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Differential Equations
Laurenz Wiskott
Institut fur Neuroinformatik
Ruhr-Universitat Bochum, Germany, EU
4 February 2017
Contents
Teaching/Material/, where you can also find other teaching material such as programming exercises. The table of contents of
the lecture notes is reproduced here to give an orientation when the exercises can be reasonably solved. For best learning effect
I recommend to first seriously try to solve the exercises yourself before looking into the solutions.
1
2.6 Nonhermitean systems (complex eigenvalues) . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
2.7.5 Exercise: System of linear differential equations with too few eigenvectors . . . . . . . 5
Classify the following differential equations. Determine their order and decide whether they are ordinary or
partial, linear, homogeneous, and/or autonomous.
(a) y = y . (1)
Classify the following differential equations. Determine their order and decide whether they are ordinary or
partial, linear, homogeneous, and/or autonomous.
2
(a)
y + 4 = 0 (1)
(b)
(1 + y 2 )y + ty + y = et (2)
(c)
g(x, y) g(x, y)
2 = 0 (3)
x y
(d)
dy(x)
= x. (4)
dx
Show that the given functions yi are indeed solutions to the given differential equations.
Write the following higher-order differential equations in standard form, i.e. as a system of first-order differ-
ential equations with exactly one derivative without prefactor per equation.
(a) y y 6 y = 0 . (1)
(b) d3 y d2 y 1 dy
+ 3 +2 = 0. (2)
dt3 dt2 2 dt
Write the following higher-order differential equations in standard form, i.e. as a system of first-order differ-
ential equations with exactly one derivative without prefactor per equation.
...
(a) 3 z 2z + z = 3
1
(b) 2 x 3y = x + y
6y + x = 2x 1
x + 2y = x + 2
(c)
2y + 4x = 2x + y
3
2.3 Superposition principle
y = y + cos(t) (1)
with positive .
Hint: At some point it may help to make the ansatz y(t) = s sin(t) + c cos(t).
(b) Discuss how the solution depends on .
Find a differential equation for which the scaled version y(t) would be a solution.
Find the solutions of the following system of linear differential equations and sketch their trajectories.
x = 3x 2y (1)
y = 2x 2y . (2)
Find the solution of the following system of linear differential equations and sketch its trajectories.
x = +x 4y (1)
y = x + y . (2)
4
2.7.5 Exercise: System of linear differential equations with too few eigenvectors
Find the general solution of the following system of linear differential equations and sketch its trajectories.
x 3x 2y = 0 (1)
y y + z = 1 (2)
z z = 0 . (3)
(a) Write the ODE in standard form and find its general solution.
(b) Discuss the qualitative behavior of the general solution as a function of > 0 on the basis of the
eigenvalues. Can you tell why it describes a damped oscillator?
Consider the metabolism of a pharmaceutical substance in the body. After oral intake the substance is
absorbed in the digestive tract and goes from there into the blood. From there it diffuses into the tissue and
is used or otherwise broken down (we dont care how this happens exactly).
The goal is to define a simple mathematical model for the time course of the amounts of substance in the
digestive tract, the blood, and the tissue. Make the following assumptions:
The substance in the digestive tract is absorbed into the blood with a rate of about 20% per hour.
The substance in the blood diffuses into the tissue with a rate of about 90% per hour.
The tissue uses or breaks down the substance with a rate of about 5% per hour.
(a) Define a system of linear differential equations that describes the dynamics of the amounts of substance
in the three different areas (digestive tract, blood, and tissue) and find the general solution.
(b) Assume that at a certain time point a certain amount has been taken in orally and that there has
been no intake ever before. Determine and sketch the time course of the amounts in the three different
areas.
(c) What happens if you give the substance as a continuous blood infusion? How much do you have to
give per hour to obtain a constant amount of 200mg in the tissue?
Consider again the model for the metabolism of a pharmaceutical substance we had recently.
(a) Describe in words what happens if over years a patient gets a pill every 48 hours. Find an estimation
for the amount of substance in the patients tissue in the long run.
Hint:
P Use the previous results from the model, the superposition principle, and the geometric series
k x
k=1 x = 1x .
5
(b) Reduce the current 3-dimensional model of this metabolic process to a 2-dimensional model. Perform
the reduction such that the predictions of the 2-dimensional model (measured by the amount of the
substance in the tissue) are as faithfull as possible.
(c) Give an estimate for how much the amount of substance in the patients tissue is fluctuating in the 48
hours between the two pills?
First calculate the fixpoints. Then linearize the differential equations around the fixpoints which are not
equal to zero.
x + x + x3 x = 0 mit > 0
Bring it into standard form, find the fixpoints depending on the parameter and investigate their stability.
x = px (1)
2
y = 0.1(x y) , (2)
(a) Sketch the nullclines and determine the only fixed point.
6
3.3.4 Exercise: Qualitative analysis the of a nonlinear ODE in 2D - FitzHugh-Nagumo model
Consider the following FitzHugh-Nagumo model of a neuron, also known as Bonhoeffer-van der Pol oscillator:
V = V V 3 /3 W + I (1)
W = 0.08(V + 0.7 0.8W ) . (2)
V denotes the membrane potential, W is a recovery variable, and I is the stimulus current. Analyze the
system as follows: