOWLTONIA.
15. Vesicatoria.

Acrid.

Leaves used as vesicants.

ADONIS.
1

G.

Vernalis.

Emmenagogue.

Gives a yellow

MEDICAL BOTANY.

240
RANUNCULUS.
1 7.

Bulbosus.

Very

Root

18. Thora.

acrid, causing blisters

very

acrid.

The

and inflammation.

juice

used

for

envenoming

weapons.
Acrid.

19. Sceleratus.

20. Acris.

acrid.

Very

Leaves used by beggars to produce ulcers.


By carrying it in the hand it has produced

inflammation.
22. Flammula.

Vesicant.

Distilled

water an excellent emetic.

HELLEBORUS.
23. Niger.

Narcotic acrid.

The

Drastic.

fibres of the

an emmenagogue and hydragogue.

as

employed
Black Hellebore.

Narcotic acrid.

25. Viridis.

Narcotic

26. Foetidus.

commended

rhizoma are
Produces

Drastic.

Leaves emetic and purgative.

acrid.

as a vermifuge against the

Re-

worm, Ascaris lumbri-

coides.

COPTIS.

rhizomata afford a tonic

Its

27. Trifolia.

bitter,

without astrin-

gency.

NIGELLA.
28. Sativa.

Seeds aromatic, sub-acrid

formerly employed instead of

pepper, and also as carminatives.

DELPHINIUM.
29. Consolida.

Seeds emetic, the leaves and stalks compose

Acrid.

cosmetics, which are injurious to the skin.

30. Staphisagria.

Seeds extremely poisonous, emetic, drastic, and

inflammatory

useful in scabies

for pediculi in the

and fungous ulcerations ;

also

head.

ACONITUM.

The

31. Anthora.

32.

33. Paniculatum

The

diuretic.

34s Napellus

root highly acrid.

Root highly acrid used to


destroy wild beasts.
Leaves bitter, acrid, narcotic, diaphoretic, and

Lycoctonum

roots are

Narcotico-acrid

proved fatal to

human

more dangerous.
;

life.

a spirituous infusion of the root has


Leaves sudorific and diuretic.

A.CTJEA.

36. Spicata

Fruit

astringent

Roots antispasmodic,
poisonous.
expectorant,
reported to have afforded relief in cases of catarrh.

MEDICAL BOTANY.

241

XANTHORHIZA.

Wood

37. Apiifolia.

and bark a pure

tonic, intensely bitter, said to

be superior to Calumba.

P/EONIA.
38. Officinalis.

Seeds emetic and cathartic.

39. Corallina.

Root antispasmOdic.

Seeds emetic and cathartic.

PODOPHYLLE^E.
PODOPHYLLUM.
40. Peltatum.

Narcotico-acrid.

The rhizoma

is

an active

cathartic.

PAPAVERACE^E.
PAPAVER.
4

1 .

Rhoeas.

Syrupus Rhoeados

Slightly narcotic.

is

prepared

froTii

the petals.
42. Somniferum.

Narcotic (Opium).

ARGEMONE.
Seeds narcotic, anodyne, and purgative.

43. Mexicana.
is

employed

and

in chronic ophthalmia

The

juice

siphilitic sores.

SANGUINARIA.
44. Canadensis.

Rhizoma

Narcotico-acrid, tonic.

emetic, escharotic,

used in cases of polypi.

CHELIDONIUM.

rific

Stimulating, aperient, diuretic, and sudo-

Juice acrid.

45. Majus.

also a deobstruent.

FUMARIEJS.
FUMARIA.
46. Officinalis.

Herbage

bitter, diaphoretic

and

aperient.

CORYDALIS.
47. Tuberosa.
48. Fabacea.

Root

bitter

and

Root bitter and

acrid.

acrid.

NYMPH^EACE^E.
NUPHAR.
49. Lutea.

Rhizoma

50.

NYMPH^EA.
Rhizoma
Alba.

Odorata.

sedative and anti-aphrodisiac.

astringent, styptic,

and sub-narcotic.

Stems very astringent, used

in poultices.

MEDICAL BOTANY.

242

MYRISTICACE^.
MYRISTICA.
Seeds aromatic, act as narcotics in over doses.

53. Officinalis.

MAGNOLIACE^E.
MAGNOLIA.
Bark
54. Glauca.

bitter

ful in chronic

and aromatic, resembling cinchona.

Use-

rheumatism.

LlRIODENDRON.

Bark

55. Tulipifera.
as a

warm

bitter, aromatic, tonic,

sudorific in chronic

and

diaphoretic.

Used

rheumatism.

WINTERACE^E.
ILLICIUM.
56. Floridanum.

Bark and leaves aromatic and

APIACE^

OR

spicy.

UMBELLIFERE^.

ASTRANTIA.
Roots acrid and purgative.

67. Major.

ERYNGIUM.
The

68. Campestre.

root

is

sweet, aromatic, and tonic

diuretic

a reputed aphrodisiac.
Root sweet, aromatic, and tonic, but of
69. Maritirnum.

also

inferior

quality.

CICUTA.

The roots are a very dangerous poison.


Roots a dangerous poison, causing true tetanic con-

70. Maculata.

71. Virosa.

vulsions.

APIUM.
72. Graveolens.

Acrid.

Rendered bland by culture.

PETROSELINUM.
73. Sativum.

The

leaves are a
stimulating diuretic.

SlSON.
78.

Amomum

Fruit pungent and aromatic.

CARUM.
79. Carui.

Fruit carminative.

PlMPINELLA.
81. Saxifraga.

Root astringent, used to relieve tooth-ache.

MEDICAL BOTANY.

24-3

PlMPINELLA.
Root astringent.

82. Dissecta.

Used

as a

masticatory to relieve

tooth-ache.

83.

Used as a masticatory

Root astringent.

Magna.

to

relieve

tooth-ache.

84. Anisum.

Effects stimulant

and carminative.

Produces Anise.

(ENANTHE.

85. Crocata.

It

very dangerous poison.

has been considered the

most energetic of the narcotico-acrid Apiacese.

86. Phellandrium.

87.

dangerous poison, but rather less energetic.

Leaves poisonous

Cynapium.

also of

a nauseous smell.

FOENICULUM.
Leaves fragrant and stimulant.

Fruit carminative.

88. Vulgare.

Produces Fennel.
89. Dulce

Fruit carminative.

Leaves fragrant and stimulant. Pro-

duces Sweet-Fennel.

ATHAMANTA.
Fruit aromatic.

91. Cretensis.

MEUM.
92.

Roots aromatic and sweet, forming an ingre-

Athamanticum.

dient in Venice treacle.

Roots aromatic and sweet, forming an ingredient in

93. Mutellina.

Venice

treacle.

ANGELICA.
Root

94. Nemorosa.

acrid.

Employed

in cases of itch.

ARCHANGELICA.
95. Officinalis.

ed

Root fragrant,
disorders.

in pectoral

pungent. Stalks employLeaves, seeds, and roots aromatic

bitterish,

tonics.

OPOPANAX.
The

96. Chironum.

produces Opopanax, a

root

foetid

gum-resin,

similar to Asafcetida.

FERULA.
97. Asafcetida.

which are

foetid

is procured
by
and antispasmodic.

gum-resin

acrid, bitter,

slicing the roots,

Produces Asa-

fcetida.

93. Persica.

Like the

last.

R2

MEDICAL BOTANY.

244
FERULA.

Yields a gum-resinous secretion.


Yields a gum-resinous secretion.

101. Ferulago.

102. Tingitana.

103.

DOREMA.
Ammoniacum. The stem and
which is Ammoniacum.

PEUCEDANUM.
The juice

104. Officinale.

The

Montanum.
an acrid

of the root

is

fcetid

gum-resin,

antispasmodic and diuretic.

Leaves and stems are bitter and aromatic.

105. Oreoselinum.
106.

fruit yielding

Foetid, stimulant.

resin.

juice of the root bitter, foetid, hardening into


remedy in epilepsy.

IMPERATORIA.
107. Ostruthium

Root acrid and

bitter,

used as a masticatory in

tooth-ache.

ANETHUM.
109. Graveolens

Fruit carminative and stimulant.

Produces

Dill.

HERACLEUM.
110. Sphondylium.

Rind and root

acrid.

CUMINUM,
112.

Fruit

Cyminum.
Produces

carminative.

Used

in

veterinary surgery.

Cummin.

LASERPITIUM.
115. Glabrum.

and

116.

The

caustic.

juice of the root

is

gum-resinous, acrid, bitter,

Violent purgative.

DAUCUS.
Gummifer
Roots

bitter

and balsamic, yielding Bdellium

sicu-

lum.
117. Gingidium.
Roots bitter and balsamic.
1 1 8. Carota.
Fruit carminative and diuretic.

Root used as a cure

for ulcers.

ANTHRISCUS.
1

9. Sylvestris.

Narcotic.

120. Vulgaris.
Deleterious.
The whole plant highly poisonous.
121. Cfcrefolium.
Roots eatable.
Produces Chervil.

CONIUM.
124. Maculatum.

Powerfully narcotico-acrid.

Is

Hemlock.

SMYRNIUM.
125. Olusatrum.

Leaves slightly aromatic.

Fruit carminative.

MEDICAL BOTANY.

245

CORIANDRUM.
126. Sativum

Fruit carminative and aromatic.

ARALIACE^E.
PANAX.
127. Quinquefolium.
mulant.

Roots aromatic, pungent, restorative, and

sti-

ABALIA.
128. Nudicaulis.

Alterative and tonic.

Tincture of the wood used against

130. Spinosa.

colic.

HEDERA.
Leaves and berries

131. Helix.

bitter,

aperient,

emetic, sudorific.

Is the Ivy.

GROSSULACE^;.
RISES.
1

33.

Rubrum.

The

134. Nigrum.
juice

juice of the fruit refrigerant.

Fruit, leaves,
is

and wood

and stimulant.

tonic

The

used against catarrhs.

BERBERACE^E.
BERBERIS.

Bark

135. Vulgaris.

astringent.

drink

is

prepared from

the

fruit.

136. Lycium.

Extract useful in cases of ophthalmia.

VITACE^.
VlTIS.

Fruit cooling and antiseptic; diuretic and laxative

137. Vinifera.

in large quantities.

COMBRETACE.E.
TERMINALIA.
146. Benzoin.
147. Belerica.
tonic,

Juice concrete, used as incense.

The

kernels of the fruit intoxicating

and attenuant.

also astringent,

Produces Myrobalans.

MYRTACE^E.
MELALEUCA.
150. Cajeputi.

Essential

Cajeputi

oil.

oil

irritating

and stimulating.

Produces

MEDICAL BOTANY.
PUNICA.

Bark of the

152. Granatum.
ers

and bark of the

root
fruit

Flowa powerful anthelmintic.


Produces
tonic and astringent.

Pomegranates.

MYRTUS.
Aromatic and astringent.

153. Communis.

CARYOPHYLLUS.
Stimulant and carminative.

154. Aromaticus.

Produces Cloves.

EUGENIA.
155. Acris.

The unripe

fruit is

oily, irritable,

and

is

used to allay

is

used to allay

tooth-ache.

The unripe fruit is oily, irritable, and


the tooth-ache.
Is the Allspice of the shops.

156. Pimenta.

EUCALYPTUS.
Bark astringent, yielding a juice resembling Kino.'

158. Resinifera.

159. Robusta.

Bark

astringent.

CORNACE.E.
CORNUS.
63. Florida.

Bark

with an astringent aromatic taste ; tonic


antiseptic, giving a scarlet dye.
1 64. Sericea.
Said to be amongst the best of tonics.
Useful in in1

bitter,

and

termittent fevers.

165. Circinata.

166. Suecica

Astringent.

Useful in diarrhoea.

Berries tonic.

CUCURBITACE^E.
LAGENAHIA.
169. Vulgaris

Fruit poisonous.

CUCUMIS.
171. Colocynthis.
Fruit acrid.
duces Colocynth.

Poisonous to

human

beings.

Pro-

BRYONIA.
177. Alba.

Root acrid and


purgative.
removing bruises. Cathartic.
178. Dioica
Root acrid and

purgative.

removing bruises.

Cathartic.

Used as a
Used

as

discutient for

a discutient

for

MEDICAL BOTANY.

247

MOMOBDICA.
179. Elaterium.

Juice

It

poisonous.

a violent cathartic and

is

hydragogue.
180. Balsamina

Fruit a dangerous

poison,

acting

as a powerful

hydragogue.

BRASSICACE^E

OR

CRUCIFER^E.

COCHLEABIA.
189. Officinalis.

Antiscorbutic, stimulant,

when

but inert
1

dried.

and

diuretic, eaten fresh;

Produces Scurvy-grass.

Root stimulant, diaphoretic, and diuretic, and exProduces Horseradish.

90. Armoracia.

ternally rubefacient.

CARDAMINE.
191. Pratensis.

Stimulant,

flowers a

remedy

diaphoretic,

and

diuretic.

The

dried

for epilepsy.

SINAPSIS.

Seeds acrid, stimulating, and bitter.


Oil purgative,
rubefacient in paralysis.
Vesicant.
Produces Mustard.

192. Nigra.
194. Alba.

Seeds acrid and pungent.


Produces Mustard.

Used

as

stimulating

ca-

thartics.

ERUCA.
1

95. Sativa.

Seeds acrid and bitter.

RAPHANUS.
Seeds emetic.

196. Sativus.

Roots diuretic and laxative.

Produces

Radishes.

CAPPARIDACE^:.
CAPPARIS.
Flower-buds antiscorbutic, stimulant, and aperient.

197. Spinosa.

Produces Capers.
Fruit poisonous.
198. Pulcherrima.
Root acrid.
199. Cynophallophora.

An

infusion

recommended

in

dropsy.

VIOLACE^:.
VIOLA.
203. Odorata.
tive.

204. Canina.

205. Tricolor.
orders.

Petals used as a laxative.

Roots emetic and purga-

Flowers anodyne, producing faintness and apoplexy.


Leaves depurative. Roots emetic.
Bruised leaves used in the cure of cutaneous dis-

MEDICAL BOTANY.

248
IONIDIUM.

Roots

206. Ipecacuanha.

emetic.

Used

as

a .substitute for true

Ipecacuanha.

MORINGACE^E.
MOBINGA.
216. Aptera.

Seeds acrid.

Employed

in fevers

and as rubefacients.

PASSIFLORACE^.
PASSIFLORA.

Root emetic.
218. Quadrangularis.
Powerfully narcotic.
in hysteria.
220. Fcetida.
serviceable
Emmenagogue,

PAPAYACE^E.
CAEICA.

The milky

221. Papaya.

juice,

and powder of the

seeds, are

power-

ful vermifuges.

BIXACE.E.
BlXA.
224. Orellana.

Demulcent.

Produces Arnotto.

CANELLE^.
CANELLA.
231. Alba.

All parts of the tree are aromatic, hot, and


pungent,
fresh.
Distilled bark aromatic, carminative, and

when

stomachic

used in scurvy.

HYPERICACE^E.
HYPERICUM.
232. Perforatum

and

Leaves astringent.

An

infusion used in

lotions.

ANDROS^MUM.
233. Officinale.

Leaves esteemed as
vulnerary.

TERNSTROMIACE.E.
THEA.
237. Viridis.

238. Bohea.

stimulant narcotic.

Stimulant.

gargle

MEDICAL BOTANY.

249

SAPINDACE^E.
CABDIOSPERMUM.
Root aperient.

239. Halicacabum.

SAPINDUS.
Fruit detersive and acrid.

240. Saponaria.

employed

in chlorosis.

Tincture of the berries

Produces Soapberries.

^SCULUS.
Bark a

246. Hippocastanum.
in gangrene

and

its

febrifuge in fevers.

powder an

decoction used

errhine.

POLYGALACE^E.
POLYGALA.
Root acid and acrid; sudorific and expectorant in
247. Senega.
small doses, but emetic and cathartic in large.
Root acid and acrid; sudorific and expectorant
and cathartic in large.

254. Chamaebuxus.

in small doses, but emetic

LINAGES.
LlNUM.
Seeds used for cataplasms.
The infusion
Produces Linseed.

261. Usitatissimum.

is

demulcent and emollient.


262. Catharticum.

Bitter, cathartic,

and purgative.

CISTACE^:.
ClSTUS.

Resin stimulant and emmenagogue.


Produces Ladanum.

264. Creticus.

Recommended

in chronic catarrh.

265. Ladaniferus.

Resin

stimulant

and emmenagogue.

Used

in

chronic catarrh.

Resin stimulant and emmenagogue.

266. Ledon.

Used

in chronic

catarrh.

267. Laurifolius.

Resin

stimulant

and emmenagogue.

Uved

chronic catarrh.

STERCULIACE.E.
KYDIA.
274. Calycina.

Bark

sudorific.

THEOBROMA.
275. Cacao.

Seeds nutritive, restorative.

Produces Chocolate.

in

MEDICAL BOTANY.

250
ADANSONIA.
278. Digitata.

Mucilaginous.

Fruit

Dried leaves useful in fevers.

sub-acid.

MALVACEAE.
ABUTILON.

Used

281. Indicum.

as

an emollient.

MALVA.
Mucilaginous and emollient.

284. Sylvestris.

Is the

Mallow.

ALTHAEA.
285. Officinalis.

Mucilaginous

and

Is

emollient.

the

Marsh

Mallow.

ABELMOSCHUS.
Mucilaginous, emollient, and demulcent.
used to form poultices.

287. Esculentus.

Leaves

Seeds cordial and stomachic.

288. Moschatus.

TILIACE^.
TILIA.

Flowers antispasmodic.

293. Europeea

The Lime-tree.

LYTHRACE^.
HEIMIA.
295.

and

Sudorific

Salicifolia.

Used

diuretic.

venereal

in

dis-

orders.

LYTHRUM.
296. Salicaria.

Astringent.

Recommended

in cases of diarrhoea.

MELIACE^E.
MELIA.
297. Azedarach.

Root bitter and nauseous.

Used

as

mintic.

GUAREA.
301. Aubletii.

Bark emetic and purgative.

CEDRELACEvE.
SwiETENIA.
305. Mahagoni.

Bark

febrifugal.

Produces Mahogany.

an anthel-

MEDICAL BOTANY.

251

AURANTIACE^E.
CITRUS.
316. Aurantium.

Peel of the fruit tonic and aromatic.

Produces

Seville Oranges.

317. Bigaradia. Peel of the fruit bitter and tonic.


318. Limetta.
Fruit fragrant.
Produces Limes.
319. Limonum. Juice of the fruit yields citric acid.
aromatic and stomachic.

The

peel

Produces Lemons.

SPONDIACE^.
SPONDIAS.

320. Mangifera.

Emollient.

RHAMNACE^E.
ZlZIPHUS.
322. Jujuba.

Fruit pectoral.

Bark used

for

Produces

diarrhoea.

Jujubes.
Fruit pectoral.

323. Vulgaris.

Bark used

for diarrhoea.

BERCHEMIA.
Roots used in cachectic disorders

324. Volubilis.

said to be anti-

siphilitic.

CEANOTHUS.
Astringent and antisiphilitic.

325. Americanus.

RHAMNUS.
326. Catharticus.

Fruit

purgative

produces

colic.

An

hydra-

The Buckthorn.

gogue.

Fruit emetic.
327. Frangula.
Fruit emetic.
328. Infectorius.

329. Saxatilis.

Fruit emetic.

331. Oleoides.

Fruit emetic.

332. Buxifolius.

Fruit emetic.

EUPHORBIACE^E.
Buxus.
350. Sempervirens.

Leaves and wood bitter and nauseous

and purgative.

sudorific

Produces Box -wood.

CICCA.

351. Disticha.

Leaves

sudorific.

Seeds cathartic.

Fruit sub-acid.

CROZOPHORA.
359. Tinctoria.
ties.

Plants with emetic, drastic, and corrosive proper-

Seeds cathartic.

MEDICAL BOTANY.

252
CROTON.

Bark aromatic and

360. Cascarilla.

Bark

361. Eleuteria.

fragrant.

aromatic, tonic, stimulant.

bitter,

Produces

Cascarilla.

Seeds drastic.
363. Tiglium.
Bark of the root aromatic and purgative.
Aromaticum.
369.
RICINUS.
374. Communis.

Seeds cathartic.

Produces Castor-oil.

JATROPHA.
375. Curcas.

emetic and drastic.

Seeds

and

Leaves rubefacient

discutient.

377

a.

The

Multifida

seeds are excellent emetics

and purgatives.

JANIPHA.
378. Manihot.

Expressed juice poisonous.


duces Cassava and Tapioca.

Pro-

Fecula nutritive.

MERCURIALIS.
384. Perennis.
385. Annua.

Very

poisonous, producing vomiting

and

diarrhoea.

Poisonous.

HIPPOMANE.
Juice caustic and venomous.

389. Mancinella.

Acrid.

Manchineel.

HURA.
390. Crepitans.
Milk very venomous, producing blindness.
a drastic purgative.
An emetic.

Seeds

EUPHORBIA.
393. Tirucalli.

Milk a remedy for siphilis; cathartic and emetic.


Bark of the root purgative. Produces Euphor-

395. Antiquorum.
bium.

Milk purgative.

396. Canariensis.
397. Heptagona.
398. Officinarum.

400. Nereifolia.

The milk

Produces Euphorbium.

a mortal poison.
Milk purgative. Produces Euphorbium.

Juice

is

of the

leaves

purgative, deobstruent,

and

diuretic.

401. Gerardiana.
402. Lathyris.

Bark of the root cathartic and emetic.


Seeds drastic.

Bark of the

root

and stems

cathartic

and emetic.
403. Esula.
A dangerous poison.
404. Cyparissias.
virulent poison.
405. Thymifolia.
Violent

406. Ipecacuanha.

purgative.
Vulnerary, anthelmintic.
Root powerfully emetic and cathartic.

MEDICAL BOTANY.

253

EUPHORBIA.
407. Peplis
408. Peplus.

All the parts purgative.


All the parts purgative.

409. Falcata.

All the parts purgative.

410. Corollata.

Emetic,

expectorant,

and

cathartic.

The bruised

root excites inflammation.

411. Linearis

Juice employed for siphilitic ulcers.

PEDILANTHUS.
412. Tithymaloides.

Antivenereal, emetic.

CELASTRACE^E.
MAYTENUS.
Leaves stimulant.

415. Chilensis.

SILENACE^E.
SlLENE.

Root anthelmintic.

418. Virginica.

SAPONARIA.
420. Officinalis.

Saponaceous.

GYPSOPHILA.

42 1

Struthium.

Saponaceous.

TAMARICACE^E.
TAMARIX.
Bark
Manna.

422. Gallica.

bitter

and

astringent.

Branches yield a kind of

SIMARUBACEJG.
QUASSIA.
424. Amara.

Wood

bitter

and

tonic.

Infused flowers stomachic.

PlCRjENA.
427. Excelsa.

Wood

bitter, tonic,

and stomachic.

Produces Quassia

chips.

RUTACEJE.
RUTA.

Used as an

429. Graveolens.

emmenagogue, antispasmodic

and

anthelmintic.

BAROSMA.
436. Crenulata.

Leaves an excellent aromatic, stomachic, and


Produces Diosma leaves.

cious diuretic.

effica-

MEDICAL BOTANY.

254
BABOSMA.
'437.

Leaves an excellent aromatic, stomachic, and

Serratifolia.

cacious diuretic.

Leaves an excellent aromatic, stomachic, and

438. Crenata.

effi-

Produces Diosma leaves.


effica-

Produces Diosma leaves.

cious diuretic.

ZYGOPHYLLACE^E.
ZYGOPHYLLUM.
439. Fabago.

Esteemed as a vermifuge.

GUAIACUM.
440. Officinale.

Wood

yielding a bitter, acrid, stimulant gum-resin,

as a diaphoretic

employed

and

alterative.

XANTHOXYLACE.E.
PTELEA.
442. Trifoliata.
bitter.

Young

shoots anthelmintic.

Fruit aromatic and

substitute for hops.

XANTHOXYLON.
444. Fraxineum.

Bark aromatic and pungent.

Used

as a

remedy

in chronic rheumatism.

Infusion antispasmodic.

445. Clava Herculis.

Tincture febrifugal.

Decoction antisiphilitic.

BRUCEA.
450. Antidysenterica.

Tonic, astringent.

GERANIACE.E.
GERANIUM.
Root
451. Maculatum

astringent, containing Tannin.

A remedy in

452. Robertianum.

nephritic complaints.

OXALIDACEvE.
OxALIS.
453. Acetosella.

Plant refrigerant, antiscorbutic.

CORIARIACE.E.
CORIARIA.
454. Myrtifolia.

Fruit a dangerous poison.

ROSACES.
POTENTILLA.
455. Reptans.
Root very astringent.
456. Tormentilla. Root very astringent.

255

MEDICAL BOTANY.
GEUM.
457. Rivale.

Useful in diarrhoea.

Stomachic.

458. Urbanum.

Useful in diarrhoea.

Stomachic.

459. Canadense.

Root and leaves a mild

tonic.

Useful in

Bitter.

diarrhoea.

AGRIMONIA.
460. Eupatoria.

Astringent, anthelmintic.

RUBUS.
Bark of the

461. Villosus.

root

astringent.

Useful

in

cholera,

diarrhoea, &c.

ROSA.
462. Canina.

Laxative.

463. Centifolia.

Laxative.

464. Gallica.

Petals astringent

and

tonic.

GILLENIA.
465. Trifoliata.

Roots emetic.

SPIRAEA.

467. Ulmaria. Aromatic, tonic.


468. Filipendula.
Aromatic, tonic.

AMYGDALE^E.
AMYGDALUS.
Oil of the
470. Communis.
bitter

471. Persica.

Produces

seeds extremely poisonous.

almonds.
Oil, flowers,

and seeds extremely poisonous.

CERASUS.
472. Laurocerasus.

Leaves, bark, and seeds poisonous.

Produces

acid.

hydrocyanic
Leaves poisonous. Bark febrifugal.
473. Virginiana.
474. Padus. Abounds in the oil of bitter almonds, and

is

therefore

poisonous.

Bark

475. Capollim.

febrifugal.

PRUNUS.

The bark

477. Cocumilia.
478. Spinosa.

is

a remedy for the fevers of Calabria.

Fruit acid, astringent, and austere.

POMEJE.
PYRUS.
479. Aucuparia.

Leaves poisonous.

MEDICAL BOTANY.

2o6
CYDONIA.

Seeds demulcent.

480. Vulgaris.

SANGUISORBE^E.
ALCHEMILLA.
Decoction slightly tonic.

481. Vulgaris.

FABACE^E
TRIBE

OR
I.

LEGUMINOS^E.

PAPILIONACEJE.

ANAGYRIS.
Seeds poisonous.

482. Fcetida.

BAPTISIA.

Roots and herbage

483. Tinctoria.
thartic,

antiseptic,

sub-astringent, ca-

and emetic.

GENISTA.
484. Tinctoria

Produces a yellow dye.

Bitter.

CYTISUS.
485. Laburnum.

Seeds poisonous, narcotico-acrid.

486. Alpinus.
Seeds poisonous, narcotico-acrid.
487. Scoparius.
Decoction of the shoots diuretic
Seeds emetic.

and

cathartic.

Produces broom-tops.

ANTHYLLIS.
488. Hermannia.

Root

One

489. Vulneraria.

diuretic.

of the best styptics.

TRIGONELLA.
490. Foenum Graecum.

Decoction of the seeds an emollient.

Used

in veterinary medicine.

MELILOTUS.
491. Officinalis.

Decoction emollient.

Used

in lotions

and enemas.

TRIFOLIUM.
492. Alpinum.

Roots sweet and demulcent.

INDIGOFERA.

The dye is a dangerous vegetable poison.


The dye is a dangerous vegetable
Powdered
poison.

494. Tinctoria.
495. Anil

leaf used in hepatitis.

496. Argentea.

The dye

is

a dangerous vegetable poison.

CLITORIA.
498. Tematea.

Roots emetic.

MEDICAL BOTANY.

257

Roots sweet, tonic, demulcent.

Produces Liquorice.
Produces an

GLYCYRRHIZA.
500. Glabra.
501. Echinata.

Roots

sweet, tonic, demulcent.

less

inferior sort of Liquorice.

AGATI.
507. Grandiflora.

Bark

bitter

and

tonic.

PISCIDIA.

508. Erythrina.

Tincture

of the

Bark astringent and

bark narcotic and diaphoretic.

irritating.

COLUTEA.
Leaves purgative.

509. Arborescens.

ASTRAGALUS.
512. Tragacantha.

Produces a kind of Tragacanth.

Emollient.

CORONILLA.
Leaves cathartic.

514. Emerus.
515. Varia.

Leaves diuretic and cathartic.

Juice poisonous.

ARTHROLOBIUM.
Leaves vesicant.

516. Scorpioides.

ALHAGI.

From

518. Maurorum.

branches exudes

the

substance of the

nature of Manna.

ERVUM.
Seeds poisonous.

519. Ervilia.

LATHYRUS.
Seeds narcotic, producing head-ache

520. Aphaca.

if

eaten in a ripe

state.

521. Cicera.

Seeds narcotic.

ABRUS.
Root

522. Precatorius

and leaves employed as a substitute

for

Liquorice.

MUCUNA.
526. Pruriens.

Hairs

irritating.

TRIBE

II.

Produces Cowitch.

CjESALPINIEjE.

ANDIRA.
533. Inermis.
gative,

Bark anthelmintic, mucilaginous,

and narcotic

drastic, emetic, pur-

poisonous in large doses.


8

MEDICAL BOTANY.

258
CASSIA.

An

536. Acutifolia.

excellent

purgative.

Produces

Alexandrian

Senna.

An excellent purgative.

539. Obovata.

Produces Black-leaved Senna.

Leaves purgative.
Leaves purgative.
544. Marilandica.

540. Tora.

CATH ARTOCA RPUS.


545. Fistula.

Extract of the pulp laxative.

an excellent

Seeds purgative.

Roots

febrifuge.

POINCIANA.
Root acrid and poisonous.

548. Pulcherrima.

emmenagogue

Leaves a powerful

also purgative.

H^EMATOXYLON.
549. Campeachianum.

Decoction used in
powerful astringent.
Produces Logwood.

and dysentery.

diarrhoea

BAUHINIA.
55 1

Toraentosa.

Astringent.

TAMARINDUS.

The pulp of the

552. Indica.

sub-acid

fruit is cooling and laxative.


Leaves
employed as an anthelmintic. Produces Tamarinds.

HYMEN JEA.
553. Courbaril.

Fruit purgative.

TRIBE

III.

Bark anthelmintic.
MIMOSEjE.

ACACIA.
557. Catechu.

558. Vera.

Astringent.

The bark

566

a. Mollissima.

566

b.

yields

Gum

arable.

Astringent.

Melanoxylon.

Astringent.

VACHELLIA.
567. Famesiana.

Bark yields a gum

like

Gum

SAXIFRAGACE.E.
HEUCHERA.
572. Americana.

Root a powerful
astringent.

CRASSULACE^E.
SEMPERVIVUM.
573. Tectorum.

Leaves astringent

refrigerant.

arable.

MEDICAL BOTAXY.

259

SEDUM.
574. Telephium.

and

Refrigerant

Leaves

astringent.

useful

in

diarrhoea.

575. Acre.

Leaves

Recommended

acrid.

in

cancerous

cases

and

epilepsy.

ANACARDIACE^E.
MANGIFERA.
584. Indica.

Gum-resin

slightly bitter

and pungent.

ANACARDIUM.

Gum

586. Occidentale.

Juice acrid.

astringent.

nuts abound in a caustic thick

The

coats of the

Produces Cashew.

oil.

RHUS.
589. Toxicodendron.

Yields a narcotic, acrid, milky juice, extremely

poisonous.

590. Glabrum.

Yields

narcotic,

acrid,

milky

juice,

extremely

poisonous.

SCHINUS.
595. Molle.

Acrid.

PlSTACIA.

596. Vera.

Fruit emollient.

Produces Pistacia nuts.

597. Terebinthus.

Yields Cyprus turpentine.


598. Lentiscus.
Produces a sweet, fragrant, stimulant resin, called
Mastich, used to preserve the teeth.

CORYLACE^:.
QUERCUS.
Bark astringent
haemorrhage and diarrhoea.

599. Pedunculata.

600.

Bark

Sessiliflora.

the powder employed in passive

From

astringent.

this the oak-galls are ob-

tained.

Feeds the Kermes

602. Coccifera.

Bark and leaves

603. Falcata.

insect.

astringent.

Employed

in cases of

gangrene.

BETULACE^E.
BETULA.
604. Alba.

Bark

tonic.

Employed

as a febrifuge.

ALNUS.
605. Glutinosa.

Bark

tonic.

decoction employed as a gargle.


s 2

MEDICAL BOTANY.

260

URTICACE^:.
URTICA.
607. Dioica.

The whole plant

is

and

astringent

Is the

diuretic.

Nettle.

HUMULUS.
Ripe catkins

609. Lupulus.

narcotic

and

Infusion

bitter.

and

Produces Hops.

tincture aromatic, tonic.

Ficus.

Bark

611. Indica

tonic.

relieve tooth-ache.

Juice applied to the teeth and


Is the Banyan-tree.

gums

to

Yields Caoutchouc.

612. Elastica.

Seeds cooling and alterative.

616. Religiosa.

Fruit

617. Carica.

pectoral,

Produces

and laxative.

demulcent,

Figs.

CANNABIS.

an
very powerful, stimulating narcotic, used as
Produces
Hemp.
intoxicating drug.

618. Sativa

MOBUS.
619. Nigra.

Bark cathartic and anthel-

Fruit cooling and laxative.


Produces Mulberries.

mintic.

620. Alba.

Root said to be an excellent vermifuge.

DORSTENIA.
621. Contrayerva.

Root stimulant,

eruptive and other

Root stimulant,

622. Brasiliensis.

Root stimulant,

624. Drakena.

and tonic; used

sudorific,

in

diseases.
sudorific,

sudorific,

and

and

tonic.

tonic.

ULMACE^E.
ULMUS.
626. Effusa. The inner bark demulcent and
and a

tringent

627. Campestris.
astringent

diuretic

slightly as-

feeble tonic.

The inner bark demulcent and


and a

diuretic

slightly

feeble tonic.

MYRICACE^E,
MYRICA.
628. Gale.

Infusion used as a vermifuge

Hops
629. Cerifera.

leaves as a substitute for

in brewing.

Bark of the

root acrid

mulating and very acrid.

and

astringent.

Powder

sti-

MEDICAL BOTANY.

261

COMPTONIA.

Used

Tonic and astringent.

630. Asplenifolia.

in diarrhoea.

JUGLANDACE^;.
JUGLANS.
Inner bark of the root a mild and efficacious laxative;

631. Cinerea.

of the stem, rubefacient.

632. Regia.

The young

Produces Walnuts.

fruit purgative.

CHLORANTHACE^:.
CHLOBANTHUS.
633.

All the parts powerfully aromatic.

Officinalis.

Root an active

stimulant.

633

a.

Brachystachys.

All the parts

powerfully aromatic.

Roots

active stimulants.

PIPERACE^:.
PIPER.

Produces Round Pepper.


Produces Long Pepper.

Pungent and stimulant.


Pungent and stimulant.

634. Nigrum.
636. Longum.

Leaves and shoots discutient.

639. Amalago.

Root

sudorific, dia-

Fruit pungent.
phoretic.
642. Betel.
By chewing the leaf intoxicating effects are produced.
Stimulant.

SALICACE^E.
SALIX.
648. Russelliana.

Bark

febrifugal.

Bark slightly febrifugal.


649. Fragilis.
650. Purpurea.
Bark febrifugal.
651. Alba.

Bark

652. Pentandra.

653. Caprea.

febrifugal.

Bark aromatic and

Bark

febrifugal.

febrifugal.

POPULUS.
Leaf-buds bitter, aromatic.
654. Nigra.
655. Dilatata. Leaf-buds bitter, aromatic.
656. Balsamifera.
657. Candicans.
659. Tremuloides.

Buds
Buds

diuretic

diuretic

and

and

antiscorbutic.

antiscorbutic.

Bark esteemed as a

febrifuge.

MEDICAL BOTANY.

262

BALSAMACE^E.
LlQUIDAMBAE.
Bark pungent, bitter, expectorant. Produces Storax.
Almost inert.
662. Styraciflua.

661. Orientate.

THYMELACE.E.
DAPHNE.
All the parts excessively acrid, acting as an

666. Mezereum

irri-

tant poison.

All the parts excessively acrid, acting as an irritant

667. Laureola.
poison.

All the parts excessively acrid, acting as an irritant

668. Gnidium
poison.

DIRCA.

Bark

670. Palustris.

acrid, cathartic, vesicant.

Fruit narcotic.

HERNANDIACE^E.
HEENANDIA.
671. Sonora.

Juice of leaves a

Bark, seed, and leaves purgative.

powerful depilatory.

LAURACE^E.
ClNNAMOMUM.
Aromatic, stimulant.

674. Zeylanicum.

Produces Cinnamon.

CAMPHORA.
685. Omcinarum.

Yields Camphor.

PERSEA.
686. Gratissima.
astringent.

Leaves balsamic, pectoral, and vulnerary.


Yields the Avocado Pear.

Seeds

SASSAFRAS.
697. Officinale.
diuretic

Dried leaves mucilaginous.


Plant employed as a
sudorific.
Produces Sassafras.

and

BENZOIN.
699. Odoriferum.

Bark aromatic, stimulant, and

the twigs a vermifuge.

Fruit aromatic,

tonic.

oil

Infusion of

a stimulant.

LAURUS.
701. Nobilis.

Leaves and

fruit

aromatic.

Fixed

oil

a stimulant.

MEDICAL BOTANY.

263

ARISTOLOCHIACE^E.
ARISTOLOCHIA.

704

a.

The root has a disagreeable smell, and a strong


Cymbifera.
bitter aromatic taste.

706. Trilobata.

sudden and powerful sudorific.


root has a penetrating smell and bitter

The

708. Serpentaria.

acting as a stimulant, tonic, diaphoretic.

antispasmodic and anodyne.


709. Pallida.
slight aromatic stimulant tonic.

Sudorific

ployed as an emmenagogue in amenorrhoea.


712. Sempervirens.
slight aromatic stimulant tonic.

employed
A
713. Rotunda.

em-

Sudorific;

an emmenagogue in amenorrhoea.

as

slight aromatic stimulant tonic.

ployed as an
714. Clematitis.

taste,

In some cases an

emmenagogue

Sudorific;

em-

in amenorrhcea.

Roots powerfully stimulating.

ASARUM.
Roots purgative, emetic, and diuretic.
716. Europaeum,
leaves used to provoke sneezing.

Rhizoma

717. Canadense.-

aromatic.

A warm

Powdered

stimulant diapho-

retic.

CHENOPODIACE^E.
CHENOPODIUM.
719. Olidum.
Employed as an antispasmodic and emmenagogue.
721. Botrys.
Expectorant, employed in catarrh and humoral asthma.
The seeds yield an oil which is powerfully
722. Anthelminticum.
anthelmintic.

723. Ambrosioides.

Stimulant, corroborant.

ATBIPLEX.
724. Angustifolia. Seeds emetic.
Seeds emetic.
725. Hortensis.

SALSOLA.
726. Kali.

Yields Soda.

Yields Soda.

727. Sativa.
728. Soda.

Yields Soda.

729. Tragus.

Yields Soda.

PHYTOLACCACE^E.
PHYTOLACCA.
730. Decandra.

Root emetic.

Said to cure psora and tsenia capitis.

MEDICAL BOTANY.

264

POLYGONACE^E.
COCCOLOBA.
Leaves, wood, and bark are astringent; the decoc-

731. Uvifera.

Jamaica Kino.

tion forms

RHEUM.
732. Emodi.

Roots

tonic,

and

astringent,

purgative.

Furnishes

Indian Rhubarb.

Root

737. Rhaponticum.

bitter,

astringent,

Rhubarb

and aromatic

when

inferior.

chewed, mucilaginous.
Roots purgative and tonic.
738. Undulatum.
739. Caspicum. Roots purgative and tonic.

Roots purgative and tonic.


740. Compactum.
Roots purgative and tonic.
741. Palmatum.
Roots purgative and tonic.
742. Crassinervium.

All produce

Rhubarb

Nos. 741 and 735


the best.

RUMEX.
Root astringent
cure for the itch.

743. Crispus.

used in the form of ointment as a

Root astringent

744. Obtusifolius.

employed as a

Plant agreeably acid.


Produces Sorrel.
diuretic.

745. Acetosa.

746. Alpinus.

dentifrice.

Acting as a refrigerant and

Root purgative.

POLYGONUM.
Leaves so acrid as to act as vesicants.

747. Hydropiper.

ful diuretic.

748. Bistorta.

power-

Dyes wool yellow.

powerful astringent.

and leucorrhoea

Decoction employed in gleet

also in passive haemorrhages

and

diarrhoea.

Fruit emetic and cathartic.

749. Aviculare.
751. Amphibium.

Yields a false Sarsaparilla.

PETIVERIACE^].
PETIVERIA.
752. Alliacea.

All the parts acrid, sudorific,


emmenagogue.
remedy for tooth-ache.

roots used as a

NYCTAGINACE^E.
MlRABILIS.
754. Jalapa
Root purgative.
755. Longiflora.
Root exceedingly
purgative.

The

MEDICAL BOTANY.

265

PYROLACE^E.
CHIMAPHILA.
Leaves, stalks, and roots bitter-sweet, pungent.
Fresh leaves acrid, acting as vesicants and rube-

775. Corymbosa.
Diuretic.

Stomachic and

facients.

tonic.

ERICACEAE.
RHODODENDRON.

Maximum.

Acting as a poison.
Astringent, narcotic.
Reported to be deleterious.
Astringent, narcotic.
Leaves narcotic in a high degree ; useful in
778. Chrysanthum.
776.

777. Ponticum.

and venereal complaints.

chronic rheumatism

AZALEA.
779. Pontica.

Qualities of the plant deleterious.

LEDUM.

The

780. Latifolium.
delirium

leaves infused in beer produce head-ache

although they have been

in agues, dysentery,

781. Palustre.

and

and

used with advantage

diarrhoea.

Ditto.

KALMIA.
Leaves poisonous

782. Latifolia.

shoots poisonous to
to

them

man.

to

animals

narcotic.

Young

brown powder which adheres

acts as a sternutatory.

GAULTHEBIA.
783. Procumbens.

Fruit contains an aromatic, sweet, pungent, vo-

which

latile oil,

is

antispasmodic and diuretic.

tincture

useful in diarrhoea.

ARBUTUS.
784. Unedo.

wine

is

made from

the fruit, reported to be narcotic.

ARCTOSTAPHYLOS.
785.

Uva

ursi.

Leaves astringent and

and calculous

cases.

bitter.

Used

Diuretic.

LOISELEURIA.
786. Procumbens.

Useful as an astringent medicine.

in

nephritic

MEDICAL BOTANY.

266

VACCINACE^E.
VACCINIUM.

The

Fruit narcotic.

787. Uliginosum.

berries yield

an intoxicating

liquor.

PRIMULACE^E.
CYCLAMEN.
Root acrid

788. Hedersefolium.

acting as a drastic purgative,

em-

menagogue.

PRIMULA.
789. Veris.

Flowers sedative.

Produces Cowslips.

ANAGALLIS.
Prescribed in epilepsy and dropsy.

Acrid.

790. Arvensis.

SAPOTACE^;.
ACHKAS.
795. Sapota.

Bark a powerful astringent.

Seeds diuretic.

EBENACE^E.
DlOSPYRUS.

Bark a powerful astringent and

798. Virginiana.

febrifuge.

STYRACE^E.
STYRAX.

799. Officinale.

stimulating expectorant.

Produces Storax.

AQUIFOLIACE^].
ILEX.

801. Aquifolium.

Root and bark emollient, expectorant, and diu-

Leaves febrifugal.

retic.

PRINOS.
804-.

Bark a valuable

Verticillatus.

tonic.

Berries emetic,

tonic,

corroborant.

CONVOLVULACE^;.
IPOM^A.
807. Macrorhiza.
matter.

809. Purga.

Roots

consisting

Roots purgative.

Produces jalap.

BATATAS.
815. Paniculata.

of saccharine

Laxative.

Roots cathartic.

and farinaceous

MEDICAL BOTANY.

267

PHARBITIS.
816. Nil.

Said to be a quick cathartic.

Seeds purgative.

CONVOLVULUS.
817. Scammonia.

Roots cathartic.

818. Althseoides.

Roots purgative.

Produces Scammony.

CALYSTEGIA.
819. Sepium. Roots purgative.
820. Soldanella. Roots purgative.

LOBELIACE.E.
LOBELIA.
823. Inflata.

An

asthma.
824. Siphilitica.

acrid narcotic, and powerful emetic.


Used in
In small doses expectorant and diaphoretic.
Root acrid and emetic.
Used as a remedy for

siphilis.

HlPPOBROMA. (ISOTOMA.)
Acrid, venomous.

825. Longiflorum.

TUPA.
826. Feuillsei.

venomous.

Acrid,

The

of the

smell

flowers said

to produce vomiting.

CINCHONACE.E.
HYMENODICTYON.
Bark

856. Excelsum.

and

bitter

astringent.

EXOSTEMA.
Juice of the capsules produces a burning itching in
Bark febrifugal and emetic.
lips.

857. Caribaeum.

the nostrils and

858. Floribundum.

Bark

febrifugal

and emetic

rather drastic.

MANETTIA.
Bark of the

862. Cordifolia.

dysentery, acting as

root a valuable

remedy

in dropsy

and

an emetic.

RANDIA.
864. Dumetorum.

Fruit narcotic, emetic.

GARDENIA.
865. Campanulata.

Fruit employed as a cathartic and anthelrnintic.

COFFEA.
876. Arabica.

Stimulating, aromatic.

Produces Coffee.

MEDICAL BOTANY.

268

CAPRIFOLIACE^E.
TRIOSTEUM.

Bark of the root emetic and

896. Perfoliatum.

Leaves

cathartic.

diaphoretic.

SAMBUCUS.
Roots cathartic.

897. Ebulus.

Juice of the fruit cooling, laxative, and diuretic.

898. Nigra.

purgative

Flowers diaphoretic

emetic.

Bark

employed as ex-

Produces Elder- berries.

pectorants.

GALIACE^E,

OR

STELLATE.

RUBIA.
Root used

899. Tinctorum.

Said to be tonic, diuretic,

for dyeing.

Produces Madder.

and emmenagogue.

ASPERULA.
900. Odorata.

Diuretic.

ASTERACE^.
LlATRIS.

Roots have a terebinthinous odour, and are diuretic

904. Squarrosa

and

antisiphilitic.

905. Scariosa.

Diuretic, antisiphilitic.

EUPATORIUM.
All the parts bitter.

907- Perfoliatum.

In

warm

valuable tonic stimulant.

infusion or decoction emetic, sudorific,

and

aperient.

TUSSILAGO.

The

913. Farfara.

leaves,

smoked
It is

against dyspnoea.

have been employed


bitter, and a slight tonic.

like tobacco,

demulcent,

ERIGERON.
914. Philadelphicum

Used

as a diuretic.

STENACTIS.
915. Annua.

Employed

as a diuretic.

SOLIDAGO.
9 1 6. Odora.

Leaves yielding a volatile

oil,

which

is

aromatic,

sti-

mulant, diaphoretic, and carminative.

INULA.
919. Helenium

and other

Tonic, diuretic,
diseases.

diaphoretic.

Used

Produces Elecampane.

in

dyspepsia,

MEDICAL BOTANY.
PULICARIA.
920. Dysenterica

Astringent, diuretic.

BIDENS.

The whole

921. Tripartita.

When

plant acrid.

chewed,

it

excites

salivation.

SPILANTHES.

The whole

923. Oleracea.

plant acts as a powerful stimulant of the

salivary organs.

ANTHEMIS.
925. Nobilis.

Produces Chamomile heads.

Tonic, stimulant, emetic.

MARUTA.
926. Cotula.

Every part

is

and

foetid

active bitter, producing vomiting

Its

acrid.

decoction

is

an

and sweating.

ANACYCLUS.
Root hot,

927. Pyrethrum.

acrid oil in the bark,

acrid,

and permanent, depending on an


it a rubefacient and sti-

which renders

mulant.

PTARMICA.

The whole plant

928. Vulgaris.

is

pungent,

stimulant.

Dried

leaves produce sneezing.

PYRETHRUM.
The whole

930. Parthenium.

plant
mulating, and anti-hysteric.

is

bitter

considered tonic, sti-

ARTEMISIA.
932. Maritima.

Bitter, tonic, aromatic.

936. Glacialis.

Bitter, tonic, aromatic.

Leaves pungent and stimulating.

941. Dracunculus.

Is

Tarra-

gon.

943. Abrotanum.
944. Moxa.

powerful anthelmintic.
Furnishes a kind of Moxa.

945. Absinthium.
Is

powerful

Is Southern-wood.

bitter, tonic; extolled as

a stomachic.

Wormwood.

TANACETUM.
946. Vulgare.

Every part

cordial nature.

Is

bitter.

The

qualities are of a tonic

and

Tansy.

ARNICA.
948. Montana.

virulent plant, acting as a narcotico-acrid agent.

MEDICAL BOTANY.

270
DORONICUM.

Narcotico-acrid.

919. Pardalianches.

CALENDULA.

Employed as a carminative.

950. Officinalis.

CYNARACE^E.
CENTAUREA.
951. Calcitrapa. Bitter, febrifugal.
952. Centaurium.
Bitter, febrifugal.
954. Jacea.

Bitter, febrifugal.

SlLYBUM.
956. Marianum.

Leaves

sudorific

and

aperient.

LAPPA.
Root

957. Minor.
ter

and

tonic, aperient, sudorific,

acrid

and

diuretic.

Fruit bit-

also used as a diuretic.

CNICUS.
958. Benedictus.

Febrifugal.

CICHORACE^l.
LACTUCA.
959. Virosa.

Narcotic.

960. Sativa.

Sedative.

TARAXACUM.
The

961. Dens leonis.


are tonic

and

Produces Thridax.

infusion, decoction,

aperient.

and extract of the root

Diuretic.

CICHORIUM.
962. Intybus.

Root tonic and aperient. Used in decoction in chroand cutaneous diseases.

nic visceral

VALERIANACEJE.
VALERIANA.
964. Officinalis.

Roots

foetid,

as an anthelmintic.

stimulant,

and

narcotic.

Employed

Produces Valerian-root.

PLANTAGINACE^.
PLANTAGO.
968. Psy Ilium. Seeds mucilaginous, demulcent.
970. Cynops. Seeds mucilaginous, demulcent.
Leaves and roots bitter, astringent.
971. Lanceolata.
expectorant and vulnerary.

Used

as

an

MEDICAL BOTANY.

27 1

GLOBULARIACE7E.
GLOBULARIA.

A bitter, drastic purgative.


A bitter, drastic purgative,

972. Alypum.
973. Vulgaris.

employed as a resolvent

and vulnerary.

PLUMBAGINACEJE.
STATICE.

Root intensely astringent.

976. Caroliniana.

ABMERIA.
Flowers an active

977. Vulgaris.

diuretic.

PLUMBAGO.
978. Europaea

Very

acrid; used to remove tooth-ache.

An

effec-

tual emetic.

979. Rosea.

Acrid, vesicant.

981. Zeylanica.

Acrid, vesicant.

BORAGINACE^:.
BORAGO.

Root mucilaginous.

984. Officinalis.

Pectoral, emollient.

Is

Bo-

rage.

SYMPHYTUM.
986. Officinale

Reputed vulnerary, esculent.

CYNOGLOSSUM.
987. Officinale

Fffitid, narcotic,

LAMIACE^

antispasmodic.

OR

? ?

LABIATE.

LAVANDULA.
995. Vera.

Flowers carminative, stimulant, and tonic ; used with


Produces Lavender.

the leaves as sternutatories.

996. Spica. Yields oil of spike.


997. Stoechas. Considered expectorant and antispasmodic.

MENTHA.
999. Viridis.

Aromatic and carminative.

Produces Spearmint.

MEDICAL BOTANY.

272
MENTHA.

An

1000. Piperita.
dic.

aromatic stimulant.

Volatile

oil

antispasmo-

Produces Peppermint.

1001. Pulegium.
Aromatic, antispasmodic.
Furnishes a fragrant oil.
1002. Citrata.

Stomachic and emmenagogue.

1003. Rotundifolia.

Stomachic and emmenagogue.


Stomachic and emmenagogue.

1004-. Aquatica.

1005. Arvensis.

LYCOPUS.

1006

a.

Europseus.

febrifuge,

commended

as

an astringent.

SALVIA.
1008.

Officinalis.

Qualities aromatic, bitter,

1009. Grandiflora.

and stomachic.

Qualities aromatic, bitter,

and stomachic.

ROSMARINUS.
1010.

Officinalis.

Employed

as a cephalic medicine.

MONAEDA.
1011. Fistulosa

Bitter, aromatic, febrifugal.

AMABACUS.
Aromatic and

1013. Dictamnus.

tonic.

ORIGANUM.
1014. Vulgare.

Pungent, stimulant, and fragrant.

Produces Mar-

joram.

THYMUS.
1015. Vulgaris.

Pungent, stimulant, and fragrant.

1016. Serpyllum.

Pungent, stimulant, and fragrant.

HYSSOPUS.
1017.

Officinalis.

Stimulating, stomachic, carminative.

MELISSA.
1020. Calamintha.

Aromatic,

bitter, febrifugal.

SCUTELLABIA.
1021. Lateriflora

Reputed

to

be a remedy

for

hydrophobia.

NEPETA.
1022. Cataria

It acts as

a real aphrodisiac on cats.

Used

also in

amenorrhoea.

1023. Glechoma
Ivy.

Expectorant,

anti-hysterical.

Produces

Ground

MEDICAL BOTANY.

273

LEONURUS.
1024. Cardiaca.

Stimulant.

Formerly used against canine madness.

STACHYS.
1026. Betonica.

Root emetic and purgative.

Stimulating.

MABBUBIUM.
1027. Vulgare.

Herb,

lating

and

bitter,

tonic.

aromatic.

Expectorant.

Recommended
Is

as stimu-

Horehound.

VERBENACE.E.
VlTEX.
1030.

Leaves powerfully discutient.

Trifolia.

1031. Agnus castus.

Fruit acrid.

Fruit acrid, stimulant.

STACHYTABPHA.

The expressed

1036. Jamaicensis.
clysters,

and

as

juice purgative;

employed

for

an anthelmintic.

BIGNONIACE^E.
CATALPA.
Leaves and bark

1037. Syringifolia.

bitter, expectorant.

BIGNONIA.
1038. Antisiphilitica.

Discutient, anti- venereal.

ACANTHACE^;.
RHINACANTHUS.
Milk boiled on the roots

1039. Communis.
siacal

is

considered aphrodi-

also alexipharmic.

ACANTHUS.
1043. Mollis.

Leaves emollient.

ADHATODA.
1045. Vasica.

Flowers, leaves, and roots anti spasmodic, bitter, and

sub-aromatic.

ANDOGBAPHIS.
1046. Paniculata.

Stomachic,

dysentery.

used as a remedy

for

cholera and

Said to be alexipharmic.

SCROPHULARIACE^E.
DIGITALIS.

1047. Purpurea

Diuretic, narcotic.

Is Foxglove.

MEDICAL BOTANY.

27-i

SCROPHULARIA.
Leaves and

1048. Nodosa.

roots purgative

and emetic, with a

bitter

taste.

Leaves and roots purgative and emetic, but less


1049. Aquatica.
so than the last.

HEBPESTES.
1

Antirheumatic.

050. Monniera.

CALCEOLARIA.
Leaves purgative and emetic.

1053. Pinnata.

LINARIA.
1054. Vulgaris.
Bitter, purgative, and diuretic.
a wash for chronic diseases of the skin.

Recommended

1055. Cymbalaria.

Flowers used as

as an antiscorbutic.

Diuretic.

EUPHRASIA.
1059.

Officinalis.

1060.

Officinalis.

Slightly bitter

and aromatic, ophthalmic.

GRATIOLA.
Bitter, acting as

a purgative and emetic.

Useful

in cases of hypochondriasis.

SCOPARIA.
1061. Dulcis.

-Febrifugal

Expressed juice mucilaginous, and used

as a cooling laxative.

VERSA SCUM.
1062. Nigrum.

Sub-narcotic.

SOLANACE^l.
HYOSCYAMUS.
1065. Niger.

powerful narcotic.

Is

Henbane.

ATROPA.

1066. Belladonna.

dangerous narcotic.
Every part of the plant
In medicine it is narcotic,
diaphoretic, and

poisonous.
Is

diuretic.

Deadly Nightshade.

CAPSICUM.
1067.

Annuum.

Fruit and seeds stimulant,


pungent.
Fruit and seeds
stimulant, more pungent.
Baccatum. Fruit and seeds
stimulant, very pungent.

1067

a. Frutescens.

1067

b.

DATURA.
1068. Tatula

violent narcotic
poison.

an anodyne and sedative.

Employed externally

as

MEDICAL BOTANY.

275

DATURA.

1069. Stramonium.

violent narcotic

nally as an anodyne

and

poison.

Employed exter-

sedative.

PHYSALIS.
1072. Alkekengi.

Diuretic,

employed in veterinary

practice.

NICANDBA.
1073. Physaloides.

Diuretic.

SOLANUM.
1074. Nigrum. Stimulating, narcotic.
1076. Dulcamara.
Berries bitter and poisonous.

Plant narcotic and

diaphoretic.

NICOTIAN A.

1081. Tabacum.

infusion as

stimulant narcotic, employed as an errhine ; in


an expectorant and sedative ; in vapour as an

Produces Virginian Tobacco.


antispasmodic.
stimulant narcotic, much more mild in

1082. Rustica.
tion.

1083. Persica

its

opera-

Produces Syrian Tobacco.

stimulating narcotic, less mild in

its

operation.

Produces Persian Tobacco.

CESTRACE^E.
OESTRUM.
Febrifugal, used externally as an astringent.

1087. Laurifolium.

GENTIANACE^:.
GENTIANA.
1088. Catesbsei.

Bitter, tonic, febrifugal.

1089. Amarella.

Bitter, tonic, febrifugal.

1090. Campestris.
Bitter, tonic, febrifugal.
1091. Purpurea.
Bitter, tonic, febrifugal.
Root
1095. Lutea.
Bitter, tonic, febrifugal.

anthelmintic.

Pro-

duces the Gentian-root of the shops.

FRAZERA.
1097. Carolinensis.

Root

bitter, emetic, cathartic.

ERYTHR^A.
1099. Centaurium.

Bitter, tonic, febrifugal.

Used

in rustic phar-

macy.
T 2

MEDICAL BOTANY.

276

MENYANTHES.
All the plant

1105. Trifoliata.

bitter.

valuable tonic, emetic,

diaphoretic.

VILLARSIA.

Stems

1106. Nymphseoides.

bitter, tonic,

and

febrifugal.

SPIGELIA.
1 1

Root and leaves are active anthelmintics

07. Marilandica.

also

Produces Wormseed.

purgative and narcotic.

APOCYNACE^E.
CERBERA.
Kernels

1111. Manghas

emetic

and poisonous;

the milky

employed as a purgative.
1113. Thevetia. Bark bitter, cathartic, and a powerful

sap

febrifuge.

ALLAMANDA.

An

1125. Cathartica.

infusion

of the leaves a valuable cathartic.

In over-doses emetic and purgative.

NERIUM.
1128. Oleander.

Acrid, stimulating, poisonous.

APOCYNUM.
1130. Androssemifolium.

Every part

lactescent.

Root

bitter, tonic,

acting as an emetic.

1131. Cannabinum.

Emetic

in decoction diuretic

and

diaphoretic.

PLUMIERA.
1137. Rubra.

Milk

corrosive.

ASCLEPIADACE^:.
ASCLEPIAS.

1141. Tuberosa.
catarrh,

Root expectorant and


diaphoretic;

pneumony, and

pleurisy.

employed in

Useful as a tonic and

stimulant.

1143. Curassavica.

Roots purgative and emetic.

CALOTROPIS.

1144.

Gigantea.The juices
tives

1145. Procera

of the root and bark are used as altera-

and purgatives.

Produces Mudar.

Juice acrid; also a


powerful depilatory.

CYNANCHUM.
1148. Vincetoxicum.

Emetic and purgative, celebrated as an an-

tidote to poisons.

MEDICAL BOTANY.

277

OLEACE.E.
OLEA.
1157. Europsea.

The

emollient,

fruit

and

duces Olive

yields

an

Bark

laxative.

which

oil,

bitter

and

is

demulcent,

astringent.

Pro-

oil.

OBNUS.
1158. Europaea.
The branches yield true Manna. A gentle laxative.
1159. Rotundifolia.
The branches yield Manna of a better quality.

FBAXINUS.
1

160. Excelsior.

Leaves cathartic.

Bark

tonic

and

febrifugal.

SYRINGA.
1161. Vulgaris.

Bark

tonic, bitter,

and

febrifugal.

CYCADACE^E.
CYCAS.
1 1

62. Revoluta.

Farinaceous.

ZAMIA.
1

Yields a kind of Arrow-root.

169. Furfuracea.

:,

OR

CONIFERS.

PIN us.
1170. Sylvestris.

Terebinthinous,

resinous.

Produces Turpentine,

resinous.

Produces

Pitch.

1171. Pumilio

-- Terebinthinous,

Hungarian

balsam.
1

72. Pinaster.

Produces Bordeaux tur-

Terebinthinous, resinous.

pentine.

1173. Cembra.

Terebinthinous,

resinous.

Produces

Carpathian

balsam.
ABIES.

1174. Picea.

Terebinthinous,

resinous.

Produces Strasburgh tur-

pentine.

1175. Balsamea.

Terebinthinous, resinous.

Produces Canada bal-

sam.
1176. Larix.

Terebinthinous, resinous.

Produces Venice turpentine.

CALLITRIS.

1177. Quadrivalvis.

Resinous.

Produces Sandarach.

MEDICAL BOTANY.

278
JUNIPERUS.
1

78.

Fruit sudorific, carminative

Communis.

the

oil

a very pow-

Produces Juniper-berries.
The oil is a powerful stimulant, acting as a
1179. Virginiana.
In amenorrhoea it acts as an
rubefacient and vesicant.
erful diuretic.

emmenagogue.

Is Savin.

Diuretic.

Oil a powerful stimulant, acting as a rubefacient and


In amenorrhoea it acts as an emmenagogue.
vesicant.

1180. Sabina.

Is Savin.

Diuretic.

TAXACE^E.
TAXUS.
Leaves fetid, very poisonous, acting like Digitalis.

1181. Baccata.

Berries harmless.

Is the

Yew.

ZINGIBERACE.E.
ZlNGIBEB.
1 1

82. Officinale.

valuable aromatic, carminative, stimulant, sia-

Produces Ginger.

lagogue.

CURCUMA.
Root aromatic, stomachic, carminative.

1186. Zedoaria.

Produces

Zedoary.
1189. Longa. Bitter, aromatic, stimulant, tonic.
Produces Turmeric.

Used

in

dyeing.

K^EMPFEHIA.
1 1

The

92. Galanga.

roots

have an agreeable smell, and

warm

have an agreeable smell, and

warm bitter

bitter

aromatic taste.

The

1193. Rotunda.

roots

aromatic taste.

MARANTACEjE.
MARANTA.
1204. Arundinacea.
mic.

Amylaceous.

Reckoned a powerful alexiphar-

Produces Arrow-root.

CANNA.
1

205. Edulis.

Amylaceous.

1206. Coccinea.

Amylaceous.

Produces Tous

les

mois.

AMARYLLIDACE.E.
CRINUM.
1207. Asiaticum.

Bulbs powerfully emetic, poisonous.

MEDICAL BOTANY.

279

OPORANTHUS.
1208. Luteus

Bulbs purgative.

BRUNSVIGIA.
1209. Toxicaria.

Juice of the bulbs a dangerous poison.

Used

to

envenom arrows.
NARCISSUS.

1210. Poeticus.

Bulbs emetic, poisonous.


Bulbs and flowers emetic, poisonous.
Emetic and poisonous.

1211. Pseudo-narcissus.

1212. Tazzetta.

PANCRATIUM.
1213. Maritimum.

Emetic.

ALSTR(EMERIA.
1214.

Salsilla.

Diuretic

and

diaphoretic.

IRIDACE^E.
IRIS.

Rhizoma nauseous and

1216. Versicolor.

acrid

an active cathartic.

Useful as a diuretic.

Rhizoma acrid; possessing purgative and


emetic properties.
Rhizoma a sub-acrid, aromatic, bitter substance.
1218. Florentina

1217. Pseud-acorus.

Produces Orris-root.

CROCUS.
1219. Sativus.

Stimulant.

and emmenagogue.

Used

as

carminative,

antispasmodic.

Produces Saffron.

ORCHIDACE^E.
ORCHIS.

Amylaceous, demulcent.

1221. Mascula.

Produces Salep.

BLETIA.
1225. Verecunda.

Bitter, stimulant, stomachic.

PALMACE^E.
CARYOTA.
1232. Urens.

Amylaceous.

CALAMUS.
1233. Draco.

Astringent.

Produces Sago.

MEDICAL BOTANY.

280
ELAIS.

1234. Guineensis.

Palm

Produces

Oleaginous, astringent.

oil.

MELANTHACE^E.
VERATRUM.
1236. Viride.

Roots an acrid emetic, stimulant, sedative.

White

1237. Album.

Produces

Hellebore.

small

acts as

dose

a large causes

an emetic;

Produces White Hellebore.

vomiting and purging.

Seeds acrid; used as anthelmintics.

1238. Sabadilla.

dangerous

stimulant.

HELONIAS.

Used

Plant a narcotic poison.

1241. Erythrosperma.

for destroying

flies.
1

242. Dioica.

Root in infusion anthelmintic

and

in tincture bitter

tonic.

GYKOMIA.
Root

1243. Virginica.

diuretic,

hydragogue.

TRILLIUM.
1244. Erectum.

Rhizoma

violently emetic

fruit suspicious.

COLCHICUM.
1246. Autumnale.

Sudorific,

thelmintic.

emetic,

purgative.

narcotico-acrid

poison.

Used
Is

the

as an an-

Meadow

Saffron.

LILIACE.E.
ERYTHRONIUM.
1247. Americanum. Root and

leaves emetic.

ALETRIS.
1249. Farinosa.

Very

machic.

bitter.

Used

in infusion as

a tonic and

sto-

Emetic.

SQUILL A.
1250. Maritima

Bulbs acrid, vesicant, emetic, diuretic,


expectorant.

Produces Squills.

ALLIUM.
1255. Sativum.

Bulbs stimulant,
expectorant,
as anthelmintics.
Produces Garlic.

and

diuretic.

1256. Cepa.
Stimulant, diuretic, expectorant, and rubefacient.
duces Onions.

Used
Pro-

MEDICAL BOTANY.

281

DRACAENA.
1258. Draco.

Tonic,

astringent,

resinous,

employed

in

diarrhoea.

Produces Dragon's blood.


Roots astringent, useful in dysentery.
1259. Terminalis.
1260. Ferrea. Roots astringent, useful in dysentery.

ALOE.
1261. Vulgaris.
1262. Socotrina.

Purgative.

Produces Barbadoes Aloes.


Produces

Purgative, bitter, aromatic.

Socotrine

and Mocha Aloes.


1263. Purpurascens.

Purgative,

aromatic.

bitter,

Produces Soco-

trine Aloes.

1264. Spicata.

Produces Cape Aloes and Horse Aloes.


Produces Cape Aloes and Horse
Purgative.

Purgative.

1265. Arborescens.
Aloes.

1266. Commelyni.

Cape Aloes and Horse

Purgative.

Produces

Purgative.

Produces Cape

Aloes.

1267. Mitriformis.

Aloes

and Horse

Aloes.

SMILACE^E.
SMILAX.
1269. Aspera.

Emetic, diaphoretic, narcotic.

Produces Italian Sar-

saparilla.

1270. Sarsaparilla.

1272.

Emetic, diaphoretic, narcotic.


Emetic, diaphoretic, narcotic.

Siphilitica.

Produces

Lisbon

Sarsaparilla.

ARACE.E.
ARUM.
Tubers amylaceous, stimulant, diaphoretic, and
Produces Portland
expectorant; juice acrid, poisonous.

1280. Maculatum.

Sago.

COLOCASIA.
1282. Esculenta.

Acrid, sialagogue, amylaceous.

SYMPLOCARPUS.
Tubers

1285. Foetidus.

acrid, antispasmodic,

hydragogue.

DlEFFENBACHIA.
1289. Seguina.

An

parts an

exceedingly venomous plant.

indelible stain to linen.

The

juice

im-

MEDICAL BOTANY.

282

ACORACE^E.
ACORUS.

Rhizoma aromatic,

1290. Calamus.

bitter, stomachic.

Adapted

to

Produces Calamus aromaticus.

cases of dyspepsia.

GRAMINACE^E.
LOLIUM.

1292. Temulentum.

narcotico-acrid poison.

Used

as a sedative

Produces Darnel.

poultice.

TBITICUM.
1293. Vulgare.

Produces Wheat.

Amylaceous.

HORDEUM.
Produces Barley.

Nutritive.

1294. Vulgare.

SECALE.
Origin of Ergot.

1295. Cereale.

BROMUS.
1296. Mollis.

Narcotic.

Emetic.

1297. Purgans.

AVENA.
1300. Sativa.

Nutritive.

Produces Groats.

ANDROPOGON.
1302. Schoenanthus.

Leaves stomachic, aromatic,

bitter.

SACCHARUM.
1305. Officinarum.

Nutritive.

An

antidote to arsenic.

Produces

Sugar.

CYPERACE^:.
CYPERUS.
1
1

Stomachic.
306. Longus.
Stomachic.
307. Rotundus.

Tubers useful in cholera.

CAREX.
1308. Arenaria.
tive.

Creeping stems diaphoretic, demulcent, and alteraProduces German Sarsaparilla.

Creeping stems diaphoretic, demulcent, and alterative.


Produces German Sarsaparilla.

1309. Hirta.

1310. Intermedia.
terative.

Creeping stems diaphoretic, demulcent, and alProduces German Sarsaparilla.

MEDICAL BOTANY.

283

XYRIDACE^E.
XYRIS.
1311. Indica.

Used against ringworm.

FILICALES.
ADIANTUM.
1315. Capillus Veneris.

Rhizoma

astringent

and aromatic, pectoral

the decoction emetic.

1316. Pedatum.

Rhizoma

astringent

and aromatic, pectoral

the

Used

as a

decoction emetic.

PTEBIS.

1317. Aquilina.

Rhizoma

astringent

and anthelmintic.

substitute for Hops.

NEPHKODIUM.
Rhizoma anthelmintic.

1318. Filix mas.

OSMUNDA.
Rhizoma

1319. Regalis.

rachitis.

tonic

and

styptic,

useful

in

cases

of

INDEX.
ACANTHACE.E, 154, 273.
Acanthales, 217.
Aceraceae, 109.
Acerales, 203.
Achenium, 5.9, 63.

Achlamydeous, 41.
Aclilamydosse, 209.
Acoraceae, 185, 282.
Acotyledonous, 67.

Acrogenous, 12.
Acrogens, 18, 68, 191.

Actinenchyma,
Acuminate, 29.

3.

Annulus, 69, 73.


Anonaceae, 92.
Anonales, 199.
Anther, 45, 46.

Antheridia, 71.
Anthocarpous, 59.
Antitropous, 67.
Apex, 54, 57.
Apiaceae, 134, 242.
Apocarpous, 50.

Apocynaceae, 146, 276.


Apophysis, 69.
Apothecia, 71.

Acute, 29.

Appendages

Acute-angled, 20.
Adherent, 230.
Adventitious leaf-buds, 23.
-ffisculaceae, 109, 249.

Araceae, 186, 281.


Arales, 222.

-(Estivation, 36.

jEtheogamous plants, 91.


Age of Exogenous trees, 1 4.
Aggregosae, 213.

of the axis, 9.
Aquifoliaceae, 144, 266.

Araliaceae, 133, 245.

Araneose, 9.
Arcuate, 66.
Aril, 63.
Aristolochiaceae, 162, 263.

Aigrette, 41.-

Aristolochiales, 212.

Air-vessels, 6.

Arrow-headed, 27.

Albumen, 65.
Alburnum, 15.

Articulated, 18.

Algaceae, 69, 195.


Algae, 195.

Alismaceae, 185.
Alismales, 221.
Alliances of plants, 197.
Alsineae, 104.
Alternate, 37.

Amarantaceae, 158.
Amaryllidaceae, 177, 278.
Amentales, 209.

Amentum,

38.

pitted tissue,
Artocarpeae, 167.

Ascending, 18.
Asci, 71.

Ascidium, 25.
Asclepiadacese,

46, 276.

Assimilating power, 32.


Asteraceae, 138, 268.
Asterales, 215.

Atractenchyma,

3.

Atropous, 55.

Amomales, 220.

Aurantiaceae, 106, 251.


Auriculate, 25, 26.

Amphigamous

Axis, 9.

plants, 91.

Amphitropous, 55.
Amplexicaul, 25.
Ampulla, 25.
Amygdalae, 123, 255.
Amyridaceae, 124.
Anacardiaceae, 119, 259.
Analytical method, 78.
Anatropous, 55.
Andraeaceae, 193.
Andrceceum, 45.
Angular, 27.
Aiuiular, 6.

Balanophoracege, 174.
Balausta, 62.
Balsamaceae, 262.
Balsamales, 208.

Balsaminaceae, 114.

Bark, 15.
Base, 57.
Basidia, 72.

Begoniaceae, 158.

Begoniales, 202.
Berberaceae, 94, 245.

4.

285

INDEX.
Berberales, 200.

Cedrelaceae,

Berry, 61.

Celastraceae, 117, 253.


Cellular, 2.

Betulaceae,

168,259.

111,250.

Bignoniaceae, 147, 273.


Bignoniales, 217.
Bijugute, 28.

Cellular plants, 91.


Cells, 47.
Central, 50.

Bipinnate, 28.

Centrifugal, 39.
Centripetal, 39.

Biserrate, 29.
Biternate, 28.

Cercodiaceae, 127.

BixaceJE, 100, 248.


Bixales, 203.

Boraginaceae, 150, 271.

Bothrenchyma,
Bourgeon, 20.
Bouton, 20.

4.

Cestraceae, 275.
Chalaza, 55, 65.

Chamaelaucieae, 126.
Channelled, 25.
Characeae, 192.
Charales, 224.

Bractlets, 35.
Bracteolae, 35.

Chenopodales, 212.
Chenopodiaceae, 156, 263.
Chloranthaceae, 164, 261.

Branched

Chlorophyll, 2.

Bract, 35.

cellular tissue, 3.

Brassicaceae, 97, 247.

Brexiales, 213.
Bromeliaceae, 179.

Bromeliales, 220.

Brunoniales, 216.
Bryaceae, 192.

Bud, embryo,
Buds, 20.

9.

Chromogen, 2.
Chromule, 2.
Chrysobalanaceae, 124.
Cichoraceae, 1 39, 270.
Cinchonaceae, 136, 267.
Cinchonales, 215.
6.

Cinenchyma,

Circaeeae, 127.

Bulbills, 21.
Bulblets, 21.
Bulbs, 21, 22.

Cistaceae, 100, 249.

Butomaceae, 183.

destines,

Cactaceae, 131.

Closed, 6.
Cloves, 21.

Caesalpinieae, 120.
Calcarate, 42.
Calceolate, 42.

Callitrichales,211.

Calycanthaceae, 124.
Calyciflone, 91, 117.
Calyptra, 70.
Calyptrate, 42.
Calyx, 40, 41, 54.

Cambium,

17.

Campanales, 2 1 5.
Campanulaceae, 142.
Campanulate, 42.
Campylotropous, 55.
Canellese, 248.

Cistales, 204.

Cladenchyma,

Clusiaceae, 108.

Cocci, 60.
Cocculales, 212.
Collar, 66, 73.

Colpenchyma,

Capparidaceae, 98, 247.


Capriales, 215.
Caprifoliaceae, 136, 268.
Capsule, 60, 63.
Carpel, 50.

Carunculae, 64.

Caryophyllaceae, 104.
Caryopsis, 60, 63.
Casuarales, 210.
Caudate, 60.
Caudicle, 48.
Cauliculus, 66.

3.

Columella, 58, 69.


Column, 46.
Columnosae, 209.

Coma, 64.
Combretaceae, 124, 245.
Commelynaceae, 183.
Commelynales, 221.

Commissure, 58, 61.


Compositae, 138.
Compound, 40.

Capillitium, 73.

Capitulum, 38.

3.

3.

organs, 9.

Compressed, 20.
cellular tissue, 3.

Cone, 63.

Conenchyma,

3.

Conical, 11.
cellular tissue, 3.

Coniferce, 171, 277.


Coniocysta, 72.

Conjugato-pinnate, 28.
Connective, 46.
Consolidated, 230.

Continuous pitted (tissue),


Convolvulaceae, 1 49, 266.

4.

INDEX.

286
Convolute, 37, 67.
Coralline, 11.

Cordate, 26.
Cordate-acuminate, 26.
Cordate-ovate, 26.
Cordiaceae, 150.

254.

Coriariaceae, 117,

Coriales, 207.

Corm, 19, 21.

Dicarposae, 213.
Dicotyledonous, 67.
Dictyogens, 12, 230.

Didynamous, 46.
Digitate, 28.
Dilated, 42.
Dilleniacese, 95.
Dioscoreaceae, 178.
Diploe, 26.

Cornales, 201.

Dipsaceoe, 137.
Dipsales, 216.
Dipteraceae, 106.

Corolla, 40, 42.


Coroniform, 42.

Disciform, 13.
Disk, 48.

Cortina, 73.

Dissepiments, 52.
Dissolved, 230.
Disunited, 230.

Cormogens, 230.
Cornaceae, 133, 246.

Corylaceae, 169,259.
Corymb, 39.

Divergent, 67.
Dorsal suture, 58.

Corymbiferae, 139.
Cotyledons, 66.
Coussinet, 25.
Crassales, 208.

Dotted ducts,
Double, 42.

Crassulaceae, 131,258.
Creeping, 18.

Droseraceae, 101.
Drupe, 59, 63.

Cremocarp, 61.

Ducts, 4, 5.

Crenate, 29.
Crisp, 29.
Cruciales, 202.

Duplicato-dentate, 29.

Cruciferae, 97, 247.

Echiales, 216.
Elaeagnacese, 160.

Cryptogamous

4.

Ebenaceae, 144, 266.


plants, 90.

Cucurbitaceee, 129, 246.

Elatinaceae, 104.

Cucurbitales, 202.
Cuneate, 27.
Cup, 48.

Elementary organs,
Emarginate, 29.

10, 65, 66.


buds, 23.
Empetraceae, 162.

Embryo,

Cupule, 61.
Cupuliferae, 169.

Curled, 29.

Curvembryosae, 209.
Cuticle, 7.

Cycadaceae, 170, 277.

Cylindrenchyma,

3.

Cylindrical tissue,
Cyme, 39.

3.

Endocarp, 57.
Endogenae, 173.
Endogenous, 12.
Endogens, 17.

Endophkeum,

15.

Endopleura, 64.
Endorhizae, 68.

Cynaraceae, 139, 270.

Endothecium, 48.

Cyperaceae, 188,282.
Cyrtandraceae, 147.
Cystidia, 72.
Cytinaceae, 174.

Entangled

Cytoblast, 2, 4.

Dasdalenchyma,

3.

Danaeaceae, 191.

cellular tissue, 3.

Entire, 29.
Epicarp, 57.

Epidermis, 15.
Epigynosae, 213, 219.
Epigynous, 46.

Epiphkeum, 15.

Decompound, 28.

Epiphyllous, 37.
Equisetaceae, 192.
Erect, 66.
Ericaceae, 143, 265.

Deformed, 18.

Ericales, 213.

Dehisce, 58.
Dehiscence, 47, 58.
Dehiscent, 58.
Dentate, 29.

Escalloniaceae, 133.
Etaerio, 61.

Daphnales, 211.
Datiscales, 210.

Diplecolobeae, 97.
Diachyma, 26.

Diadelphous, 46.

2.

Etiolated, 33.

Euphorbiaceae, 162, 251.


Euphorbiales, 205.
Excentrical, 66.

Excipulus, 72.

INDEX.
Exogenae, 91.
Exogens, 13.
Exogenous, 12.

Gesneraceae, 142.

Gland, 61.
Glandular hairs, 8, 9.
Gleicheniaceae, 191.

Exorhizae, 68.
Exostome, 55.
Expansion, order of, 37.
Extra-axillary, 37.

Globulariaceae, 154, 271.


Globuli, 72.

Glochidiate, 9.

Glomeruli, 72.
Glumaceae, 188.

Fabaceae, 119,256.

Glumes, 35.

False-bark, 17.
Fasciculate, 11.

Glumosae, 219.
Gongyli, 71, 72.
Goodeniaceae, 141.
Goodeniales, 215.

Female organs, 49.


Ferns, 69.
Ferula, 13.
Funiculus, 54.

Graminaceae, 189, 282.


Granada, 72.
Grossales, 200.
Grossulaceae, 132, 245.
Guttales, 203.

Fibro-cellular tissue, 3.

Ficoidales, 202.

Ficoideae, 131.
Fig, 63.
Filaments, 45, 46.
Filicales,

287

Guttiferae, 108.

Gymnogens, 230.

191,223,283.

Filices, 191.

Gynandrosae, 219.

Flacourtiaceae, 100.

Gynandrous, 46.
Gynobase, 49.
Gynoeceum, 49.
Gynophore, 50.
Gyroma, 71.

Flat, 67.
Flocci, 73.

Florkeales, 207.
Floral envelopes, 35, 40.
Floridae, 174.

Flower-bearing stems,
Flower-buds, 20, 34.

8.

8.

Half-terete, 20.

Flower-leaf, 35.

Halorageae, 128.
Hamamelaceae, 133.

Flowering plants, 90.


Flowerless plants, 68, 90.
Fluviales,

Hairs,

185,222.

Hastate, 27.
Heart- wood, 15.

Foliola, 28.

Heliacal, 66.
Hepaticae, 194.

Follicle, 59.

Hernandiaceae, 262.

Food

Heterotropous, 67.
Hilum, 64.

Foliaceous, 67.

of plants, 30.

Foramen, 55.
Forked, 46.

Hippurideae, 127.
Homogens, 230.
Humiriaceae, 112.
Hydrales, 220.

Frankeniaceae, 103.
Fruit, 57.
Fuchsieae, 127.
Fumarieae, 96, 241.
Fungaceae, 69, 72, 195.

Hydrocharaceae, 174.
Hydrophyllaceae, 151.

Fungales, 224.
Fungi, 195.

Fungous, 67.
Funnel-shaped, 42.
Fusiform cellular tissue,
Galbulus, 63.
Galeate, 42.
Galiaceae,

141,268.

Gamopetalous, 42.
Gamosepalous, 41.

Gemmation, 30.
Gemmule, 66.

Hymenium,

73.

Hypericaceae, 107, 248.


Hypha, 72.

Hypogynosae, 219.
3.

Hypogynous, 45.
Hypothecium, 71.
Illecebraceae, 130.

Imbricate, 37.

Impregnation, 56.
Indehiscent, 58.
Indifferent, 44.
Induplicate, 37.
Indusium, 69.

Gentianaceae, 148, 275.


Gentianales, 218.

Inenchyma,

Geraniaceae, 113, 254.

Inferior, 42, 54.

Geraniales, 206.

Inflated, 25.

3.

INDEX.

288
Inflorescence, 37.
Intercellular passages, 2.
Internodes, 19.

Interruptedly pinnate, 28.


Inverted, 66.
Involucre, 35.
Iridaceae, 179, 279.
Irregular, 43.
Irritability, 33.
Ixiales, 220.

Jasminacese, 146.
Jointed, 25.
Juglandacese, 170, 261.
Juncaceae, 182.
Juncaginaceae, 184.
Juncales, 221.

Jungermanniaceae, 193.
knotted, 18.
Labiales, 217.
Labiatae, 271.
Labiate, 42.

Lobes, 46.
Loculicidal, 58.

Loganiales, 218.
Lomentaceous, 59.

Loranthaceae, 135.

Lycopodales, 224.
Lycopodiaceae, 191.

Lymphatic

hairs, 8, 9.

Lyrate, 28.
Lythracea:, 124, 250.

Mace, 63.
Magnoliaceae, 95, 242.
Male organs, 45.
Malpighiaceae, 110.

Malvaceae, 105, 250.


Malvales, 204.
Marantaceae, 177, 278.
Marchantiacese, 194.

Marginal placenta, 50.


Medical Botany, 239.
Medulla, 3.
Medullary plates, 16.
rays, 16.
sheath, 13.

Labiatiflorae, 139.

Lacunae, 72.
Lamiaceae, 153, 271.
Lamina, 24, 25.
Lanceolate, 26.

Melanthaceaa, 181, 280.


Melastomaceae, 124.

Latex, 6, 16.

Meliales, 205.

Laticiferous, 6.

Menispermaceae, 93.

Lauraceae, 158, 262.


Laureales, 211.

Merenchyma,

Leaf, simple, 26.


Leaf-buds, 10, 20, 22.
adventitious, 23

deciduous, 21.
normal, 22.
Leafless, 18, 25.
Leaflets, 28.

Leafy, stem, 18.


Leaves, 23.

compound, 28.
Legume, 59, 63.
Leguminosse, 119, 256.
Lentibales, 217.
Lentibulaceae, 154.
Lenticular, 67.
glands, 6.

Leptospermeae, 126.
Liber, 4, 15.

Meliaceae, 112,250.

3.

Mericarps, 61.

Mesembryaceae, 131.

Mesophkeum,

15.

Mesophyllum, 26.
Mesosperm, 64.
Micropyle, 65.
Midrib, 26.
Mimoseae, 121.

Monadelphous, 46.
Moniliform, 11.
Monimiales, 210.

Monochlamydeae, 91.
Monocotyledonous, 67.
Monopetalous, 42.
Monophyllous, 41.
Monosepalous, 41.
Moringaceae, 248.

Mosses, 69, 70.


Mucronate, 29.

Lichenaceae, 194.
Lichenes, 194.

Mucus,

Lichens, 69.

Multiple, 59.

organic, 2, 4.

Multangular, 20.

Lignification, matter of, 2.

Muriform

Liliaceae, 180, 280.


Liliales, 221.

Loasaceae, 129.

Musaceae, 177.
Muscales, 224.
Musci, 192.
Mycelia, 73.
Myricaceae, 168, 260.
Myristicaceae, 160, 242.
Myrsinaceae, 145.

Lobeliaceae, 142, 267.

Myrtaceae, 126, 24

Limb, 41, 43.


Linacese, 114, 249.

Linnean Sexual System, 76.


Lirella, 71.

cellular tissue,

>.

6.

289

INDEX.
Myrtales, 201.
Myrteze, 126.
Naiadaceae, 185.
Naked, 41.
Narcissales, 220.
Natural System of
Nectary, 43, 49.
Nelumbiaceae, 95.
Nepenthales, 212.
Nervures, 26.

De

Candolle, 90.

Nodes, 19.
Nodose, 11.

Paleae, 35.
Paleaceous, 60.
Palmaceae, 183, 279.
Palmales, 221.
Palmate, 28.
Pandales, 222.
Panduriform, 28.
Panicle, 39.

Papaveraceae, 98, 241.


Papayaceae, 248.
Papilionaceae, 120.
Papilionaceous, 42, 43.

Nolanales, 214.

Pappose, 60.
Pappus, 41.

Normal

Parenchyma,

leaf-buds, 22.

8.

Notorhizeae, 97.

Parietal, 52.

Nucamentosae, 213.

Passifloraceae, 129,

Nuclei, 21.
Nucleus, 54, 71.
Nuculanium, 61.

Passionales, 203.
Patellula, 71.
Pedate, 28.

Nut, 61, 63.


Nyctaginacese, 157, 264.
Nymphaeaceae, 94, 241.

Pedicels, 36.
Peduncle, 36.

248.

Pelta, 71.
Peltate, 9, 26.

Oblique, 26, 42.


Oblong, 26.

Penaeales, 211.
Pepo, 62.

Obscurely triquetrous, 20.


Obtuse, 29.

Perianthium, 41.

angled, 20.
Ochreae, 29.
Octangular, 20.

CEnothereae, 127.
Oleacese, 145, 277.

Oleales,218.

Pericarp, 57.
Peridiolum, 72.
Perigonium, 41.

Perigynous, 45.
Perisperm, 65.
Peristome, 69.
Perithecium, 71, 73.

Onagraceae, 126.
Onagrales, 201.
One-lipped, 43.

Petals, 42.
Petiole, 24.

Operculum, 70.

Petivales, 212.
Petiveriaceae, 156, 264.

Ophioglossaceae, 191.
Oplarium, 71.

Opophora,

6.

Opposite, 23.
the leaves (inflorescence), 37.
Orange, 62.
Orbicular, 26.
Orbiculi, 73.
Orbilla, 71.

Orchidaceae, 279.

Organs, compound, 9.
elementary, 2.
Orobanchaceae, 152.
Orthoploceae, 97.

Petiolar, 37.

Pha?nogamous plants, 90.


Phaceliales, 216.
Phanerogamous plants, 90.
Philadelphaceae, 125.
Phragmata, 53.

Phycomater, 72.
Phyllodium, 24.
Physiological Botany,

Pinaceaa, 171,277.

Pine-apple, 63.
Pinnate, 28.

Orthotropous, 55, 67.


Osmundaceae, 191.

Pinnatitid, 28.

Ostiolum, 73.
Oval, 26.

Piperales, 210.
Pistil, 49.

cellular tissue, 3.

Ovary, 49, 54.


Ovate, 26.

Ovenchyma,

3.

Ovule, 48, 50, 54.


Oxalidaceae, 114, 254.

1.

Phytolaccaceae, 156, 263.


Pileus, 73.

Piperaceae, 164, 261.

Pistillidium, 71.

Pitcher, 25.
Pith, 13.
Pitted tissue, 4.
Pittosporaceae, 114.
Pittosporales, 200.

INDEX.

290
Placentiform, 11.

Piano-compressed, 20.
Plantaginaceae, 155, 270.
Plantales, 216.
Platanaceae, 166.
4.

Pleurenchyma,

Pleurorhizese, 97.
Plicative, 37.

Plumbaginaceae, 154, 271.


Plumbales, 216.

Receptacle, 39, 49.

Rectembryosae, 209.
Regular leaf-buds, 22.
Reniform, 26.

Replum, 59.
Resedaceae, 99.
Reservoirs of oil, 6.
Reticulated, 6.

Plumose, 60.
Plumule, 66.

Retosae, 219.
Retuse, 29.

Podetia, 71.

Rhamnaceae, 118, 251.


Rhamnales, 205.

Podophylleae, 241.
Podospenn, 54.

Podostemales, 210.
Polemoniaceae, 148.

Rhizanthae, 173.
Rhizanths, 230.

Rhizoma,

19.

Pollen, 46.

Ribs, 26.

Polyadelphous, 46.
Polycarposae, 213.
Polygakceae, 102, 249.
Polygonaceae, 159, 264.

Ringent, 42.
Root, 10.

Polygonales,212.
Polypetalous, 42.
Polypodiaceae, 191.
Polysepalous, 41.

Pome,

62.

Pomeae, 123, 255.


Pores, 47.
Portulacaceae, 130.
Potentilleae, 122.

Rootshaped, 18.
Rootstock, 19.
Rosacese, 122, 254.
Resales, 207.
Roseae, 122.
Rostrate, 60.

Roundish oblong, 26.


Ruminated, 65.
Rutaceae, 115, 253.
Rutales, 206.

Praefloration, 36.

Sac of the amnios, 55.

Prickles, 9, 19.

Sagittale, 27.

Primine, 54.

Salicaceae, 165, 261.

Primulaceae, 155, 266.


Primulales, 214.
Prismatical cellular tissue, 3.

Salicales,

Prismenchyma,

3.

Proper juice, 34.


vessels, 6,

Prosenchyma,

2, 3.

210.
Samara, 61.
Sanguisorbeae, 123, 256.
Santalaceae, 161.
Santalales, 211.
Sap, 33.

Sapindaceae, 110, 249.

Pulp, 2.

Sapotaceae, 145, 266.


Sarcocarp, 57.
Sarraceniaceae, 96.

Putamen, 57.

Saururaceae, 165.

Pyrolaceae, 265.
Pyxis, 61.

Saxales, 208.

Quadrangular, 20.

Scalariform, 6.
Scales, 35.

Proteaceae, 160.
Proteales, 211.

Raphe, 55, 65.


Raphides, 3, 9.

Saxifragaceae, 132, 258.


Scaevolaceae, 141.

Quadrilateral, 20.

Quadriquetrous, 20.
Quincuncial, 37.
Quinquangular, 20.
Quinquelateral, 20.

Scaly bulbs, 21.

Raceme, 38.

Sclerales, 212.

Racliis, 36.

Ranales, 199.

Sclerogen, 2, 3, 4.
Scrophulales, 217.
Scrophulariaceae, 152, 273.
Scutate, 9.
Scutellum, 71.

Ranunculaceee, 91, 239.

Scypha, 71.

Radicle, 66.
Rafflesiaceae, 173.

Ramentaceous,

9.

root, 19.

Scape, 18.
Scitamineae, 175.
Scleranthaceae, 130.

INDEX.

201

Secretions of plants, 30.


Secundine, 54.
Seed, 63.

Stellate cellular tissue, 3.


Stem, 11.

Semiterete, 67.

endogenous, 12.
exogenous, 12, 13.
Sterculiaceae, 249.

Semivascular plants, 9 1 .
Sepals, 41.

acrogenous, 12.

Separated, 230.
Septate, 6.
Septicidal, 58.

Stigma, 49.

Serrate, 29.
Seta, 70.

Stomates, 7,
Stroma, 73.

Setaceous, 60.
Seven-lobed, 28.

Strophiolae, 64.
Structural Botany,

Sexes, 35.

Sheathing, 25.

Struma, 25, 69.


Strumose, 46.

Shields, 71.
Silenacese, 253.

Style, 49.

Silenales, 206.

Stylidiaceae, 142.

Silenese, 104.

Styraceas, 144, 266.


Succulent, 18.
Superior, 42, 54.

Stipes, 73.
Stipules, 24, 29.
8.

1.

Stupose, 46.

Siliqua, 61.
Silicula, 61, 63.
Simarubaceae, 116, 253.

Simple, 40, 59.


Sinuated, 28.
Sinuous cellular, 3.
Ske*tch of a new distribution of the
Vegetable Kingdom, 226.
Smilaceae, 180, 281.
Solanaceae, 151, 274.
Solanales, 218.

Supervolutive, 37.
Suspensor, 66.
Suture, 58.

Syncarpous, 50.
Syngenesious, 47.
System, Natural, 85.
natural, of De Candolle, 90.
Linnaean Sexual, 76.

Systematical Botany, 75.

Solitary, 37.

and

axillary, 38.

Soredia, 72.
Son, 69.

Tamaricaceae, 101, 253.


Taper, 25.

Taxaceae, 170, 278.

Tela cellulosa, 2.
Tendril, 25, 40.

Spadicosae, 219.
Spadix, 38.

Spathe, 35.

Terete, 20.

Spermoderm, 64.

Terminal, 37.
Ternate, 28.

Sphaerenchyma,

3.

Spheroidal, 67.
cellular tissue, 3.

Spike, 38.
Spine, 19.
Spinescent, 25.
Spiny, 42.
Spirseeae, 122.
Spiral, 60.
vessels, 5.

Spondiaceae, 251.

Spongelets, 10.
Spongioles, 10.
Sporangia, 69, 70, 71, 72, 73.
Sporidia, 72.

Spofogens, 230.
Spojes, 69.
Spurious dissepiments, 53.
Stamens, 45.
Staminidia, 71.
Staphyleaceae, 117.
Stellales, 215.
Stellatae, 141, 268.
Stellate, 9, 60.

Ternstromiaceae, 106, 248.


Testa, 64.
Testiculate, 11.

Tetradynamous, 46.
Thalamiflorse, 91.
Thallodes, 72.

Thallogens, 230.
Thallus, 71, 73.

Theales, 203.
Theca, 69, 70.
Thecaphore, 50.

Thymelaceae, 161, 262.


Thyrsus, 39.
Tiliacese, 105, 250.
Tissu cellulaire, 2.
Tissue, 2.
cellular, 2.

laticiferous, 6.

muriform

cellular, 16.

pitted, 4.

vascular, 5.

woody,
'

Toothed, 46.

4,

3.

INDEX.

292

Vague, 69.

Toothed at the apex, GO.

Toms,

Valerianaceae, 138, 270.


Valvate, 37.
and involute, 37.

49.

Trachenchyma,

5.

Transverse, 58, 66.

Valves, 47, 58.

Transversely articulate, 25.


Triangular, 20, 27.

Vasa

contracta, 6.

expansa, 6.
Vascular, 2.

Trica, 71.

Tridentate, 29.
Trilateral, 20.
Tripinnate, 28.
Triquetrous, 20.

tissue, 5.

Vasiform, 4.
Veins, 25.

Velum, 73.

Tritemate, 28.
Truncate, 29, 60.

Ventral suture, 58.


Verbenacese, 153, 273.
Vermicular, 66.
Vernation, 30.

Tube, 41.
Tubercular, 11.

Tuberculum, 71.
Tuberous, 11.

Verticillate, 23.

Tubiferosae, 209.

Vesicate, 42.
Vesiculae, 72.

Tubular, cellular tissue, 3.


Turneracese, 130.
Two-edged, 20.
Two-lipped, 43.

Vessels, proper, 6.
Vexillary, 37.
Vitaceee, 113,245.
Vitellus, 66.

Typhacese, 187.
Typhales, 222.

Violacese, 102, 247.


Violales, 202.

Ulmacese, 260.
Ulmales, 210.

Volvales, 214.
Vrille, 25.

Tubers, 19.

Volva, 73.

Umbel,

38.

Umbellales, 199.
Umbelliferse, 134, 242.
Umbilicus, 64.
Unilateral, 66.
Unguiculate, 43.

Wedge-shaped, 27.

Winged, 25.
Winteracese, 95, 242.

Wood, 14.
Woody, 2.

Unguis, 43.

tissue, 16.

Unsymmetrical, 43.
Urn, 70.
Urticaceae, 260.
Urticales, 209.

Xanthoxylaceas, 116, 254.


Xyridacese, 283.

Urticeae, 167.
Utricle, 60.

Zellengewebe, 2.
Zingiberaceae, 175, 278.

Zygophyllaceae, 116, 254,

Vaccinacese, 143, 266.

THE END.

LONDON

PRINTED BY SAMUEL BENTLEY,


Bangor House, Shoe Lane.

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