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NOMINA INSECTA NEARCTICA 551

ODONATA
ROSSER W. GARRISON
RESEARCH ASSOCIATE
NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY
900 EXPOSITION BOULEVARD
LOS ANGELES, CALIFORNIA 90007
552 NOMINA INSECTA NEARCTICA
NOMINA INSECTA NEARCTICA 553

1991 and 1 February 1993, respectively. The first


INTRODUCTION edition was reviewed by Garrison in Argia: 1991, Vol.
3 No. 1: 23-24. The second and third editions
incorporated corrections and additions brought to the
This list includes all described Odonata of North author's attention. The third edition has a new section,
America north of the Mexican border. Species Part XIII, which includes figures of wings of almost all
occurring in the Antilles and Bahamas are considered to genera of Odonata.]
be Neotropical and are not included. Species
misidentifications are not included. I have used the This list generally reflects current taxonomic views. I
following sources: have listed all subgenera as synonyms although several
are considered as valid subgenera or even genera by
Kirby, W.F. 1890. A synonymic catalogue of some workers. I have sent several versions of the list to
Neuroptera Odonata, or dragonflies. Gurney and the following Odonatists, who have freely given their
Jackson, London, ix + 202 pp. advice, opinions, and constructive criticisms: G. Bick
(Florida), C. Cook (Kentucky), J. Daigle (Florida), J.
Muttkowski, R.A. 1910. Catalogue of the Odonata.
De Marmels (Venezuela), T. Donnelly (New York), S.
of North America. Bull. Publ. Mus. Milwaukee, l(l):1-
Dunkle (Florida), K. Tennessen (Alabama), S. Tsuda
207.
(Japan).
Paulson, D.R. 1969. The Odonata of North and
Problems have confronted me for correct dates and
Middle America. (Unpublished list).
authorship of various taxa. The Hagen/Se1ys
Paulson, D.R. 1977. Odonata in Biota Acuatica de authorship controversy has been particularly
Sudamerica Austral, S.H. Hurlbert, ed., San Diego troublesome; and I have decided on status by examining
State University, 170-184. each original description by these two authors. Philippe
Machet, of Saint-Cloud, France, has been particularly
Paulson, D.R. 1982. Odonata in Aquatic Biota of
helpful in providing correct dates for certain poorly
Mexico, Central America and the West Indies. H.H.
known published taxa described by Blanchard and
Hurlbert and A. Villalobos-Figueroa, eds. San Diego
others.
State Univ., San Diego.
I owe much appreciation to the late Mr. Charles
Paulson, D.R. 1983. Working list of the Odonata of
Bridges, of the University of Illinois at Urbana-
South America. (Unpublished list). Champaign, who corresponded with me over the years
Davies, D.A.L. and R Tobin. 1984. The dragonflies of regarding the status of many New World Odonata name
the world: a systematic list of the extant species of changes; he provided me with much help during the
Odonata. Vol. 1. Zygoptera, Anisozygoptera. evolution of this list. To Don Frack of Covina,
Societas Internationalis Odonatologica Rapid Comm. California, I owe a special debt. He provided me with a
(Suppl.) 3. DBASE program, TAXON, which made the present list
possible. My wife, Jo, who has always helped me over
Davies, D.A.L. and P. Tobin. 1985. The dragonflies the years, helped with the data input.
of the world: a systematic list of the extant species of
Odonata. Vol. 2. Anisoptera. Societas Internationalis
Odonatologica Rapid Comm. (Suppl.) 5.
Tsuda, Shigeru. 1986. A distributional list of world
Odonata. Preliminary edition. Privately published.
Osaka.
Tsuda, Shigeru. 1991. A distributional list of world
Odonata. Privately published. Osaka. [Note: This is
the revised edition of the 1986 work cited above; a
review of this work can be found in Argia: 1991, Vol. 3
No. 1: 23-24.]
Bridges, Charles A. 1994. Catalogue of the family-
group, genus-group and species group names of the
Odonata of the world (third edition). Privately
published by the author (now deceased) [Note: The first
and second editions of this work appeared on 15 June

ODONATA: INTRODUCTION
554 NOMINA INSECTA NEARCTICA

ALTERNATIVE FAMILY NAMES


The purpose of this section is to list some alternative
family names the user might encounter in the Odonata.
This list is not an exhaustive list of family group names,
but is only included for the convenience of the users of
the check list. The abbreviation (not NA) stands for not
North American, i.e. the family is not found in the
Nearctic region north of the Mexican border.
Aeshnidae
Aeschnidae (see Aeshnidae)
Agrionidae (see Calopterygidae)
(Amphipterygidae not NA)
Argiidae (see Calopterygidae)
(Austropetaliidae not NA)
Calopterygidae
(Chlorocyphidae not NA)
Coenagrionidae
Cordulegasteridae (see Cordulegastridae)
Cordulegastridae
Corduliidae
(Dicteriadidae not NA)
(Epiophlebiidae not NA)
(Euphaeidae not NA)
Gomphidae
(Hemiphlebiidae not NA)
(Isostictidae not NA)
Lestidae
(Lestoideidae not NA)
Libellulidae
Macrodiplactidae (see Libellulidae)
(Megapodagrionidae not NA)
(Neopetaliidae not NA)
(Perilestidae not NA)
Petaluridae
(Platycnemididae not NA)
Platystictidae
(Polythoridae not NA)
Protoneuridae
(Pseudolestidae not NA)
(Pseudostigmatidae not NA)
Pueliidae (see Coenagrionide)
(Synlestidae not NA)

ODONATA: ALTERNATIVE FAMILY NAMES


NOMINA INSECTA NEARCTICA 555

STATISTICS

This section contains some raw statistics without comment. A list of the species group
names proposed by decade is listed in the table below followed by a graph of the
numbers. Allowance must be made in interpreting the graph for the truncated decades at
the beginning and end of the graph. The tables on the following page are a break down of
the number of valid species and names by family.

Species group names (valid and synonymous) described by decade.

1758 - 1759 3
1760 - 1769 1
1770 - 1779 17
1780 - 1789 1
1790 - 1799 6
1800 - 1809 2
1810 - 1819 0
1820 - 1829 0
1830 - 1839 71
1840 - 1849 53
1850 - 1859 57
1860 - 1869 153
1870 - 1879 72
1880 - 1889 24
1890 - 1899 56
1900 - 1909 56
1910 - 1919 53
1920 - 1929 19
1930 - 1939 18
1940 - 1949 17
1950 - 1959 16
1960 - 1969 6
1970 - 1979 3
1980 - 1989 13
1990 - 10

160
140
120
100
80
60
40
20
0
1758- 1780- 1810- 1840- 1870- 1900- 1930- 1960- 1990
1789 1819 1849 1879 1909 1939 1969 -

ODONATA: STATISTICS
556 NOMINA INSECTA NEARCTICA

Family # Names # Valid


Aeshnidae 63 38
Calopterygidae 33 8
Cordulegastridae 11 8
Corduliidae 93 58
Gomphidae 138 97
Lestidae 23 17
Libellulidae 216 108
Petaluridae 2 2
Platystictidae 1 1
Protoneuridae 2 2
Total 727 435

ODONATA: STATISTICS
GarrisonNew World Odonata List (NWOL)

CLASSIFICATION: ODONATA
The classification adopted here, with the exception of the Gomphidae, is largely traditional and is based primarily
on: Fraser (1957. A reclassification of the Order Odonata. Royal Zoological Society of New South Wales,
133pp). The classification of the Gomphidae follows: Carle (1986. The classification, phylogeny and biogeography
of the Gomphidae (Anisoptera). I. Classification. Odonatologica 15(3): 275-326). Parts of the classification are
unsatisfactory and phylogenetic studies will alter the classification considerably. Such a new classification has
recently been proposed by Bechly (1996. Morphologische Untersuchungen am Flgelgeder der rezenten Libeeen
und deren Stammgruppenvertreter (insecta; Pterygota; Odonata). Petalura, spec.-vol. 2: 402 pp., 111 figs., 3 tabs.
[revised second edition with an English appendix: "Phylogenetic System and List of Synapomorphies"]). Drastic
changes in the higher classification are advocated. An abstracted English version (The phylogenetic system of fossil
and recent Odonatoptera).is available at the following web site:
http://members.aol.com/odonatadat/phylogeny/bechly.htm

SUBORDER ZYGOPTERA Selys, 1854

CALOPTERYGIDAE Selys, 1850


Calopteryginae Selys, 1850: Calopteryx.
Hetaerininae Selys, 1853: Hetaerina.
LESTIDAE Calvert, 1901
Lestinae Calvert, 1901: Archilestes, Lestes.
PLATYSTICTIDAE Tillyard & Fraser, 1938
Palaemnematinae Tillyard & Fraser, 1938: Palaemnema.
PROTONEURIDAE Tillyard, 1917
Protoneurinae Tillyard, 1917: Neoneura, Protoneura.
COENAGRIONIDAE Kirby, 1890
Argiinae Tillyard, 1917: Argia.
Coenagrioninae Kirby, 1890: Chromagrion, Coenagrion, Nehalennia, Neoerythromma.
Ischnurinae Fraser, 1957: Acanthagrion, Amphiagrion, Enallagma, Hesperagrion, Ischnura, Zoniagrion.
Pseudagrioninae Tillyard, 1917: Apanisagrion, Telebasis.

SUBORDER ANISOPTERA Selys, 1854

PETALURIDAE Tillyard, 1917


Petalurinae Tillyard, 1917: Tachopteryx.
Tanypteryginae Davies & Tobin, 1985: Tanypteryx.
AESHNIDAE Rambur, 1842
Aeshninae Rambur, 1842: Aeshna, Anax Coryphaeschna, Gynacantha, Remartinia, Triacanthagyna.
Brachytroninae Tillyard, 1917: Basiaeschna, Boyeria, Epiaeschna, Gomphaeschna, Nasiaeschna,
Oplonaeschna.
GOMPHIDAE Rambur, 1842
Hageniinae Davies & Tobin, 1985: Hagenius.
Octogomphinae Carle & Cook, 1984: Lanthus, Octogomphus, Stylogomphus.
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GarrisonNew World Odonata List (NWOL)

Gomphinae Rambur, 1842: Arigomphus, Dromogomphus, Gomphus, Stylurus.


Onychogomphinae Chao, 1984: Erpetogomphus, Ophiogomphus.
Lindeniinae Selys, 1854: Aphylla, Phyllogomphoides, Progomphus.
CORDULEGASTRIDAE Tillyard, 1917
Cordulegastrinae Tillyard, 1917: Cordulegaster.
CORDULIIDAE Kirby, 1890
Macromiinae Tillyard, 1917: Didymops, Macromia.
Corduliinae Kirby, 1890: Cordulia, Dorocordulia, Epitheca, Helocordulia, Neurocordulia, Somatochlora,
Williamsonia.
LIBELLULIDAE Rambur, 1842
Brachydiplactinae Tillyard, 1917: Micrathyria, Nannothemis.
Leucorrhiniinae Tillyard, 1917: Brachymesia,, Celithemis, Leucorrhinia.
Libellulinae Rambur, 1842: Cannaphila, Libellula, Orthemis.
Sympetrinae Tillyard, 1917: Crocothemis, Erythemis, Erythrodiplax, Pachydiplax, Pseudoleon,
Sympetrum.
Trithemistinae Tillyard, 1917: Brechmorhoga, Dythemis, Macrothemis, Paltothemis.
Palpopleurinae Tillyard, 1917: Perithemis.
Trameinae Tillyard, 1917: Idiataphe, Miathyria, Pantala, Tauriphila Tholymis, Tramea.
Urothemistinae Lieftinck, 1954: Macrodiplax.

Annotations and comments on the Check List


General Comments:

1. Date of publication of Hagen and Se1ys, Monographie des Gomphines: I follow Calvert (1905. Biologia Centrali
Americana: 148), who states in a footnote: "It is usual to quote the "Monographie des Gomphines" with the date
1857, as is done by Kirby [1890] for example, and "November 1857" is, indeed, that which appears on the title
page of the volume; but the original paper cover bore the date "Mai 1858," which has usually not been preserved in
bound copies. Moreover, plates xx-xxii are dated at the bottom as having been drawn in 1858." Cowley (1937. J.
Soc. Bibliography Nat. Hist. 1(3):73-74) opted for 1857, but further evidence of an 1858 publication date was
brought to my attention by P. Machet. In the "appendice" (page 651 [391 separate]) is stated: "Some
circumstances having put off the conclusions of this volume, considerable and rather numerous materials have
reached us during printing [italics mine]. One comes from a trip made by H. Hagen to London in 1857; others
from a shipment of species from Texas, made by the Smithsonian Institution."
2. I give synonyms, nomina nuda, etc., only for New World taxa for Holarctic species such as Enallagma
cyathigerum and Libellula quadrimaculata. The single exception is Crocothemis servilia servilia, an Old World
species recently introduced into peninsular Florida.

Specific Comments:

1. Aeshna clepsydra with synonyms arundinacea Se1ys 1872 and maxima Heikal in Hisinger 1861. The latter
two names have often been included as synonyms of A. clepsydra. My reason for their deletion is based on a
footnote by Walker to Muttkowski (1910. Cat. Odon.: 109):
"Walker in litteris Aug. 14, 1909: 'The synonyms given by you do not belong to the true clepsydra. A.
arundinacea and maxima I have not seen: It is very probable that the first at least is a good species."' And,

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GarrisonNew World Odonata List (NWOL)

"Walker in litteris, Sept. 24, 1909: 'Some of the references ... do not refer to true clepsydra...I am quite sure
arundinacea is not clepsydra, but as I have not seen the type it is probably best to let it stand as a synonym of
the latter on the authority of Martin'."
Curiously, I can find no mention of these little-used names in Walker's 1912 monograph. The original
description of arundinacea is based on a female from "Irkutzk", and Se1ys states it is similar to Siberian A.
juncea. According to Dr. Matti Hmlinen, University of Helsinki, in a letter to me, A. maxima is a junior
synonym of the Old World Aeshna crenata Hagen. I have found no records of clepsydra occurring anywhere in
the Palearctic, and I strongly suspect that arundinacea refers to a species other than the North American
clepsydra.
2. Triacanthagyna obscuripennis Blanchard. Williamson, in his monograph of this genus (1923. Univ. Mich.
Mus. Zool. Misc. Publ. 9:1-80), believed this to be a possible synonym of T. septima; but P. Machet believes,
after examining the short description and figure by Blanchard, that it may be of the trifida group, and possibly a
synonym of one of the later described species. Should this be the case (if the type is found), I would not be in
favor of using this name, I list obscuripennis as a distinct species, but it should, in my opinion, be considered a
nomen dubium or a nomen obletum, if and when its status is determined.
3. Gomphus complex. The status of the various genera/subgenera of North American entities has not been
satisfactorily resolved. I have not studied the problems, but have largely followed the advice of others. In this
list, I do not recognize the subgenus Stenogomphurus nor do I accept elevation of the subgenus Gomphurus to
generic status since there appears to be questions of their validity and changes in the literature as follows: Carle
(Carle, F. L. 1986. The classification, phylogeny and biogeography of the Gomphidae (Anisoptera). I.
Classification. Odonatologica 15(3): 275-326.) proposed the subgenus Stenogomphurus (type species:
Gomphus consanguis) under the large genus Gomphus. However, G. consanguis has always been included
under the subgenus Gomphurus of Gomphus. Carle (1986) also raises Gomphurus to generic status, but does
not key or diagnose between Gomphus and Gomphurus. He does not explain his reason for removing G.
consanguis from Gomphus and placing it in the new subgenus, Stenogomphurus. It would appear that he
never diagnosed Stenogomphurus from Gomphus. Carle (1986) does not discuss the relationship between
Gomphurus and Gomphus sensu lato, and gives no reason for the elevation of Gomphurus to generic status.
Therefore, I will continue to treat Gomphurus as a subgenus of Gomphus. Phanagomphus is also not
recognized in this list for the following reason: Carle (Carle, F. L. 1986. The classification, phylogeny and
biogeography of the Gomphidae (Anisoptera). I. Classification. Odonatologica 15(3): 275-326.) proposed the
subgenus Phanogomphus (type species: Gomphus minutus Rambur), which Carle & May (Carle, F. L. & M. L.
May. 1987. Gomphus (Phanogomphus) westfalli spec. nov. from the gulf coast of Florida (Anisoptera:
Gomphidae). Odonatologica 16(1): 67-75) later expanded to include other species of Gomphus (Gomphus).
Many Odonatists, including myself, consider Phanogomphus a subjective synonym of the type species of
Gomphus, G. vulgatissimus (Linnaeus). I treat Arigomphus as a separate genus from Gomphus.
Hylogomphus is considered a synonym of Gomphus (Gomphus), following Carle (1986). I realize that this
arrangement will probably not be satisfactory to many, but a final resolution may not be achieved for several
years.
Carle (in. May, M and F. Carle, 1996. An annotated list of the Odonata of New Jersey, with an appendix on
nomenclature in the genus Gomphus. Bulletin of Odonatology 4(1):1-35) has defended his actions regarding the
introduction of the names discussed above. That such an appendix was written and the acknowledgement that
several workers were in doubt about the proper introduction of new taxa and changes illustrates, in my opinion,
the continued confusion by this author's actions.
4. CORDULEGASTRIDAE: In a recent paper, 1992. Revision der Cordulegastridae. 1. Entwurfeiner neuen
Klassifizierung der Familie (Odonata: Anisoptera). Opuscula Zoologica Fluminensia 96:1-18, H. Lohmann has
performed a cladistic analysis on the family, with the result that our eight species are allocated to six genera as
follows:

Archegaster (gen. nov.)


sayi

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GarrisonNew World Odonata List (NWOL)

Kalyptogaster (gen. nov.)


erronea (with synonym fasciata)
Lauragaster (gen. nov.)
diadema
dorsalis
Pangaegaster (gen. nov.)
maculata
Taeniogaster (Selys, 1854)
obliquua
Zoraena Kirby 1890
bilineata
diastatops

While I admire the treatment of this family by Lohmann, I do not think it serves science to split our American
species from one into six genera. Carle, in an earlier paper (1983. Ann. Entomol. Soc. Amer. 76(1):61-68) also
recognized Zoraena and Taeniogaster, but his divisions do not seem to have been followed by Odonatists in this
country. I still consider all of our species in one genus, Cordulegaster. However, I include the above data for
those who wish to make changes.
5. Both Dr. Ken Tennessen and Mr. Carl Cook inform me that Neurocordulia clara Muttkowski is more closely
related to N. molesta, although Cook believes N. clara a good species. I have treated it as a synonym of N.
molesta.
6. Ken Tennessen (in litt.) informs me that he believes Epitheca (Tetragoneuria) williamsoni Muttkowski to be a
probable synonym of E. costalis.
7. Erythrodiplax connata complex. I include some nomenclatoral changes which are not in accord with Borror
1942 (Ohio State Univ. Contr. Zool. Entomol., No. 4). Other Odonatists and I treat E. minuscula and E. fusca
as separate species based on an absence of intergrades with E. connata (despite the fact that Borror reports such
specimens in his monograph)..
8. Libellula bifasciata Blanchard. The type for this old, forgotten name was discovered by Lieftinck to be L.
semifasciata, not L. pulchella, according to Dr. M.J. Westfall, Jr. (pers. comm.). As with Triacanthagyna
obscuripennis, I think it best to consider L. bifasciata a nomen obletum.

Status of specific names not mentioned in this list:


1. Enallagma optimolocus Miller & Ivae, 1995 (Proceedings of the Entomological Society of Washington
97(4):833-838). I reviewed an earlier draft of the manuscript, have one paratype in my collection and also
examined a specimen supposedly of this species from Oregon (Crook County, South Fork Crooked River, nr.
confluence with North Fork, 10 Aug. 1994, Ken J. Tennessen, collector). I strongly believe that this taxon is only
a hybrid between E. anna and E. carunculatum for the following reasons: No females of E. optimolocus have
been collected and this "species" has always been collected where the two parental types occur. Examination of
two paratypes and the Oregon male, the last kindly loaned to me by Dr. Ken Tennessen, show appendage
morphology intermediate between the parental types. Hybridization between other Enallagma species is known
and hybrids intermediate in appendage morphology have been produced in the laboratory and occur in nature for
two closely related species, Ischnura denticollis and I. gemina (Leong, J. M. & J. E. Hafernick, Jr. 1992.
Hybridization between two damselfly species (Odonata: Coenagrionidae): morphometric and genitalic
differentiation. Annals of the Entomological Society of America 85(6): 662-670).
2. Gomphus williamsoni Muttkowski, 1910. This name based on only one known specimen, is almost certainly a
natural hybrid between Gomphus (Gomphus) graslinellus and Gomphus (Gomphus) lividus.
3. Macromia wabashensis Williamson, 1909. Until recently, consensus among most U. S. Odonatists was to treat
Macromia wabashensis as a synonym or color form (polymorph) of M. taeniolata. Secondary genitalia
including the shape of the head of the hamule for M. pacifica, M. wabashensis, and M. taeniolata is similar.

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GarrisonNew World Odonata List (NWOL)

Williamson (1909 Proc. U. S. Nat. Museum 37:369-398.) reports captures of M. wabashensis with Macromia
taeniolata and M. pacifica, which suggests the possibility that M. wabashensis is a hybrid between M. pacifica
and M. taeniolata. Other mostly non-published data suggest the occurrence of both M. pacifica and M.
taeniolata wherever M. wabashensis has been collected. Like Dunkle (1995. Conservation of dragonflies
(Odonata) and their habitats in North America. page 23 in Corbet, P. S., S. W. Dunkle, H. Ubukata, eds.
Proceedings of the International symposium on the conservation of dragonflies and their habitats. Japanese
Society for the Preservation of Birds, Kushiro, xii + 1-70), I am doubtful of according species status to M.
wabashensis. Acquisition of more specimens and the discovery of the female may help to determine its true
status. I suggest that multivariate morphometric analyses of available specimens of M. wabashensis and samples
of M. taeniolata and M. pacifica from zones of sympatry with M. wabashensis may provide evidence about
whether M. wabashensis is a hybrid.

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562 NOMINA INSECTA NEARCTICA

AESHNIDAE Coryphaeschna Williamson 1903

Coryphaeschna adnexa Hagen 1861 (Aeschna)


Aeschna cyanifrons Hagen 1861 Unav.
Aeshna Fabricius 1775 Aeschna macromia Brauer 1865 Syn.
Cnemophila Wallengren 1894 Syn. Coryphaeschna guyanensis Machet 1991 Syn.
Rhionaeschna Frster 1909 Syn. Coryphaeschna ingens Rambur 1842 (Aeschna)
Neureclipa Navas 1911 Syn. Aeschna abboti Hagen 1863 Syn.
Hesperaeschna Cockerell 1913 Syn. Coryphaeschna viriditas Calvert 1952 (Coryphaeschna)
Secundaeschna Gtz 1923 Syn. Aeschna virens Rambur 1842 Homo.
Marmaraeschna Calvert 1952 Syn.
Schizuraeschna Calvert 1952 Syn. Epiaeschna Hagen 1877
Adversaeschna Watson 1992 Syn.
Epiaeschna heros Fabricius 1798 (Aeschna)
Aeshna californica Calvert 1895 (Aeshna) Aeschna multicincta Say 1839 Syn.
Aeshna canadensis Walker 1908 (Aeshna)
Anax maritimum Provancher 1895 Nomob. Gomphaeschna Selys 1871
Aeshna clepsydra Say 1839 (Aeshna)
Aeshna constricta Say 1839 (Aeshna) Gomphaeschna antilope Hagen 1874 (Aeschna)
Aeshna dugesi Calvert 1905 (Aeshna) Gomphaeschna furcillata Say 1839 (Aeschna)
Aeshna eremita Scudder 1866 (Aeshna) Gynacantha quadrifida Rambur 1842 Syn.
Aeshna hudsonica Selys 1875 Syn.
Aeshna interrupta Walker 1908 (Aeshna) Gynacantha Rambur 1842
Aeshna interna Walker 1908 Syn. (subspecies) Acanthagyna Kirby 1890 Syn.
Aeshna lineata Walker 1908 Syn. (subspecies) Selsiophlebia Frster 1904 Syn.
Aeshna nevadensis Walker 1908 Syn.
Aeshna juncea Linnaeus 1758 (Libellula) Gynacantha nervosa Rambur 1842 (Gynacantha)
Aeshna americana Bartenef 1929 Syn.
Aeshna multicolor Hagen 1861 (Aeshna) Nasiaeschna Selys 1900
Aeshna furcifera Karsch 1891 Syn.
Aeshna mutata Hagen 1861 (Aeshna) Nasiaeschna pentacantha Rambur 1842 (Aeschna)
Aeshna palmata Hagen 1856 (Aeshna)
Aeshna arida Kennedy 1918 Syn. Oplonaeschna Selys 1883
Aeshna persephone Donnelly 1961 (Aeshna) Hoplonaeschna Karsch 1891 Syn.
Aeshna psilus Calvert 1947 (Aeshna)
Aeshna dominicana Hagen 1861 Unav. Oplonaeschna armata Hagen 1861 (Aeschna)
Aeshna septentrionalis Burmeister 1839 (Aeshna) Oplonaeschna polernista Navas 1917 Syn.
Aeshna sitchensis Hagen 1861 (Aeshna)
Aeshna subarctica Walker 1908 (Aeshna) Remartinia Navas 1911
Aeshna tuberculifera Walker 1908 (Aeshna)
Aeshna umbrosa Walker 1908 (Aeshna) Remartinia luteipennis Burmeister 1839 (Aeschna)
Aeshna occidentalis Walker 1912 Syn. (subspecies) Aeschna florida Hagen 1861 Syn. (subspecies)
Aeshna verticalis Hagen 1861 (Aeshna)
Aeshna propinqua Scudder 1866 Syn. Triacanthagyna Selys 1883
Aeshna walkeri Kennedy 1917 (Aeshna)
Triacanthagyna septima Selys 1857 (Gynacantha)
Anax Leach 1815

Anax amazili Burmeister 1839 (Aeschna)


Anax maculatus Rambur 1842 Syn.
Anax junia Drury 1770 (Libellula) CALOPTERYGIDAE
Anax spiniferus Rambur 1842 Syn.
Anax longipes Hagen 1861 (Anax)
Anax walsinghami McLachlan 1883 (Anax)
Anax validus Hagen 1875 Unav. Calopteryx Leach 1815
Agrion Fabricius 1775 Syn. (?)
Basiaeschna Selys 1883 Sylphus Hagen 1853 Syn.
Anaciagrion Kennedy 1920 Syn.
Basiaeschna janata Say 1839 (Aeschna)
Aeschna minor Rambur 1842 Syn. Calopteryx aequabilis Say 1839 (Calopteryx)
Calopteryx virginica Selys 1853 Syn.
Boyeria McLachlan 1896 Calopteryx hudsonica Hagen 1875 Syn.
Fonscolombia Selys 1883 Homo. Calopteryx yakima Hagen 1889 Syn.
Calopteryx coloradica Cockerell 1913 Syn.
Boyeria grafiana Williamson 1907 (Boyeria) Calopteryx californica Kennedy 1917 Syn.
Boyeria vinosa Say 1839 (Aeschna)
Aeschna quadriguttata Burmeister 1839 Syn.

ODONATA: AESHNIDAE, CALOPTERYGIDAE


NOMINA INSECTA NEARCTICA 563

Calopteryx amata Hagen 1889 (Calopteryx) Argia bipustulata Kirby 1890 Missp.
Calopteryx angustipennis Selys 1853 (Sylphis) Argia cupreum Hagen 1861 (Agrion)
Sylphis elegans Hagen 1853 Syn. Argia emma Kennedy 1915 (Argia)
Calopteryx dimidiata Burmeister 1839 (Calopteryx) Argia extraneum Hagen 1861 (Agrion)
Calopteryx apicalis Burmeister 1839 Syn. Argia mista Navas 1934 Syn.
Calopteryx cognata Rambur 1842 Syn. Argia fumipennis Burmeister 1839 (Agrion)
Calopteryx maculata Beauvois 1805 (Agrion) Argia obscura Rambur 1842 Syn.
Calopteryx holosericea Burmeister 1839 Syn. Agrion violaceum Hagen 1861 Syn. (subspecies)
Calopteryx materna Say 1839 Syn. Argia atra Gloyd 1968 Syn. (subspecies)
Calopteryx opaca Say 1839 Syn. Argia hinei Kennedy 1918 (Argia)
Calopteryx papilionaria Rambur 1842 Syn. Argia immundum Hagen 1861 (Agrion)
Calopteryx floridana Huggins 1927 Syn. Argia lacrimans Hagen 1861 (Agrion)
Argia leonorae Garrison 1994 (Argia)
Hetaerina Hagen 1853 Argia lugens Hagen 1861 (Hyponeura)
Argia moestum Hagen 1861 (Agrion)
Hetaerina americana Fabricius 1798 (Agrion) Agrion putridum Hagen 1861 Syn.
Lestes basalis Say 1839 Syn. Argia intruda Williamson 1912 Syn.
Hetaerina basalis Hagen 1859 Homo. Argia munda Calvert 1902 (Argia)
Hetaerina californica Hagen 1859 Syn. Argia rita Kennedy 1919 Syn.
Hetaerina pseudamericana Walsh 1863 Syn. Argia nahuana Calvert 1902 (Argia)
Hetaerina sclerata Walsh 1863 Syn. Argia saalasi Valle 1942 Syn.
Hetaerina texana Walsh 1863 Syn. Argia oenea Hagen 1865 (Argia)
Hetaerina titia Drury 1773 (Libellula) Argia pallens Calvert 1902 (Argia)
Calopteryx tricolor Burmeister 1839 Syn. Argia solita Kennedy 1918 Syn.
Hetaerina septentrionalis Selys 1853 Syn. Argia pima Garrison 1994 (Argia)
Hetaerina limbata Selys 1853 Syn. Argia plana Calvert 1902 (Argia)
Hetaerina rupamnensis Walsh 1863 Syn. Argia spegazzinii Navas 1934 Syn.
Hetaerina rupinsulensis Walsh 1863 Syn. Argia rhoadsi Calvert 1902 (Argia)
Hetaerina bipartita Selys 1873 Syn. Argia sabino Garrison 1994 (Argia)
Hetaerina vulnerata Hagen 1853 (Hetaerina) Argia sedulum Hagen 1861 (Agrion)
Argia tarascana Calvert 1902 (Argia)
Argia tezpi Calvert 1902 (Argia)
Argia tibialis Rambur 1842 (Platycnemis)
Agrion fontium Hagen 1861 Syn.
COENAGRIONIDAE Agrion binotatum Walsh 1862 Syn.
Argia tonto Calvert 1902 (Argia)
Argia translata Hagen 1865 (Argia)
Argia espinalensis Navas 1935 Syn.
Acanthagrion Selys 1876 Argia vivida Hagen 1865 (Argia)
Myagrion Frster 1914 Syn. Argia kurilis Hagen 1865 Syn.

Acanthagrion quadratum Selys 1876 (Acanthagrion) Chromagrion Needham 1903

Amphiagrion Selys 1876 Chromagrion conditum Selys 1876 (Erythromma)

Amphiagrion abbreviatum Selys 1876 (Pyrrhosoma) Coenagrion Kirby 1890


Amphiagrion saucium Burmeister 1839 (Agrion) Agrion Leach 1815 Homo.
Agrion discolor Burmeister 1839 Syn. Austrocoenagrion Kennedy 1920 Syn.
Amphiagrion amphion Selys 1876 Syn. Chloragrion Hartinov and Hartinova 1990 Syn.

Apanisagrion Kennedy 1920 Coenagrion angulatum Walker 1912 (Coenagrion)


Coenagrion interrogatum Selys 1876 (Agrion)
Apanisagrion lais Brauer 1876 (Nehalennia) Coenagrion resolutum Selys 1876 (Agrion)
Agrion servum Selys 1876 Syn.
Argia Rambur 1842 Enallagma walkeri Muttkowski 1911 Syn.
Hyponeura Selys 1854 Syn.
Diargia Calvert 1909 Syn. Enallagma Charpentier 1840
Chalcargia Kennedy 1920 Syn. Aenallagma Selys 1875 Syn.
Cyanargia Kennedy 1920 Syn. Thermagrion Frster 1906 Syn.
Heliargia Kennedy 1920 Syn. Africallagma Kennedy 1920 Syn.
Leptargia Kennedy 1920 Syn. Amphiallagma Kennedy 1920 Syn.
Micrargia Kennedy 1920 Syn. Ischnallagma Kennedy 1920 Syn.
Proischnura Kennedy 1920 Syn.
Argia agrioides Calvert 1895 (Argia) Teleallagma Kennedy 1920 Syn.
Argia alberta Kennedy 1918 (Argia) Sobobapteron Pierce 1965 Syn.
Argia apicale Say 1839 (Agrion)
Argia barretti Calvert 1902 (Argia)
Argia bipunctulatum Hagen 1861 (Agrion)

ODONATA: CALOPTERYGIDAE, COENAGRIONIDAE


564 NOMINA INSECTA NEARCTICA

Ischnura utahensis Muttkowski 1910 Syn.


Enallagma anna Williamson 1900 (Enallagma) Ischnura cervula Selys 1876 (Ischnura)
Enallagma culicinorum Byers 1927 Syn. Ischnura damula Calvert 1902 (Ischnura)
Enallagma antennatum Say 1839 (Agrion) Ischnura demorsum Hagen 1861 (Agrion)
Enallagma fischeri Kellicott 1895 Syn. Ischnura denticollis Burmeister 1839 (Agrion)
Enallagma aspersum Hagen 1861 (Agrion) Ischnura exstriata Calvert 1895 Syn.
Enallagma basidens Calvert 1902 (Enallagma) Ischnura erratica Calvert 1895 (Ischnura)
Enallagma boreale Selys 1875 (Enallagma) Ischnura gemina Kennedy 1917 (Celaenura)
Enallagma calverti Morse 1895 Syn. Ischnura hastata Say 1839 (Agrion)
Enallagma carunculatum Morse 1895 (Enallagma) Agrion anomalum Rambur 1842 Syn.
Enallagma civile Hagen 1861 (Agrion) Agrion venerinotata Haldeman 1844 Syn.
Enallagma plebium Selys 1876 Syn. Ischnura kellicotti Williamson 1898 (Ischnura)
Enallagma simile Selys 1876 Syn. Ischnura perparva Selys 1876 (Ischnura)
Enallagma clausum Morse 1895 (Enallagma) Ischnura positum Hagen 1861 (Agrion)
Enallagma coecum Hagen 1861 (Enallagma) Ischnura prognatum Hagen 1861 (Agrion)
Enallagma cardenium Hagen 1876 Syn. Ischnura ramburii Selys 1850 (Agrion)
Enallagma krugii Kolbe 1888 Syn. Ischnura tuberculata Selys 1857 Syn.
Enallagma concisum Williamson 1922 (Enallagma) Agrion aurantiaca Hagen 1861 Syn.
Enallagma cyathigerum Charpentier 1840 (Agrion) Agrion credulum Hagen 1861 Syn.
Agrion hastulatum Stephens 1835 Homo. Agrion defixum Hagen 1861 Syn.
Agrion annexum Hagen 1861 Syn. Agrion expertum Hagen 1861 Unav.
Enallagma robustum Selys 1875 Syn. Agrion iners Hagen 1861 Syn.
Enallagma vernale Gloyd 1943 Syn. (subspecies) Ischnura verticalis Say 1839 (Agrion)
Enallagma daeckii Calvert 1903 (Telagrion) Agrion canadense Provancher 1876 Syn.
Enallagma davisi Westfall 1943 (Enallagma)
Enallagma divagans Selys 1876 (Enallagma) Nehalennia Selys 1850
Enallagma doubledayi Selys 1850 (Agrion) Argiallagma Selys 1876 Unav.
Enallagma dubium Root 1924 (Enallagma) Argiallagma Calvert 1907 Syn.
Enallagma durum Hagen 1861 (Agrion)
Enallagma ebrium Hagen 1861 (Agrion) Nehalennia gracilis Morse 1895 (Nehalennia)
Enallagma exsulans Hagen 1861 (Agrion) Nehalennia integricollis Calvert 1913 (Nehalennia)
Enallagma geminatum Kellicott 1895 (Enallagma) Nehalennia irene Hagen 1861 (Agrion)
Enallagma piscinarium Williamson 1899 Syn. Nehalennia carolott Butler 1914 Syn.
Enallagma hageni Walsh 1863 (Agrion) Nehalennia pallidula Calvert 1913 (Nehalennia)
Enallagma laterale Morse 1895 (Enallagma)
Enallagma minusculum Morse 1895 (Enallagma) Neoerythromma Kennedy 1920
Enallagma novaehispaniae Calvert 1907 (Enallagma)
Enallagma ypsilon Needham 1942 Syn. Neoerythromma cultellatum Selys 1876 (Enallagma)
Enallagma pallidum Root 1923 (Enallagma)
Enallagma pictum Morse 1895 (Enallagma) Telebasis Selys 1865
Enallagma pollutum Hagen 1861 (Agrion) Erythragrion Selys 1876 Syn.
Enallagma praevarum Hagen 1861 (Agrion)
Enallagma recurvatum Davis 1913 (Enallagma) Telebasis byersi Westfall 1957 (Telebasis)
Enallagma semicirculare Selys 1876 (Enallagma) Telebasis salvum Hagen 1861 (Agrion)
Enallagma signatum Hagen 1861 (Agrion) Telebasis boucardi Selys 1868 Syn.
Agrion dentiferum Walsh 1863 Syn.
Enallagma sulcatum Williamson 1922 (Enallagma) Zoniagrion Kennedy 1917
Enallagma traviatum Selys 1876 (Enallagma)
Enallagma westfalli Donnelly 1964 Syn. (subspecies) Zoniagrion exclamationis Selys 1876 (Agrion)
Enallagma vesperum Calvert 1919 (Enallagma)
Enallagma laurenti Calvert 1919 Syn.
Enallagma weewa Byers 1927 (Enallagma)

Hesperagrion Calvert 1902


CORDULEGASTRIDAE
Hesperagrion heterodoxum Selys 1868 (Amphiagrion)
Agrion flavescens Selys 1868 Syn.
Cordulegaster Leach 1815
Ischnura Charpentier 1840 Taeniogaster Selys 1854 Syn.
Anomalagrion Selys 1857 Syn. Thecagaster Selys 1854 Syn.
Micronympha Kirby 1890 Syn. Thecaphora Charpentier 1840 Syn.
Celaenura Kennedy 1917 Syn. Thecaphora Selys 1854 Homo.
Anomalura Kennedy 1920 Syn. Zoraena Kirby 1890 Syn.
Nanosura Kennedy 1920 Syn. Kuldanagaster Yousuf and Yunus 1974 Syn.
Bedfordia Mumford 1942 Homo. Archegaster Lohmann 1992 Syn.
Ischnuridia Donnelly 1965 Syn. Kalyptogaster Lohmann 1992 Syn.

Ischnura barberi Currie 1903 (Ischnura)

ODONATA: COENAGRIONIDAE, CORDULEGASTRIDAE


NOMINA INSECTA NEARCTICA 565

Lauragaster Lohmann 1992 Syn. Helocordulia selysii Hagen 1878 (Cordulia)


Pangaeagaster Lohmann 1992 Syn. Helocordulia uhleri Selys 1871 (Cordulia)
Sonjagaster Lohmann 1992 Syn.
Macromia Rambur 1842
Cordulegaster bilineata Carle 1983 (Zoraena) Phyllomacromia Selys 1878 Syn.
Cordulegaster diadema Selys 1868 (Cordulegaster) Pseudogomphus Kirby 1889 Syn.
Cordulegaster diastatops Selys 1854 (Thecaphora) Ceratopyga Nunney 1895 Syn.
Cordulegaster lateralis Scudder 1866 Syn. Hylaeschna Sjstedt 1900 Syn.
Cordulegaster dorsalis Hagen 1858 (Cordulegaster)
Cordulegaster deserticola Cruden 1969 Syn. (subspecies) Macromia alleghaniensis Williamson 1909 (Macromia)
Cordulegaster erroneus Hagen 1878 (Cordulegaster) Macromia annulata Hagen 1861 (Macromia)
Cordulegaster maculatus Selys 1854 (Cordulegaster) Macromia caderita Needham 1950 Syn.
Cordulegaster obliqua Say 1839 (Aeschna) Macromia illinoiensis Walsh 1862 (Macromia)
Cordulegaster fasciatus Rambur 1842 Syn. (subspecies) Epophthalmia georgina Selys 1878 Syn. (subspecies)
Cordulegaster sayi Selys 1854 (Cordulegaster) Macromia australensis Williamson 1909 Syn.
Macromia magnifica McLachlan 1874 (Macromia)
Macromia rickeri Walker 1937 Syn.
Macromia margarita Westfall 1947 (Macromia)
Macromia pacifica Hagen 1861 (Macromia)
CORDULIIDAE Macromia flavipennis Walsh 1862 Syn.
Macromia taeniolata Rambur 1842 (Macromia)

Neurocordulia Selys 1871


Cordulia Leach 1815 Platycordulia Williamson 1908 Syn.

Cordulia shurtleffi Scudder 1866 (Cordulia) Neurocordulia alabamensis Hodges 1955 (Neurocordulia)
Cordulia bifurcata Hagen 1861 Unav. Neurocordulia molesta Walsh 1863 (Cordulia)
Neurocordulia clara Muttkowski 1910 Syn.
Didymops Rambur 1842 Neurocordulia obsoleta Say 1839 (Libellula)
Libellula polysticta Burmeister 1839 Syn.
Didymops floridensis Davis 1921 (Didymops) Neurocordulia virginiensis Davis 1927 (Neurocordulia)
Didymops transversa Say 1839 (Libellula) Neurocordulia xanthosoma Williamson 1908 (Platycordulia)
Epophthalmia cinnamomea Burmeister 1839 Syn. Neurocordulia yamaskanensis Provancher 1875 (Aeshna)
Didymops servillei Rambur 1842 Syn. Epitheca jamascarensis Hagen 1878 Syn.

Dorocordulia Needham 1901 Somatochlora Selys 1871


Chlorosoma Charpentier 1840 Homo.
Dorocordulia libera Selys 1871 (Cordulia)
Dorocordulia lepida Hagen 1871 (Cordulia) Somatochlora albicincta Burmeister 1839 (Epophthalmia)
Cordulia eremita Scudder 1866 Syn.
Epitheca Burmeister 1839 Somatochlora massettensis Whitehouse 1941 Syn.
Tetragoneuria Hagen 1861 Syn. Somatochlora brevicincta Robert 1954 (Somatochlora)
Epicordulia Selys 1871 Syn. Somatochlora calverti Williamson and Gloyd 1933 (Somatochlora)
Somatochlora cingulata Selys 1871 (Epitheca)
Epitheca canis McLachlan 1886 (Epitheca) Somatochlora elongata Scudder 1866 (Cordulia)
Epitheca costalis Selys 1871 (Cordulia) Cordulia saturata Hagen 1861 Unav.
Epitheca williamsoni Muttkowski 1911 Syn. Somatochlora ensigera Martin 1907 (Somatochlora)
Epitheca cynosura Say 1839 (Libellula) Somatochlora charadraea Williamson 1907 Syn.
Libellula tomentosa Fabricius 1775 Nomob. Somatochlora filosa Hagen 1861 (Cordulia)
Epophthalmia lateralis Burmeister 1839 Syn. Somatochlora forcipata Scudder 1866 (Cordulia)
Cordulia complanata Rambur 1842 Syn. Cordulia chalybea Hagen 1861 Unav.
Cordulia basiguttata Selys 1871 Syn. Somatochlora franklini Selys 1878 (Epitheca)
Cordulia diffinis Selys 1871 Syn. Somatochlora macrotona Williamson 1909 Syn.
Epitheca simulans Muttkowski 1911 Syn. Somatochlora georgiana Walker 1925 (Somatochlora)
Epitheca petechialis Muttkowski 1911 (Epitheca) Somatochlora hineana Williamson 1931 (Somatochlora)
Epitheca princeps Hagen 1861 (Epitheca) Somatochlora hudsonica Hagen 1871 (Epitheca)
Cordulia regina Hagen 1871 Syn. (subspecies) Somatochlora incurvata Walker 1918 (Somatochlora)
Epitheca semiaquea Burmeister 1839 (Libellula) Somatochlora kennedyi Walker 1918 (Somatochlora)
Epitheca diffinis Hagen 1861 Unav. Somatochlora linearis Hagen 1861 (Cordulia)
Epitheca calverti Muttkowski 1915 Syn. Epitheca procera Selys 1871 Syn.
Epitheca sepia Gloyd 1933 (Epitheca) Somatochlora lateralis Needham 1901 Missp.
Epitheca spinigera Selys 1871 (Epitheca) Somatochlora margarita Donnelly 1962 (Somatochlora)
Epitheca indistincta Morse 1895 Syn. Somatochlora minor Calvert 1898 (Somatochlora)
Epitheca spinosa Hagen 1878 (Cordulia) Somatochlora ozarkensis Bird 1933 (Somatochlora)
Epitheca stella Williamson 1911 (Epitheca) Somatochlora provocans Calvert 1903 (Somatochlora)

Helocordulia Needham 1901

ODONATA: CORDULEGASTRIDAE, CORDULIIDAE


566 NOMINA INSECTA NEARCTICA

Somatochlora sahlbergi Trybom 1889 (Somatochlora) Gomphus Leach 1815


Somatochlora walkeri Kennedy 1917 Syn. Thanatophora Selys 1850 Syn.
Somatochlora semicircularis Selys 1871 (Epitheca) Gomphurus Needham 1901 Syn.
Epitheca nasalis Selys 1874 Syn. Hylogomphus Needham 1951 Unav.
Somatochlora septentrionalis Hagen 1861 (Cordulia) Hylogomphus Needham and Westfall 1955 Syn.
Cordulia richardsoni Hagen 1861 Unav. Phanogomphus Carle 1986 Syn.
Somatochlora tenebrosa Say 1839 (Libellula) Stenogomphurus Carle 1986 Syn.
Cordulia tenebrica Hagen 1861 Unav.
Somatochlora walshii Scudder 1866 (Cordulia) Gomphus abbreviatus Hagen 1878 (Gomphus)
Somatochlora whitehousei Walker 1925 (Somatochlora) Gomphus adelphus Selys 1858 (Gomphus)
Somatochlora williamsoni Walker 1907 (Somatochlora) Gomphus brevis Hagen 1878 Syn.
Gomphus apomyius Donnelly 1966 (Gomphus)
Williamsonia Davis 1913 Gomphus australis Needham 1897 (Arigomphus)
Gomphus borealis Needham 1900 (Gomphus)
Williamsonia fletcheri Williamson 1923 (Williamsonia) Gomphus cavillaris Needham 1902 (Gomphus)
Diplax vacua Hagen 1867 Unav. Gomphus brimleyi Muttkowski 1911 Syn.
Williamsonia lintneri Hagen 1878 (Cordulia) Gomphus consanguis Selys 1879 (Gomphus)
Gomphus crassus Hagen 1878 (Gomphus)
Gomphus walshii Kellicott 1899 Syn.
Gomphus descriptus Banks 1896 (Gomphus)
Gomphus argus Needham 1943 Syn.
GOMPHIDAE Gomphus mortimer Needham 1943 Syn.
Gomphus dilatatus Rambur 1842 (Gomphus)
Gomphus diminutus Needham 1950 (Gomphus)
Gomphus exilis Selys 1854 (Gomphus)
Aphylla Selys 1854 Gomphus flavocaudatus Walker 1940 Syn.
Gomphus externus Hagen 1858 (Gomphus)
Aphylla angustifolia Garrison 1986 (Aphylla) Gomphus consobrinus Walsh 1863 Syn.
Aphylla protracta Hagen 1859 (Cyclophylla) Gomphus fraterna Say 1839 (Aeshna)
Gomphoides ambigua Selys 1873 Syn. Gomphus manitobanus Walker 1958 Syn. (subspecies)
Aphylla williamsoni Gloyd 1936 (Gomphoides) Gomphus geminatus Carle 1979 (Gomphus)
Gomphus gonzalezi Dunkle 1992 (Gomphus)
Arigomphus Needham 1897 Gomphus graslinellus Walsh 1862 (Gomphus)
Orcus Needham 1897 Homo. Gomphus hodgesi Needham 1950 (Gomphus)
Gomphus hybridus Williamson 1902 (Gomphus)
Arigomphus cornutus Tough 1900 (Gomphus) Gomphus kurilis Hagen 1858 (Gomphus)
Gomphus whedoni Muttkowski 1913 Syn. Gomphus confraternus Selys 1873 Syn.
Arigomphus furcifer Hagen 1878 (Gomphus) Gomphus sobrinus Selys 1873 Syn.
Arigomphus lentulus Needham 1902 (Gomphus) Gomphus donneri Kennedy 1917 Syn.
Gomphus subapicalis Williamson 1914 Syn. Gomphus lineatifrons Calvert 1921 (Gomphus)
Arigomphus maxwelli Ferguson 1950 (Gomphus) Gomphus lividus Selys 1854 (Gomphus)
Arigomphus pallidus Rambur 1842 (Gomphus) Gomphus sordidus Hagen 1854 Syn.
Gomphus pilipes Hagen 1858 Syn. Gomphus umbratus Needham 1897 Syn.
Arigomphus submedianus Williamson 1914 (Gomphus) Gomphus lynnae Paulson 1983 (Gomphus)
Arigomphus villosipes Selys 1854 (Gomphus) Gomphus militaris Hagen 1858 (Gomphus)
Gomphus minutus Rambur 1842 (Gomphus)
Dromogomphus Selys 1854 Gomphus modestus Needham 1942 (Gomphus)
Gomphus oklahomensis Pritchard 1935 (Gomphus)
Dromogomphus armatus Selys 1854 (Dromogomphus) Gomphus ozarkensis Westfall 1975 (Gomphus)
Dromogomphus spinosus Selys 1854 (Dromogomphus) Gomphus parvidens Currie 1917 (Gomphus)
Dromogomphus spoliatus Hagen 1858 (Gomphus) Gomphus carolinus Carle 1979 Syn.
Gomphus quadricolor Walsh 1862 (Gomphus)
Erpetogomphus Selys 1858 Gomphus alleni Howe 1922 Syn.
Herpetogomphus Walsh 1862 Emend. Gomphus rogersi Gloyd 1936 (Gomphus)
Calogomphus Carle 1992 Syn. Gomphus sandrius Tennessen 1983 (Gomphus)
Erpetocyclops Carle 1992 Syn. Gomphus septima Westfall 1956 (Gomphus)
Gomphus spicatus Hagen 1854 (Gomphus)
Erpetogomphus compositus Hagen 1858 (Erpetogomphus) Gomphus vastus Walsh 1862 (Gomphus)
Erpetogomphus coluber Williamson and Williamson 1930 Syn. Gomphus ventricosus Walsh 1863 (Gomphus)
Erpetogomphus crotalinus Hagen 1854 (Ophiogomphus) Gomphus viridifrons Hine 1901 (Gomphus)
Erpetogomphus designatus Hagen 1858 (Erpetogomphus) Gomphus westfalli Carle and May 1987 (Gomphus)
Erpetogomphus eutainia Calvert 1905 (Erpetogomphus)
Erpetogomphus diadophis Calvert 1905 Syn. Hagenius Selys 1854
Erpetogomphus heterodon Garrison 1994 (Erpetogomphus)
Erpetogomphus lampropeltis Kennedy 1918 (Erpetogomphus) Hagenius brevistylus Selys 1854 (Hagenius)
Erpetogomphus natrix Williamson and Williamson 1930 Syn.
(subspecies)

ODONATA: CORDULIIDAE, GOMPHIDAE


NOMINA INSECTA NEARCTICA 567

Lanthus Needham 1897 Stylurus amnicola Walsh 1862 (Gomphus)


Gomphus abditus Butler 1914 Syn.
Lanthus parvulus Selys 1854 (Gomphus) Stylurus intricatus Hagen 1858 (Gomphus)
Lanthus vernalis Carle 1980 (Lanthus) Stylurus ivae Williamson 1932 (Stylurus)
Stylurus laurae Williamson 1932 (Stylurus)
Octogomphus Selys 1873 Stylurus notatus Rambur 1842 (Gomphus)
Gomphus fluvialis Walsh 1862 Syn.
Octogomphus specularis Hagen 1859 (Neogomphus) Gomphus jucundus Needham 1943 Syn.
Stylurus olivaceus Selys 1873 (Gomphus)
Ophiogomphus Selys 1854 Gomphus nevadensis Kennedy 1917 Syn.
Ophionuroides Carle 1986 Syn. Stylurus plagiatus Selys 1854 (Gomphus)
Ophionurus Carle 1986 Syn. Gomphus elongatus Selys 1854 Syn.
Stylurus potulentus Needham 1942 (Gomphus)
Ophiogomphus acuminatus Carle 1981 (Ophiogomphus) Stylurus scudderi Selys 1873 (Gomphus)
Ophiogomphus bouchardi Louton 1982 Syn. Stylurus spiniceps Walsh 1862 (Macrogomphus)
Ophiogomphus anomalus Harvey 1898 (Ophiogomphus) Stylurus segregans Needham 1897 Syn.
Ophiogomphus arizonicus Kennedy 1917 (Ophiogomphus) Stylurus townesi Gloyd 1936 (Stylurus)
Ophiogomphus aspersus Morse 1895 (Ophiogomphus)
Ophiogomphus australis Carle 1992 (Ophiogomphus)
Ophiogomphus bison Selys 1873 (Ophiogomphus)
Ophiogomphus sequoiarum Butler 1914 Syn.
Ophiogomphus carolus Needham 1897 (Ophiogomphus) LESTIDAE
Ophiogomphus colubrinus Selys 1854 (Ophiogomphus)
Ophiogomphus edmundo Needham 1951 (Ophiogomphus)
Ophiogomphus howei Bromley 1924 (Ophiogomphus)
Ophiogomphus incurvatus Carle 1982 (Ophiogomphus) Archilestes Selys 1862
Ophiogomphus alleghaniensis Carle 1982 Syn. (subspecies) Cyptolestes Williamson 1921 Syn.
Ophiogomphus mainensis Packard 1863 (Ophiogomphus) Superlestes Williamson 1921 Syn.
Ophiogomphus johannus Needham 1897 Syn.
Ophiogomphus fastigiatus Donnelly 1987 Syn. (subspecies) Archilestes californica McLachlan 1895 (Archilestes)
Ophiogomphus morrisoni Selys 1879 (Ophiogomphus) Archilestes grandis Rambur 1842 (Lestes)
Ophiogomphus nevadensis Kennedy 1917 Syn.
Ophiogomphus occidentis Hagen 1882 (Ophiogomphus) Lestes Leach 1815
Ophiogomphus phaleratus Needham 1902 Syn. Puella Brulle 1832 Syn.
Ophiogomphus californicus Kennedy 1917 Syn. Anapates Charpentier 1840 Syn.
Ophiogomphus rupinsulensis Walsh 1862 (Herpetogomphus) Africalestes Kennedy 1920 Syn.
Ophiogomphus carolinus Hagen 1885 Syn. Chalcolestes Kennedy 1920 Syn.
Herpetogomphus pictus Needham 1897 Syn. Paralestes Schmidt 1951 Syn.
Ophiogomphus severus Hagen 1874 (Ophiogomphus) Xerolestes Fraser 1951 Syn.
Ophiogomphus montanus Selys 1878 Syn. (subspecies) Icterolestes Pinhey 1980 Syn.
Ophiogomphus susbehcha Vogt and Smith 1993 (Ophiogomphus) Malgassolestes Pinhey 1980 Syn.
Ophiogomphus westfalli Cook and Daigle 1985 (Ophiogomphus) Pseudochalcolestes Pinhey 1980 Syn.

Phyllogomphoides Belle 1970 Lestes alacer Hagen 1861 (Lestes)


Lestes sublata Hagen 1862 Syn.
Phyllogomphoides albrighti Needham 1950 (Gomphoides) Lestes congener Hagen 1861 (Lestes)
Phyllogomphoides stigmata Say 1839 (Aeschna) Lestes disjuncta Selys 1862 (Lestes)
Lestes australis Walker 1952 Syn. (subspecies)
Progomphus Selys 1854 Lestes dryas Kirby 1890 (Lestes)
Gomphoides Selys 1850 Syn. ? Lestes uncatus Kirby 1890 Syn.
Alloprogomphus Carle 1986 Syn. Lestes eurinus Say 1839 (Lestes)
Archaeoprogomphus Carle 1986 Syn. Lestes forcipata Rambur 1842 (Lestes)
Eoprogomphus Carle 1986 Syn. Lestes hamata Hagen 1861 Syn.
Neaprogomphus Carle 1986 Syn. Lestes forficula Rambur 1842 (Lestes)
Lestes striata Selys 1862 Syn.
Progomphus alachuensis Byers 1939 (Progomphus) Lestes inaequalis Walsh 1862 (Lestes)
Progomphus bellei Knopf and Tennessen 1980 (Progomphus) Lestes virgo Calvert 1897 Unav.
Progomphus borealis McLachlan 1873 (Progomphus) Lestes rectangularis Say 1839 (Lestes)
Progomphus meridionalis Hagen 1885 Unav. Lestes spumaria Hagen 1862 (Lestes)
Progomphus obscurus Rambur 1842 (Diastatomma) Lestes stulta Hagen 1861 (Lestes)
Lestes tenuata Rambur 1842 (Lestes)
Stylogomphus Fraser 1922 Lestes unguiculata Hagen 1861 (Lestes)
Lestes vidua Hagen 1861 (Lestes)
Stylogomphus albistylus Hagen 1878 (Gomphus) Lestes vigilax Hagen 1862 (Lestes)
Gomphus naevius Hagen 1878 Syn.

Stylurus Needham 1897

ODONATA: GOMPHIDAE, LESTIDAE


568 NOMINA INSECTA NEARCTICA

Libellula mithra Selys 1857 Syn.


Erythemis collocata Hagen 1861 (Mesothemis)
LIBELLULIDAE Erythemis plebeja Burmeister 1839 (Libellula)
Lepthemis verbenata Hagen 1861 Syn.
Erythemis simplicicollis Say 1839 (Libellula)
Libellula imbuta Say 1839 Syn.
Brachymesia Kirby 1889 Libellula caerulans Rambur 1842 Syn.
Cannacria Kirby 1889 Syn. Libellula maculiventris Rambur 1842 Syn.
Mesothemis gundlachi Scudder 1866 Syn.
Brachymesia furcata Hagen 1861 (Erythemis) Erythemis vesiculosa Fabricius 1775 (Libellula)
Brachymesia australis Kirby 1889 Syn. Libellula acuta Say 1839 Syn.
Cannacria smithii Kirby 1894 Syn.
Brachymesia gravida Calvert 1890 (Lepthemis) Erythrodiplax Brauer 1868
Brachymesia herbida Gundlach 1889 (Libellula) Nadiplax Navas 1917 Syn.
Cannacria batesi Kirby 1889 Syn.
Cannacria fumipennis Currie 1901 Syn. Erythrodiplax berenice Drury 1770 (Libellula)
Libellula histrio Burmeister 1839 Syn.
Brechmorhoga Kirby 1894 Dythemis naeva Hagen 1861 Syn. (subspecies)
Nothemis Navas 1915 Syn. Erythrodiplax connata Burmeister 1839 (Libellula)
Libellula communis Rambur 1842 Syn.
Brechmorhoga mendax Hagen 1861 (Dythemis) Libellula chloropleura Brauer 1865 Syn.
Brechmorhoga pertinax Hagen 1861 (Dythemis) Libellula leontina Brauer 1865 Syn.
Libellula sallaei Selys 1868 Syn. Diplax fraterna Hagen 1873 Syn.
Brechmorhoga vivax Calvert 1906 (Brechmorhoga) Erythrodiplax ponderosa Karsch 1891 Syn.
Trithemis basifusca Calvert 1895 Syn.
Cannaphila Kirby 1889 Erythrodiplax funerea Hagen 1861 (Libellula)
Neurothemis affinis Kirby 1889 Syn.
Cannaphila insularis Kirby 1889 (Cannaphila) Trithemis tyleri Kirby 1899 Syn.
Misagria funerea Carpenter 1897 Syn. (subspecies) Erythrodiplax fusca Rambur 1842 (Libellula)
Libellula incompta Rambur 1842 Syn.
Celithemis Hagen 1861 Erythrodiplax justiniana Selys 1857 (Libellula)
Diplax ambusta Hagen 1868 Syn.
Celithemis amanda Hagen 1861 (Diplax) Diplax portoricana Kolbe 1888 Syn.
Libellula pulchella Burmeister 1839 Homo. Erythrodiplax minuscula Rambur 1842 (Libellula)
Celithemis bertha Williamson 1922 (Celithemis) Erythrodiplax umbrata Linnaeus 1758 (Libellula)
Celithemis leonura Westfall 1952 Syn. Libellula unifasciata De Geer 1773 Syn.
Celithemis elisa Hagen 1861 (Diplax) Libellula fallax Burmeister 1839 Syn.
Celithemis eponina Drury 1773 (Libellula) Libellula ruralis Burmeister 1839 Syn.
Libellula camilla Rambur 1842 Syn. Libellula subfasciata Burmeister 1839 Syn.
Libellula lucilla Rambur 1842 Syn. Libellula tripartita Burmeister 1839 Syn.
Celithemis fasciata Kirby 1889 (Celithemis) Libellula flavicans Rambur 1842 Syn.
Celithemis monomelaena Williamson 1901 Syn. Libellula fuscofasciata Blanchard 1845 Syn.
Celithemis martha Williamson 1922 (Celithemis) Trithemis montezuma Calvert 1899 Syn.
Celithemis ornata Rambur 1842 (Libellula)
Celithemis verna Pritchard 1935 (Celithemis) Idiataphe Cowley 1934
Ephidatia Kirby 1889 Homo.
Crocothemis Brauer 1868
Beblecia Kirby 1900 Syn. Idiataphe cubensis Scudder 1866 (Macromia)
Erythemis specularis Hagen 1867 Unav.
Crocothemis servilia Drury 1770 (Libellula)
Libellula ferruginea Fabricius 1793 Syn. Leucorrhinia Brittinger 1850
Libellula soror Rambur 1842 Syn. Coenotiata Buchecker 1876 Syn.
Crocothemis reticulata Kirby 1886 Syn.
Leucorrhinia borealis Hagen 1890 (Leucorrhinia)
Dythemis Hagen 1861 Leucorrhinia frigida Hagen 1890 (Leucorrhinia)
Leucorrhinia glacialis Hagen 1890 (Leucorrhinia)
Dythemis fugax Hagen 1861 (Dythemis) Leucorrhinia hudsonica Selys 1850 (Libellula)
Dythemis maya Calvert 1906 (Dythemis) Leucorrhinia hageni Calvert 1890 Syn.
Dythemis nigrescens Calvert 1899 (Dythemis) Leucorrhinia intacta Hagen 1861 (Diplax)
Dythemis velox Hagen 1861 (Dythemis) Leucorrhinia patricia Walker 1940 (Leucorrhinia)
Leucorrhinia proxima Calvert 1890 (Leucorrhinia)
Erythemis Hagen 1861
Lepthemis Hagen 1861 Syn. Libellula Linnaeus 1758
Mesothemis Hagen 1861 Syn. Leptetrum Newman 1833 Syn.
Platetrum Newman 1833 Syn.
Erythemis attala Selys 1857 (Libellula) Pigiphila Buchecker 1876 Syn.
Libellula annulata Rambur 1842 Homo. Plathemis Hagen 1861 Syn.
Libellula annulosa Selys 1857 Syn. Holotania Kirby 1889 Syn.

ODONATA: LIBELLULIDAE
NOMINA INSECTA NEARCTICA 569

Ladona Needham 1897 Syn. Miathyria marcella Selys 1857 (Libellula)


Belonia Kennedy 1922 Syn.
Eolibellula Kennedy 1922 Syn. Micrathyria Kirby 1889
Eurothemis Kennedy 1922 Syn.
Neotetrum Kennedy 1922 Syn. Micrathyria aequalis Hagen 1861 (Dythemis)
Syntetrum Kennedy 1922 Syn. Micrathyria septima Selys 1900 Syn.
Micrathyria didyma Selys 1857 (Libellula)
Libellula auripennis Burmeister 1839 (Libellula) Dythemis dicrota Hagen 1861 Syn.
Libellula costalis Rambur 1842 Syn. Mesothemis poeyi Scudder 1866 Syn.
Libellula axilena Westwood 1837 (Libellula) Micrathyria pruinosa Kirby 1894 Syn.
Libellula leda Say 1839 Syn. Micrathyria hagenii Kirby 1890 (Micrathyria)
Libellula comanche Calvert 1907 (Libellula)
Libellula flavida Hagen 1861 Homo. Nannothemis Brauer 1868
Libellula composita Hagen 1873 (Mesothemis) Aino Kirby 1890 Syn.
Libellula croceipennis Selys 1868 (Libellula)
Belonia uniformis Kirby 1889 Syn. Nannothemis bella Uhler 1857 (Nannophya)
Libellula aliasignata Muttkowski 1910 Syn. Nannophya pygmaea Kirby 1889 Syn.
Libellula cyanea Fabricius 1775 (Libellula) Aino puella Kirby 1890 Syn.
Libellula quadrupla Say 1839 Syn.
Libellula bistigma Uhler 1857 Syn. Orthemis Hagen 1861
Libellula deplanata Rambur 1842 (Libellula) Neocysta Kirby 1889 Syn.
Libellula exusta Say 1839 (Libellula)
Libellula flavida Rambur 1842 (Libellula) Orthemis ferruginea Fabricius 1775 (Libellula)
Libellula plumbea Uhler 1857 Syn. Libellula discolor Burmeister 1839 Syn.
Libellula foliata Kirby 1889 (Belonia) Libellula macrostigma Rambur 1842 Syn.
Libellula forensis Hagen 1861 (Libellula)
Libellula incesta Hagen 1861 (Libellula) Pachydiplax Brauer 1868
Libellula jesseana Williamson 1922 (Libellula)
Libellula julia Uhler 1857 (Libellula) Pachydiplax longipennis Burmeister 1839 (Libellula)
Libellula luctuosa Burmeister 1839 (Libellula) Libellula socia Rambur 1842 Syn.
Libellula basalis Say 1839 Homo.
Libellula odiosa Hagen 1861 Syn. Paltothemis Karsch 1890
Libellula lydia Drury 1770 (Libellula)
Libellula trimaculata De Geer 1773 Syn. Paltothemis lineatipes Karsch 1890 (Paltothemis)
Libellula serva Fabricius 1793 Syn. Dythemis russata Calvert 1895 Syn.
Libellula needhami Westfall 1943 (Libellula)
Libellula nodisticta Hagen 1861 (Libellula) Pantala Hagen 1861
Libellula pulchella Drury 1773 (Libellula)
Libellula versicolor Fabricius 1775 Syn. Pantala flavescens Fabricius 1798 (Libellula)
Libellula confusa Uhler 1857 Syn. Libellula viridula Beauvois 1805 Syn.
Libellula quadrimaculata Linnaeus 1758 (Libellula) Libellula analis Burmeister 1839 Syn.
Libellula quadripunctata Fabricius 1781 Syn. Libellula terminalis Burmeister 1839 Syn.
Libellula ternaria Say 1839 Syn. Pantala hymenaea Say 1839 (Libellula)
Libellula saturata Uhler 1857 (Libellula) Tramea huanacina Frster 1909 Syn.
Libellula semifasciata Burmeister 1839 (Libellula)
Libellula bifasciata Fabricius 1775 Nomob. Perithemis Hagen 1861
Libellula maculata Rambur 1842 Syn.
Libellula hersilia Blanchard 1849 Syn. Perithemis domitia Drury 1773 (Libellula)
Libellula subornata Hagen 1861 (Plathemis) Libellula metella Selys 1857 Syn.
Libellula vibrans Fabricius 1798 (Libellula) Perithemis iris Hagen 1861 Syn.
Perithemis pocahontas Kirby 1889 Syn.
Macrodiplax Brauer 1868 Perithemis intensa Kirby 1889 (Perithemis)
Perithemis californica Martin 1900 Syn.
Macrodiplax balteata Hagen 1861 (Tetragoneuria) Perithemis mooma Kirby 1889 (Perithemis)
Perithemis cloe Hagen 1861 Unav.
Macrothemis Hagen 1868 Perithemis octoxantha Ris 1910 Syn.
Cendra Navas 1917 Syn. Perithemis tenera Say 1839 (Libellula)
Libellula tenuicincta Say 1839 Syn.
Macrothemis imitans Karsch 1890 (Macrothemis) Libellula chlora Rambur 1842 Syn.
Macrothemis leucozona Ris 1913 Syn. (subspecies) Perithemis seminole Calvert 1907 Syn.
Macrothemis inacuta Calvert 1898 (Macrothemis)
Macrothemis rochai Navas 1918 Syn. Pseudoleon Kirby 1889
Macrothemis inequiunguis Calvert 1895 (Macrothemis)
Macrothemis vulgipes Calvert 1898 Syn. Pseudoleon superba Hagen 1861 (Celithemis)

Miathyria Kirby 1889


Nothifixis Navas 1916 Syn.

ODONATA: LIBELLULIDAE
570 NOMINA INSECTA NEARCTICA

Sympetrum Newman 1833 Tachopteryx thoreyi Hagen 1858 (Uropetala)


Diplax Burmeister 1839 Syn.
Thecadiplax Selys 1883 Syn. Tanypteryx Kennedy 1917
Tarnetrum Needham and Fisher 1936 Syn.
Kalosympetrum Carle 1993 Syn. Tanypteryx hageni Selys 1879 (Tachopteryx)

Sympetrum ambigua Rambur 1842 (Libellula)


Libellula albifrons Charpentier 1840 Homo.
Sympetrum corrupta Hagen 1861 (Mesothemis)
Sympetrum costifera Hagen 1861 (Diplax) PLATYSTICTIDAE
Diplax atripes Hagen 1873 Syn.
Sympetrum danae Sulzer 1776 (Libellula)
Sympetrum verum Bartenef 1915 Syn.
Sympetrum illota Hagen 1861 (Mesothemis) Palaemnema Selys 1860
Sympetrum internum Montgomery 1943 (Sympetrum)
Sympetrum janeae Carle 1993 (Sympetrum) Palaemnema domina Calvert 1903 (Palaemnema)
Sympetrum madida Hagen 1861 (Diplax)
Diplax flavicosta Hagen 1875 Syn.
Diplax chrysoptera Selys 1883 Syn.
Sympetrum obtrusa Hagen 1867 (Diplax)
Diplax decisa Hagen 1874 Syn.
Sympetrum occidentale Bartenef 1915 (Sympetrum)
PROTONEURIDAE
Sympetrum californicum Walker 1951 Syn. (subspecies)
Sympetrum fasciatum Walker 1951 Syn. (subspecies)
Sympetrum pallipes Hagen 1874 (Diplax)
Neoneura Selys 1860
Sympetrum morrisoni Ris 1911 Syn.
Caenoneura Kirby 1890 Syn.
Sympetrum rubicundula Say 1839 (Libellula)
Libellula assimilata Uhler 1857 Syn.
Neoneura aaroni Calvert 1903 (Neoneura)
Sympetrum semicincta Say 1839 (Libellula)
Sympetrum signiferum Cannings and Garrison 1991 (Sympetrum)
Protoneura Selys 1857
Sympetrum vicina Hagen 1861 (Diplax)
Protoneura cara Calvert 1903 (Protoneura)
Tauriphila Kirby 1889

Tauriphila australis Hagen 1867 (Tramea)


Tramea iphigenia Hagen 1867 Syn.
Tauriphila azteca Calvert 1906 (Tauriphila)

Tholymis Hagen 1867

Tholymis citrina Hagen 1867 (Tholymis)

Tramea Hagen 1861


Trapezostigma Hagen 1849 Syn.?

Tramea abdominalis Rambur 1842 (Libellula)


Libellula basalis Burmeister 1839 Homo.
Tramea binotata Rambur 1842 (Libellula)
Tramea brasiliana Brauer 1867 Syn.
Tramea longicauda Brauer 1867 Syn.
Tramea subbinotata Brauer 1867 Syn.
Tramea paulina Frster 1910 Syn.
Tramea walkeri Whitehouse 1943 Syn.
Tramea calverti Muttkowski 1910 (Tramea)
Tramea carolina Linnaeus 1763 (Libellula)
Tramea insularis Hagen 1861 (Tramea)
Tramea lacerata Hagen 1861 (Tramea)
Tramea onusta Hagen 1861 (Tramea)

PETALURIDAE

Tachopteryx Uhler 1859

ODONATA: LIBELLULIDAE, PETALURIDAE, PLATYSTICTIDAE, PROTONEURIDAE


NOMINA INSECTA NEARCTICA 571

GENERIC INDEX
Dorocordulia Needham (Corduliidae)
Acanthagrion Selys (Coenagrionidae) Dromogomphus Selys (Gomphidae)
Acanthagyna Kirby (Aeshnidae) Gynacantha Dythemis Hagen (Libellulidae)
Adversaeschna Watson (Aeshnidae) Aeshna Enallagma Charpentier (Coenagrionidae)
Aenallagma Selys (Coenagrionidae) Enallagma Eolibellula Kennedy (Libellulidae) Libellula
Aeshna Fabricius (Aeshnidae) Eoprogomphus Carle (Gomphidae) Progomphus
Africalestes Kennedy (Lestidae) Lestes Ephidatia Kirby (Libellulidae) Idiataphe
Africallagma Kennedy (Coenagrionidae) Enallagma Epiaeschna Hagen (Aeshnidae)
Agrion Fabricius (Calopterygidae) Calopteryx Epicordulia Selys (Corduliidae) Epitheca
Agrion Leach (Coenagrionidae) Coenagrion Epitheca Burmeister (Corduliidae)
Aino Kirby (Libellulidae) Nannothemis Erpetocyclops Carle (Gomphidae) Erpetogomphus
Alloprogomphus Carle (Gomphidae) Progomphus Erpetogomphus Selys (Gomphidae)
Amphiagrion Selys (Coenagrionidae) Erythemis Hagen (Libellulidae)
Amphiallagma Kennedy (Coenagrionidae) Enallagma Erythragrion Selys (Coenagrionidae) Telebasis
Anaciagrion Kennedy (Calopterygidae) Calopteryx Erythrodiplax Brauer (Libellulidae)
Anapates Charpentier (Lestidae) Lestes Eurothemis Kennedy (Libellulidae) Libellula
Anax Leach (Aeshnidae)
Anomalagrion Selys (Coenagrionidae) Ischnura Fonscolombia Selys (Aeshnidae) Boyeria
Anomalura Kennedy (Coenagrionidae) Ischnura
Apanisagrion Kennedy (Coenagrionidae) Gomphaeschna Selys (Aeshnidae)
Aphylla Selys (Gomphidae) Gomphoides Selys (Gomphidae) Progomphus
Archaeoprogomphus Carle (Gomphidae) Progomphus Gomphurus Needham (Gomphidae) Gomphus
Archegaster Lohmann (Cordulegastridae) Cordulegaster Gomphus Leach (Gomphidae)
Archilestes Selys (Lestidae) Gynacantha Rambur (Aeshnidae)
Argia Rambur (Coenagrionidae)
Argiallagma Calvert (Coenagrionidae) Nehalennia Hagenius Selys (Gomphidae)
Argiallagma Selys (Coenagrionidae) Nehalennia Heliargia Kennedy (Coenagrionidae) Argia
Arigomphus Needham (Gomphidae) Helocordulia Needham (Corduliidae)
Austrocoenagrion Kennedy (Coenagrionidae) Coenagrion Herpetogomphus Walsh (Gomphidae) Erpetogomphus
Hesperaeschna Cockerell (Aeshnidae) Aeshna
Basiaeschna Selys (Aeshnidae) Hesperagrion Calvert (Coenagrionidae)
Beblecia Kirby (Libellulidae) Crocothemis Hetaerina Hagen (Calopterygidae)
Bedfordia Mumford (Coenagrionidae) Ischnura Holotania Kirby (Libellulidae) Libellula
Belonia Kennedy (Libellulidae) Libellula Hoplonaeschna Karsch (Aeshnidae) Oplonaeschna
Boyeria McLachlan (Aeshnidae) Hylaeschna Sjstedt (Corduliidae) Macromia
Brachymesia Kirby (Libellulidae) Hylogomphus Needham (Gomphidae) Gomphus
Brechmorhoga Kirby (Libellulidae) Hylogomphus Needham and Westfall (Gomphidae) Gomphus
Hyponeura Selys (Coenagrionidae) Argia
Caenoneura Kirby (Protoneuridae) Neoneura
Calogomphus Carle (Gomphidae) Erpetogomphus Icterolestes Pinhey (Lestidae) Lestes
Calopteryx Leach (Calopterygidae) Idiataphe Cowley (Libellulidae)
Cannacria Kirby (Libellulidae) Brachymesia Ischnallagma Kennedy (Coenagrionidae) Enallagma
Cannaphila Kirby (Libellulidae) Ischnura Charpentier (Coenagrionidae)
Celaenura Kennedy (Coenagrionidae) Ischnura Ischnuridia Donnelly (Coenagrionidae) Ischnura
Celithemis Hagen (Libellulidae)
Cendra Navas (Libellulidae) Macrothemis Kalosympetrum Carle (Libellulidae) Sympetrum
Ceratopyga Nunney (Corduliidae) Macromia Kalyptogaster Lohmann (Cordulegastridae) Cordulegaster
Chalcargia Kennedy (Coenagrionidae) Argia Kuldanagaster Yousuf and Yunus (Cordulegastridae) Cordulegaster
Chalcolestes Kennedy (Lestidae) Lestes
Chloragrion Hartinov and Hartinova (Coenagrionidae) Coenagrion Ladona Needham (Libellulidae) Libellula
Chlorosoma Charpentier (Corduliidae) Somatochlora Lanthus Needham (Gomphidae)
Chromagrion Needham (Coenagrionidae) Lauragaster Lohmann (Cordulegastridae) Cordulegaster
Cnemophila Wallengren (Aeshnidae) Aeshna Leptargia Kennedy (Coenagrionidae) Argia
Coenagrion Kirby (Coenagrionidae) Leptetrum Newman (Libellulidae) Libellula
Coenotiata Buchecker (Libellulidae) Leucorrhinia Lepthemis Hagen (Libellulidae) Erythemis
Cordulegaster Leach (Cordulegastridae) Lestes Leach (Lestidae)
Cordulia Leach (Corduliidae) Leucorrhinia Brittinger (Libellulidae)
Coryphaeschna Williamson (Aeshnidae) Libellula Linnaeus (Libellulidae)
Crocothemis Brauer (Libellulidae)
Cyanargia Kennedy (Coenagrionidae) Argia Macrodiplax Brauer (Libellulidae)
Cyptolestes Williamson (Lestidae) Archilestes Macromia Rambur (Corduliidae)
Macrothemis Hagen (Libellulidae)
Diargia Calvert (Coenagrionidae) Argia Malgassolestes Pinhey (Lestidae) Lestes
Didymops Rambur (Corduliidae) Marmaraeschna Calvert (Aeshnidae) Aeshna
Diplax Burmeister (Libellulidae) Sympetrum Mesothemis Hagen (Libellulidae) Erythemis
Miathyria Kirby (Libellulidae)

ODONATA: GENERIC INDEX


572 NOMINA INSECTA NEARCTICA

Micrargia Kennedy (Coenagrionidae) Argia Taeniogaster Selys (Cordulegastridae) Cordulegaster


Micrathyria Kirby (Libellulidae) Tanypteryx Kennedy (Petaluridae)
Micronympha Kirby (Coenagrionidae) Ischnura Tarnetrum Needham and Fisher (Libellulidae) Sympetrum
Myagrion Frster (Coenagrionidae) Acanthagrion Tauriphila Kirby (Libellulidae)
Teleallagma Kennedy (Coenagrionidae) Enallagma
Nadiplax Navas (Libellulidae) Erythrodiplax Telebasis Selys (Coenagrionidae)
Nannothemis Brauer (Libellulidae) Tetragoneuria Hagen (Corduliidae) Epitheca
Nanosura Kennedy (Coenagrionidae) Ischnura Thanatophora Selys (Gomphidae) Gomphus
Nasiaeschna Selys (Aeshnidae) Thecadiplax Selys (Libellulidae) Sympetrum
Neaprogomphus Carle (Gomphidae) Progomphus Thecagaster Selys (Cordulegastridae) Cordulegaster
Nehalennia Selys (Coenagrionidae) Thecaphora Charpentier (Cordulegastridae) Cordulegaster
Neocysta Kirby (Libellulidae) Orthemis Thecaphora Selys (Cordulegastridae) Cordulegaster
Neoerythromma Kennedy (Coenagrionidae) Thermagrion Frster (Coenagrionidae) Enallagma
Neoneura Selys (Protoneuridae) Tholymis Hagen (Libellulidae)
Neotetrum Kennedy (Libellulidae) Libellula Tramea Hagen (Libellulidae)
Neureclipa Navas (Aeshnidae) Aeshna Trapezostigma Hagen (Libellulidae) Tramea
Neurocordulia Selys (Corduliidae) Triacanthagyna Selys (Aeshnidae)
Nothemis Navas (Libellulidae) Brechmorhoga
Nothifixis Navas (Libellulidae) Miathyria Williamsonia Davis (Corduliidae)

Octogomphus Selys (Gomphidae) Xerolestes Fraser (Lestidae) Lestes


Ophiogomphus Selys (Gomphidae)
Ophionuroides Carle (Gomphidae) Ophiogomphus Zoniagrion Kennedy (Coenagrionidae)
Ophionurus Carle (Gomphidae) Ophiogomphus Zoraena Kirby (Cordulegastridae) Cordulegaster
Oplonaeschna Selys (Aeshnidae)
Orcus Needham (Gomphidae) Arigomphus
Orthemis Hagen (Libellulidae)

Pachydiplax Brauer (Libellulidae)


Palaemnema Selys (Platystictidae)
Paltothemis Karsch (Libellulidae)
Pangaeagaster Lohmann (Cordulegastridae) Cordulegaster
Pantala Hagen (Libellulidae)
Paralestes Schmidt (Lestidae) Lestes
Perithemis Hagen (Libellulidae)
Phanogomphus Carle (Gomphidae) Gomphus
Phyllogomphoides Belle (Gomphidae)
Phyllomacromia Selys (Corduliidae) Macromia
Pigiphila Buchecker (Libellulidae) Libellula
Platetrum Newman (Libellulidae) Libellula
Plathemis Hagen (Libellulidae) Libellula
Platycordulia Williamson (Corduliidae) Neurocordulia
Progomphus Selys (Gomphidae)
Proischnura Kennedy (Coenagrionidae) Enallagma
Protoneura Selys (Protoneuridae)
Pseudochalcolestes Pinhey (Lestidae) Lestes
Pseudogomphus Kirby (Corduliidae) Macromia
Pseudoleon Kirby (Libellulidae)
Puella Brulle (Lestidae) Lestes

Remartinia Navas (Aeshnidae)


Rhionaeschna Frster (Aeshnidae) Aeshna

Schizuraeschna Calvert (Aeshnidae) Aeshna


Secundaeschna Gtz (Aeshnidae) Aeshna
Selsiophlebia Frster (Aeshnidae) Gynacantha
Sobobapteron Pierce (Coenagrionidae) Enallagma
Somatochlora Selys (Corduliidae)
Sonjagaster Lohmann (Cordulegastridae) Cordulegaster
Stenogomphurus Carle (Gomphidae) Gomphus
Stylogomphus Fraser (Gomphidae)
Stylurus Needham (Gomphidae)
Superlestes Williamson (Lestidae) Archilestes
Sylphus Hagen (Calopterygidae) Calopteryx
Sympetrum Newman (Libellulidae)
Syntetrum Kennedy (Libellulidae) Libellula

Tachopteryx Uhler (Petaluridae)

ODONATA: GENERIC INDEX


NOMINA INSECTA NEARCTICA 573

SPECIES INDEX
aspersum Hagen Agrion (Coenagrionidae) Enallagma
aspersus Morse Ophiogomphus (Gomphidae) Ophiogomphus
a assimilata Uhler Libellula (Libellulidae) Sympetrum rubicundula
atra Gloyd Argia (Coenagrionidae) Argia fumipennis
aaroni Calvert Neoneura (Protoneuridae) Neoneura atripes Hagen Diplax (Libellulidae) Sympetrum costifera
abboti Hagen Aeschna (Aeshnidae) Coryphaeschna ingens attala Selys Libellula (Libellulidae) Erythemis
abbreviatum Selys Pyrrhosoma (Coenagrionidae) Amphiagrion aurantiaca Hagen Agrion (Coenagrionidae) Ischnura ramburii
abbreviatus Hagen Gomphus (Gomphidae) Gomphus auripennis Burmeister Libellula (Libellulidae) Libellula
abditus Butler Gomphus (Gomphidae) Stylurus amnicola australensis Williamson Macromia (Corduliidae) Macromia
abdominalis Rambur Libellula (Libellulidae) Tramea illinoiensis
acuminatus Carle Ophiogomphus (Gomphidae) Ophiogomphus
acuta Say Libellula (Libellulidae) Erythemis vesiculosa australis Carle Ophiogomphus (Gomphidae) Ophiogomphus
adelphus Selys Gomphus (Gomphidae) Gomphus australis Hagen Tramea (Libellulidae) Tauriphila
adnexa Hagen Aeschna (Aeshnidae) Coryphaeschna australis Kirby Brachymesia (Libellulidae) Brachymesia furcata
aequabilis Say Calopteryx (Calopterygidae) Calopteryx australis Needham Arigomphus (Gomphidae) Gomphus
aequalis Hagen Dythemis (Libellulidae) Micrathyria australis Walker Lestes (Lestidae) Lestes disjuncta
affinis Kirby Neurothemis (Libellulidae) Erythrodiplax funerea axilena Westwood Libellula (Libellulidae) Libellula
agrioides Calvert Argia (Coenagrionidae) Argia azteca Calvert Tauriphila (Libellulidae) Tauriphila
alabamensis Hodges Neurocordulia (Corduliidae) Neurocordulia
alacer Hagen Lestes (Lestidae) Lestes
alachuensis Byers Progomphus (Gomphidae) Progomphus
b
alberta Kennedy Argia (Coenagrionidae) Argia
balteata Hagen Tetragoneuria (Libellulidae) Macrodiplax
albicincta Burmeister Epophthalmia (Corduliidae) Somatochlora
barberi Currie Ischnura (Coenagrionidae) Ischnura
albifrons Charpentier Libellula (Libellulidae) Sympetrum ambigua
barretti Calvert Argia (Coenagrionidae) Argia
albistylus Hagen Gomphus (Gomphidae) Stylogomphus
basalis Burmeister Libellula (Libellulidae) Tramea abdominalis
albrighti Needham Gomphoides (Gomphidae) Phyllogomphoides
basalis Hagen Hetaerina (Calopterygidae) Hetaerina americana
aliasignata Muttkowski Libellula (Libellulidae) Libellula
basalis Say Lestes (Calopterygidae) Hetaerina americana
croceipennis
basalis Say Libellula (Libellulidae) Libellula luctuosa
alleghaniensis Carle Ophiogomphus (Gomphidae) Ophiogomphus
basidens Calvert Enallagma (Coenagrionidae) Enallagma
incurvatus
basifusca Calvert Trithemis (Libellulidae) Erythrodiplax connata
alleghaniensis Williamson Macromia (Corduliidae) Macromia
basiguttata Selys Cordulia (Corduliidae) Epitheca cynosura
alleni Howe Gomphus (Gomphidae) Gomphus quadricolor
batesi Kirby Cannacria (Libellulidae) Brachymesia herbida
amanda Hagen Diplax (Libellulidae) Celithemis
bella Uhler Nannophya (Libellulidae) Nannothemis
amata Hagen Calopteryx (Calopterygidae) Calopteryx
bellei Knopf and Tennessen Progomphus (Gomphidae) Progomphus
amazili Burmeister Aeschna (Aeshnidae) Anax
berenice Drury Libellula (Libellulidae) Erythrodiplax
ambigua Rambur Libellula (Libellulidae) Sympetrum
bertha Williamson Celithemis (Libellulidae) Celithemis
ambigua Selys Gomphoides (Gomphidae) Aphylla protracta
bifasciata Fabricius Libellula (Libellulidae) Libellula semifasciata
ambusta Hagen Diplax (Libellulidae) Erythrodiplax justiniana
bifurcata Hagen Cordulia (Corduliidae) Cordulia shurtleffi
americana Bartenef Aeshna (Aeshnidae) Aeshna juncea
bilineata Carle Zoraena (Cordulegastridae) Cordulegaster
americana Fabricius Agrion (Calopterygidae) Hetaerina
binotata Rambur Libellula (Libellulidae) Tramea
amnicola Walsh Gomphus (Gomphidae) Stylurus
binotatum Walsh Agrion (Coenagrionidae) Argia tibialis
amphion Selys Amphiagrion (Coenagrionidae) Amphiagrion
bipartita Selys Hetaerina (Calopterygidae) Hetaerina titia
saucium
bipunctulatum Hagen Agrion (Coenagrionidae) Argia
analis Burmeister Libellula (Libellulidae) Pantala flavescens
bipustulata Kirby Argia (Coenagrionidae) Argia bipunctulatum
angulatum Walker Coenagrion (Coenagrionidae) Coenagrion
bison Selys Ophiogomphus (Gomphidae) Ophiogomphus
angustifolia Garrison Aphylla (Gomphidae) Aphylla
bistigma Uhler Libellula (Libellulidae) Libellula cyanea
angustipennis Selys Sylphis (Calopterygidae) Calopteryx
boreale Selys Enallagma (Coenagrionidae) Enallagma
anna Williamson Enallagma (Coenagrionidae) Enallagma
borealis Hagen Leucorrhinia (Libellulidae) Leucorrhinia
annexum Hagen Agrion (Coenagrionidae) Enallagma cyathigerum
borealis McLachlan Progomphus (Gomphidae) Progomphus
annulata Hagen Macromia (Corduliidae) Macromia
borealis Needham Gomphus (Gomphidae) Gomphus
annulata Rambur Libellula (Libellulidae) Erythemis attala
boucardi Selys Telebasis (Coenagrionidae) Telebasis salvum
annulosa Selys Libellula (Libellulidae) Erythemis attala
bouchardi Louton Ophiogomphus (Gomphidae) Ophiogomphus
anomalum Rambur Agrion (Coenagrionidae) Ischnura hastata
acuminatus
anomalus Harvey Ophiogomphus (Gomphidae) Ophiogomphus
brasiliana Brauer Tramea (Libellulidae) Tramea binotata
antennatum Say Agrion (Coenagrionidae) Enallagma
brevicincta Robert Somatochlora (Corduliidae) Somatochlora
antilope Hagen Aeschna (Aeshnidae) Gomphaeschna
brevis Hagen Gomphus (Gomphidae) Gomphus adelphus
apicale Say Agrion (Coenagrionidae) Argia
brevistylus Selys Hagenius (Gomphidae) Hagenius
apicalis Burmeister Calopteryx (Calopterygidae) Calopteryx
brimleyi Muttkowski Gomphus (Gomphidae) Gomphus cavillaris
dimidiata
byersi Westfall Telebasis (Coenagrionidae) Telebasis
apomyius Donnelly Gomphus (Gomphidae) Gomphus
argus Needham Gomphus (Gomphidae) Gomphus descriptus
arida Kennedy Aeshna (Aeshnidae) Aeshna palmata c
arizonicus Kennedy Ophiogomphus (Gomphidae) Ophiogomphus
armata Hagen Aeschna (Aeshnidae) Oplonaeschna caderita Needham Macromia (Corduliidae) Macromia annulata
armatus Selys Dromogomphus (Gomphidae) Dromogomphus caerulans Rambur Libellula (Libellulidae) Erythemis simplicicollis

ODONATA: SPECIES INDEX


574 NOMINA INSECTA NEARCTICA

californica Calvert Aeshna (Aeshnidae) Aeshna constricta Say Aeshna (Aeshnidae) Aeshna
californica Hagen Hetaerina (Calopterygidae) Hetaerina americana cornutus Tough Gomphus (Gomphidae) Arigomphus
californica Kennedy Calopteryx (Calopterygidae) Calopteryx corrupta Hagen Mesothemis (Libellulidae) Sympetrum
aequabilis costalis Rambur Libellula (Libellulidae) Libellula auripennis
californica Martin Perithemis (Libellulidae) Perithemis intensa costalis Selys Cordulia (Corduliidae) Epitheca
californica McLachlan Archilestes (Lestidae) Archilestes costifera Hagen Diplax (Libellulidae) Sympetrum
californicum Walker Sympetrum (Libellulidae) Sympetrum crassus Hagen Gomphus (Gomphidae) Gomphus
occidentale credulum Hagen Agrion (Coenagrionidae) Ischnura ramburii
californicus Kennedy Ophiogomphus (Gomphidae) Ophiogomphus croceipennis Selys Libellula (Libellulidae) Libellula
occidentis crotalinus Hagen Ophiogomphus (Gomphidae) Erpetogomphus
calverti Morse Enallagma (Coenagrionidae) Enallagma boreale cubensis Scudder Macromia (Libellulidae) Idiataphe
calverti Muttkowski Epitheca (Corduliidae) Epitheca semiaquea culicinorum Byers Enallagma (Coenagrionidae) Enallagma anna
calverti Muttkowski Tramea (Libellulidae) Tramea cultellatum Selys Enallagma (Coenagrionidae) Neoerythromma
calverti Williamson and Gloyd Somatochlora (Corduliidae) cupreum Hagen Agrion (Coenagrionidae) Argia
Somatochlora cyanea Fabricius Libellula (Libellulidae) Libellula
camilla Rambur Libellula (Libellulidae) Celithemis eponina cyanifrons Hagen Aeschna (Aeshnidae) Coryphaeschna adnexa
canadense Provancher Agrion (Coenagrionidae) Ischnura verticalis cyathigerum Charpentier Agrion (Coenagrionidae) Enallagma
canadensis Walker Aeshna (Aeshnidae) Aeshna cynosura Say Libellula (Corduliidae) Epitheca
canis McLachlan Epitheca (Corduliidae) Epitheca
cara Calvert Protoneura (Protoneuridae) Protoneura
cardenium Hagen Enallagma (Coenagrionidae) Enallagma coecum d
carolina Linnaeus Libellula (Libellulidae) Tramea
carolinus Carle Gomphus (Gomphidae) Gomphus parvidens daeckii Calvert Telagrion (Coenagrionidae) Enallagma
carolinus Hagen Ophiogomphus (Gomphidae) Ophiogomphus damula Calvert Ischnura (Coenagrionidae) Ischnura
rupinsulensis danae Sulzer Libellula (Libellulidae) Sympetrum
davisi Westfall Enallagma (Coenagrionidae) Enallagma
carolott Butler Nehalennia (Coenagrionidae) Nehalennia irene decisa Hagen Diplax (Libellulidae) Sympetrum obtrusa
carolus Needham Ophiogomphus (Gomphidae) Ophiogomphus defixum Hagen Agrion (Coenagrionidae) Ischnura ramburii
carunculatum Morse Enallagma (Coenagrionidae) Enallagma demorsum Hagen Agrion (Coenagrionidae) Ischnura
cavillaris Needham Gomphus (Gomphidae) Gomphus denticollis Burmeister Agrion (Coenagrionidae) Ischnura
cervula Selys Ischnura (Coenagrionidae) Ischnura dentiferum Walsh Agrion (Coenagrionidae) Enallagma signatum
chalybea Hagen Cordulia (Corduliidae) Somatochlora forcipata deplanata Rambur Libellula (Libellulidae) Libellula
charadraea Williamson Somatochlora (Corduliidae) Somatochlora descriptus Banks Gomphus (Gomphidae) Gomphus
ensigera
chlora Rambur Libellula (Libellulidae) Perithemis tenera deserticola Cruden Cordulegaster (Cordulegastridae)
chloropleura Brauer Libellula (Libellulidae) Erythrodiplax connata Cordulegaster dorsalis
chrysoptera Selys Diplax (Libellulidae) Sympetrum madida designatus Hagen Erpetogomphus (Gomphidae) Erpetogomphus
cingulata Selys Epitheca (Corduliidae) Somatochlora diadema Selys Cordulegaster (Cordulegastridae) Cordulegaster
cinnamomea Burmeister Epophthalmia (Corduliidae) Didymops diadophis Calvert Erpetogomphus (Gomphidae) Erpetogomphus
transversa eutainia
citrina Hagen Tholymis (Libellulidae) Tholymis diastatops Selys Thecaphora (Cordulegastridae) Cordulegaster
civile Hagen Agrion (Coenagrionidae) Enallagma dicrota Hagen Dythemis (Libellulidae) Micrathyria didyma
clara Muttkowski Neurocordulia (Corduliidae) Neurocordulia didyma Selys Libellula (Libellulidae) Micrathyria
molesta diffinis Hagen Epitheca (Corduliidae) Epitheca semiaquea
clausum Morse Enallagma (Coenagrionidae) Enallagma diffinis Selys Cordulia (Corduliidae) Epitheca cynosura
clepsydra Say Aeshna (Aeshnidae) Aeshna dilatatus Rambur Gomphus (Gomphidae) Gomphus
cloe Hagen Perithemis (Libellulidae) Perithemis mooma dimidiata Burmeister Calopteryx (Calopterygidae) Calopteryx
coecum Hagen Enallagma (Coenagrionidae) Enallagma diminutus Needham Gomphus (Gomphidae) Gomphus
cognata Rambur Calopteryx (Calopterygidae) Calopteryx dimidiata discolor Burmeister Agrion (Coenagrionidae) Amphiagrion saucium
collocata Hagen Mesothemis (Libellulidae) Erythemis discolor Burmeister Libellula (Libellulidae) Orthemis ferruginea
coloradica Cockerell Calopteryx (Calopterygidae) Calopteryx disjuncta Selys Lestes (Lestidae) Lestes
aequabilis divagans Selys Enallagma (Coenagrionidae) Enallagma
coluber Williamson and Williamson Erpetogomphus (Gomphidae) domina Calvert Palaemnema (Platystictidae) Palaemnema
Erpetogomphus compositus dominicana Hagen Aeshna (Aeshnidae) Aeshna psilus
colubrinus Selys Ophiogomphus (Gomphidae) Ophiogomphus domitia Drury Libellula (Libellulidae) Perithemis
comanche Calvert Libellula (Libellulidae) Libellula donneri Kennedy Gomphus (Gomphidae) Gomphus kurilis
communis Rambur Libellula (Libellulidae) Erythrodiplax connata dorsalis Hagen Cordulegaster (Cordulegastridae) Cordulegaster
complanata Rambur Cordulia (Corduliidae) Epitheca cynosura doubledayi Selys Agrion (Coenagrionidae) Enallagma
composita Hagen Mesothemis (Libellulidae) Libellula dryas Kirby Lestes (Lestidae) Lestes
compositus Hagen Erpetogomphus (Gomphidae) Erpetogomphus dubium Root Enallagma (Coenagrionidae) Enallagma
concisum Williamson Enallagma (Coenagrionidae) Enallagma dugesi Calvert Aeshna (Aeshnidae) Aeshna
conditum Selys Erythromma (Coenagrionidae) Chromagrion durum Hagen Agrion (Coenagrionidae) Enallagma
confraternus Selys Gomphus (Gomphidae) Gomphus kurilis
confusa Uhler Libellula (Libellulidae) Libellula pulchella
congener Hagen Lestes (Lestidae) Lestes
e
connata Burmeister Libellula (Libellulidae) Erythrodiplax
ebrium Hagen Agrion (Coenagrionidae) Enallagma
consanguis Selys Gomphus (Gomphidae) Gomphus
edmundo Needham Ophiogomphus (Gomphidae) Ophiogomphus
consobrinus Walsh Gomphus (Gomphidae) Gomphus externus

ODONATA: SPECIES INDEX


NOMINA INSECTA NEARCTICA 575

elegans Hagen Sylphis (Calopterygidae) Calopteryx angustipennis furcata Hagen Erythemis (Libellulidae) Brachymesia
elisa Hagen Diplax (Libellulidae) Celithemis furcifer Hagen Gomphus (Gomphidae) Arigomphus
elongata Scudder Cordulia (Corduliidae) Somatochlora furcifera Karsch Aeshna (Aeshnidae) Aeshna multicolor
elongatus Selys Gomphus (Gomphidae) Stylurus plagiatus furcillata Say Aeschna (Aeshnidae) Gomphaeschna
emma Kennedy Argia (Coenagrionidae) Argia fusca Rambur Libellula (Libellulidae) Erythrodiplax
ensigera Martin Somatochlora (Corduliidae) Somatochlora fuscofasciata Blanchard Libellula (Libellulidae) Erythrodiplax
eponina Drury Libellula (Libellulidae) Celithemis umbrata
eremita Scudder Aeshna (Aeshnidae) Aeshna
eremita Scudder Cordulia (Corduliidae) Somatochlora albicincta
erratica Calvert Ischnura (Coenagrionidae) Ischnura g
erroneus Hagen Cordulegaster (Cordulegastridae) Cordulegaster
espinalensis Navas Argia (Coenagrionidae) Argia translata gemina Kennedy Celaenura (Coenagrionidae) Ischnura
eurinus Say Lestes (Lestidae) Lestes geminatum Kellicott Enallagma (Coenagrionidae) Enallagma
eutainia Calvert Erpetogomphus (Gomphidae) Erpetogomphus geminatus Carle Gomphus (Gomphidae) Gomphus
exclamationis Selys Agrion (Coenagrionidae) Zoniagrion georgiana Walker Somatochlora (Corduliidae) Somatochlora
exilis Selys Gomphus (Gomphidae) Gomphus georgina Selys Epophthalmia (Corduliidae) Macromia illinoiensis
expertum Hagen Agrion (Coenagrionidae) Ischnura ramburii glacialis Hagen Leucorrhinia (Libellulidae) Leucorrhinia
exstriata Calvert Ischnura (Coenagrionidae) Ischnura denticollis gonzalezi Dunkle Gomphus (Gomphidae) Gomphus
exsulans Hagen Agrion (Coenagrionidae) Enallagma gracilis Morse Nehalennia (Coenagrionidae) Nehalennia
externus Hagen Gomphus (Gomphidae) Gomphus grafiana Williamson Boyeria (Aeshnidae) Boyeria
extraneum Hagen Agrion (Coenagrionidae) Argia grandis Rambur Lestes (Lestidae) Archilestes
exusta Say Libellula (Libellulidae) Libellula graslinellus Walsh Gomphus (Gomphidae) Gomphus
gravida Calvert Lepthemis (Libellulidae) Brachymesia
gundlachi Scudder Mesothemis (Libellulidae) Erythemis
f simplicicollis
guyanensis Machet Coryphaeschna (Aeshnidae) Coryphaeschna
fallax Burmeister Libellula (Libellulidae) Erythrodiplax umbrata adnexa
fasciata Kirby Celithemis (Libellulidae) Celithemis
fasciatum Walker Sympetrum (Libellulidae) Sympetrum occidentale
fasciatus Rambur Cordulegaster (Cordulegastridae) Cordulegaster h
obliqua
fastigiatus Donnelly Ophiogomphus (Gomphidae) Ophiogomphus hageni Calvert Leucorrhinia (Libellulidae) Leucorrhinia hudsonica
mainensis hageni Selys Tachopteryx (Petaluridae) Tanypteryx
ferruginea Fabricius Libellula (Libellulidae) Crocothemis servilia hageni Walsh Agrion (Coenagrionidae) Enallagma
ferruginea Fabricius Libellula (Libellulidae) Orthemis hagenii Kirby Micrathyria (Libellulidae) Micrathyria
filosa Hagen Cordulia (Corduliidae) Somatochlora hamata Hagen Lestes (Lestidae) Lestes forcipata
fischeri Kellicott Enallagma (Coenagrionidae) Enallagma hastata Say Agrion (Coenagrionidae) Ischnura
antennatum hastulatum Stephens Agrion (Coenagrionidae) Enallagma
flavescens Fabricius Libellula (Libellulidae) Pantala cyathigerum
flavescens Selys Agrion (Coenagrionidae) Hesperagrion herbida Gundlach Libellula (Libellulidae) Brachymesia
heterodoxum heros Fabricius Aeschna (Aeshnidae) Epiaeschna
flavicans Rambur Libellula (Libellulidae) Erythrodiplax umbrata hersilia Blanchard Libellula (Libellulidae) Libellula semifasciata
flavicosta Hagen Diplax (Libellulidae) Sympetrum madida heterodon Garrison Erpetogomphus (Gomphidae) Erpetogomphus
flavida Hagen Libellula (Libellulidae) Libellula comanche heterodoxum Selys Amphiagrion (Coenagrionidae) Hesperagrion
flavida Rambur Libellula (Libellulidae) Libellula hineana Williamson Somatochlora (Corduliidae) Somatochlora
flavipennis Walsh Macromia (Corduliidae) Macromia pacifica hinei Kennedy Argia (Coenagrionidae) Argia
flavocaudatus Walker Gomphus (Gomphidae) Gomphus exilis histrio Burmeister Libellula (Libellulidae) Erythrodiplax berenice
fletcheri Williamson Williamsonia (Corduliidae) Williamsonia hodgesi Needham Gomphus (Gomphidae) Gomphus
florida Hagen Aeschna (Aeshnidae) Remartinia luteipennis holosericea Burmeister Calopteryx (Calopterygidae) Calopteryx
floridana Huggins Calopteryx (Calopterygidae) Calopteryx maculata
maculata howei Bromley Ophiogomphus (Gomphidae) Ophiogomphus
floridensis Davis Didymops (Corduliidae) Didymops huanacina Frster Tramea (Libellulidae) Pantala hymenaea
fluvialis Walsh Gomphus (Gomphidae) Stylurus notatus hudsonica Hagen Calopteryx (Calopterygidae) Calopteryx
foliata Kirby Belonia (Libellulidae) Libellula aequabilis
fontium Hagen Agrion (Coenagrionidae) Argia tibialis hudsonica Hagen Epitheca (Corduliidae) Somatochlora
forcipata Rambur Lestes (Lestidae) Lestes hudsonica Selys Aeshna (Aeshnidae) Aeshna eremita
forcipata Scudder Cordulia (Corduliidae) Somatochlora hudsonica Selys Libellula (Libellulidae) Leucorrhinia
forensis Hagen Libellula (Libellulidae) Libellula hybridus Williamson Gomphus (Gomphidae) Gomphus
forficula Rambur Lestes (Lestidae) Lestes hymenaea Say Libellula (Libellulidae) Pantala
franklini Selys Epitheca (Corduliidae) Somatochlora
fraterna Hagen Diplax (Libellulidae) Erythrodiplax connata
fraterna Say Aeshna (Gomphidae) Gomphus
i
frigida Hagen Leucorrhinia (Libellulidae) Leucorrhinia
fugax Hagen Dythemis (Libellulidae) Dythemis
illinoiensis Walsh Macromia (Corduliidae) Macromia
fumipennis Burmeister Agrion (Coenagrionidae) Argia
illota Hagen Mesothemis (Libellulidae) Sympetrum
fumipennis Currie Cannacria (Libellulidae) Brachymesia herbida
imbuta Say Libellula (Libellulidae) Erythemis simplicicollis
funerea Carpenter Misagria (Libellulidae) Cannaphila insularis
imitans Karsch Macrothemis (Libellulidae) Macrothemis
funerea Hagen Libellula (Libellulidae) Erythrodiplax
immundum Hagen Agrion (Coenagrionidae) Argia

ODONATA: SPECIES INDEX


576 NOMINA INSECTA NEARCTICA

inacuta Calvert Macrothemis (Libellulidae) Macrothemis leontina Brauer Libellula (Libellulidae) Erythrodiplax connata
inaequalis Walsh Lestes (Lestidae) Lestes leonura Westfall Celithemis (Libellulidae) Celithemis bertha
incesta Hagen Libellula (Libellulidae) Libellula lepida Hagen Cordulia (Corduliidae) Dorocordulia
incompta Rambur Libellula (Libellulidae) Erythrodiplax fusca leucozona Ris Macrothemis (Libellulidae) Macrothemis imitans
incurvata Walker Somatochlora (Corduliidae) Somatochlora libera Selys Cordulia (Corduliidae) Dorocordulia
incurvatus Carle Ophiogomphus (Gomphidae) Ophiogomphus limbata Selys Hetaerina (Calopterygidae) Hetaerina titia
indistincta Morse Epitheca (Corduliidae) Epitheca spinigera linearis Hagen Cordulia (Corduliidae) Somatochlora
inequiunguis Calvert Macrothemis (Libellulidae) Macrothemis lineata Walker Aeshna (Aeshnidae) Aeshna interrupta
iners Hagen Agrion (Coenagrionidae) Ischnura ramburii
ingens Rambur Aeschna (Aeshnidae) Coryphaeschna lineatifrons Calvert Gomphus (Gomphidae) Gomphus
insularis Hagen Tramea (Libellulidae) Tramea lineatipes Karsch Paltothemis (Libellulidae) Paltothemis
insularis Kirby Cannaphila (Libellulidae) Cannaphila lintneri Hagen Cordulia (Corduliidae) Williamsonia
intacta Hagen Diplax (Libellulidae) Leucorrhinia lividus Selys Gomphus (Gomphidae) Gomphus
integricollis Calvert Nehalennia (Coenagrionidae) Nehalennia longicauda Brauer Tramea (Libellulidae) Tramea binotata
intensa Kirby Perithemis (Libellulidae) Perithemis longipennis Burmeister Libellula (Libellulidae) Pachydiplax
interna Walker Aeshna (Aeshnidae) Aeshna interrupta longipes Hagen Anax (Aeshnidae) Anax
internum Montgomery Sympetrum (Libellulidae) Sympetrum lucilla Rambur Libellula (Libellulidae) Celithemis eponina
interrogatum Selys Agrion (Coenagrionidae) Coenagrion luctuosa Burmeister Libellula (Libellulidae) Libellula
interrupta Walker Aeshna (Aeshnidae) Aeshna lugens Hagen Hyponeura (Coenagrionidae) Argia
intricatus Hagen Gomphus (Gomphidae) Stylurus luteipennis Burmeister Aeschna (Aeshnidae) Remartinia
intruda Williamson Argia (Coenagrionidae) Argia moestum lydia Drury Libellula (Libellulidae) Libellula
iphigenia Hagen Tramea (Libellulidae) Tauriphila australis lynnae Paulson Gomphus (Gomphidae) Gomphus
irene Hagen Agrion (Coenagrionidae) Nehalennia
iris Hagen Perithemis (Libellulidae) Perithemis domitia
ivae Williamson Stylurus (Gomphidae) Stylurus m
macromia Brauer Aeschna (Aeshnidae) Coryphaeschna adnexa
j macrostigma Rambur Libellula (Libellulidae) Orthemis ferruginea
macrotona Williamson Somatochlora (Corduliidae) Somatochlora
jamascarensis Hagen Epitheca (Corduliidae) Neurocordulia franklini
yamaskanensis maculata Beauvois Agrion (Calopterygidae) Calopteryx
janata Say Aeschna (Aeshnidae) Basiaeschna maculata Rambur Libellula (Libellulidae) Libellula semifasciata
janeae Carle Sympetrum (Libellulidae) Sympetrum maculatus Rambur Anax (Aeshnidae) Anax amazili
jesseana Williamson Libellula (Libellulidae) Libellula maculatus Selys Cordulegaster (Cordulegastridae) Cordulegaster
johannus Needham Ophiogomphus (Gomphidae) Ophiogomphus maculiventris Rambur Libellula (Libellulidae) Erythemis
mainensis simplicicollis
jucundus Needham Gomphus (Gomphidae) Stylurus notatus madida Hagen Diplax (Libellulidae) Sympetrum
julia Uhler Libellula (Libellulidae) Libellula magnifica McLachlan Macromia (Corduliidae) Macromia
juncea Linnaeus Libellula (Aeshnidae) Aeshna mainensis Packard Ophiogomphus (Gomphidae) Ophiogomphus
junia Drury Libellula (Aeshnidae) Anax manitobanus Walker Gomphus (Gomphidae) Gomphus fraterna
justiniana Selys Libellula (Libellulidae) Erythrodiplax marcella Selys Libellula (Libellulidae) Miathyria
margarita Donnelly Somatochlora (Corduliidae) Somatochlora
margarita Westfall Macromia (Corduliidae) Macromia
k maritimum Provancher Anax (Aeshnidae) Aeshna canadensis
martha Williamson Celithemis (Libellulidae) Celithemis
kellicotti Williamson Ischnura (Coenagrionidae) Ischnura massettensis Whitehouse Somatochlora (Corduliidae) Somatochlora
kennedyi Walker Somatochlora (Corduliidae) Somatochlora albicincta
krugii Kolbe Enallagma (Coenagrionidae) Enallagma coecum materna Say Calopteryx (Calopterygidae) Calopteryx maculata
kurilis Hagen Argia (Coenagrionidae) Argia vivida maxwelli Ferguson Gomphus (Gomphidae) Arigomphus
kurilis Hagen Gomphus (Gomphidae) Gomphus maya Calvert Dythemis (Libellulidae) Dythemis
mendax Hagen Dythemis (Libellulidae) Brechmorhoga
l meridionalis Hagen Progomphus (Gomphidae) Progomphus
borealis
metella Selys Libellula (Libellulidae) Perithemis domitia
lacerata Hagen Tramea (Libellulidae) Tramea
militaris Hagen Gomphus (Gomphidae) Gomphus
lacrimans Hagen Agrion (Coenagrionidae) Argia
minor Calvert Somatochlora (Corduliidae) Somatochlora
lais Brauer Nehalennia (Coenagrionidae) Apanisagrion
minor Rambur Aeschna (Aeshnidae) Basiaeschna janata
lampropeltis Kennedy Erpetogomphus (Gomphidae) Erpetogomphus
minuscula Rambur Libellula (Libellulidae) Erythrodiplax
laterale Morse Enallagma (Coenagrionidae) Enallagma
minusculum Morse Enallagma (Coenagrionidae) Enallagma
lateralis Burmeister Epophthalmia (Corduliidae) Epitheca cynosura
minutus Rambur Gomphus (Gomphidae) Gomphus
lateralis Needham Somatochlora (Corduliidae) Somatochlora
mista Navas Argia (Coenagrionidae) Argia extraneum
linearis
mithra Selys Libellula (Libellulidae) Erythemis attala
lateralis Scudder Cordulegaster (Cordulegastridae) Cordulegaster
modestus Needham Gomphus (Gomphidae) Gomphus
diastatops
moestum Hagen Agrion (Coenagrionidae) Argia
laurae Williamson Stylurus (Gomphidae) Stylurus
molesta Walsh Cordulia (Corduliidae) Neurocordulia
laurenti Calvert Enallagma (Coenagrionidae) Enallagma vesperum
monomelaena Williamson Celithemis (Libellulidae) Celithemis
leda Say Libellula (Libellulidae) Libellula axilena
fasciata
lentulus Needham Gomphus (Gomphidae) Arigomphus
montanus Selys Ophiogomphus (Gomphidae) Ophiogomphus severus
leonorae Garrison Argia (Coenagrionidae) Argia

ODONATA: SPECIES INDEX


NOMINA INSECTA NEARCTICA 577

montezuma Calvert Trithemis (Libellulidae) Erythrodiplax umbrata pentacantha Rambur Aeschna (Aeshnidae) Nasiaeschna
mooma Kirby Perithemis (Libellulidae) Perithemis perparva Selys Ischnura (Coenagrionidae) Ischnura
morrisoni Ris Sympetrum (Libellulidae) Sympetrum pallipes persephone Donnelly Aeshna (Aeshnidae) Aeshna
morrisoni Selys Ophiogomphus (Gomphidae) Ophiogomphus pertinax Hagen Dythemis (Libellulidae) Brechmorhoga
mortimer Needham Gomphus (Gomphidae) Gomphus descriptus petechialis Muttkowski Epitheca (Corduliidae) Epitheca
multicincta Say Aeschna (Aeshnidae) Epiaeschna heros phaleratus Needham Ophiogomphus (Gomphidae) Ophiogomphus
multicolor Hagen Aeshna (Aeshnidae) Aeshna occidentis
munda Calvert Argia (Coenagrionidae) Argia pictum Morse Enallagma (Coenagrionidae) Enallagma
mutata Hagen Aeshna (Aeshnidae) Aeshna pictus Needham Herpetogomphus (Gomphidae) Ophiogomphus
rupinsulensis
pilipes Hagen Gomphus (Gomphidae) Arigomphus pallidus
n pima Garrison Argia (Coenagrionidae) Argia
piscinarium Williamson Enallagma (Coenagrionidae) Enallagma
naeva Hagen Dythemis (Libellulidae) Erythrodiplax berenice geminatum
naevius Hagen Gomphus (Gomphidae) Stylogomphus albistylus plagiatus Selys Gomphus (Gomphidae) Stylurus
nahuana Calvert Argia (Coenagrionidae) Argia plana Calvert Argia (Coenagrionidae) Argia
nasalis Selys Epitheca (Corduliidae) Somatochlora semicircularis plebeja Burmeister Libellula (Libellulidae) Erythemis
natrix Williamson and Williamson Erpetogomphus (Gomphidae) plebium Selys Enallagma (Coenagrionidae) Enallagma civile
Erpetogomphus lampropeltis plumbea Uhler Libellula (Libellulidae) Libellula flavida
needhami Westfall Libellula (Libellulidae) Libellula pocahontas Kirby Perithemis (Libellulidae) Perithemis domitia
nervosa Rambur Gynacantha (Aeshnidae) Gynacantha poeyi Scudder Mesothemis (Libellulidae) Micrathyria didyma
polernista Navas Oplonaeschna (Aeshnidae) Oplonaeschna armata
nevadensis Kennedy Gomphus (Gomphidae) Stylurus olivaceus pollutum Hagen Agrion (Coenagrionidae) Enallagma
nevadensis Kennedy Ophiogomphus (Gomphidae) Ophiogomphus polysticta Burmeister Libellula (Corduliidae) Neurocordulia
morrisoni obsoleta
nevadensis Walker Aeshna (Aeshnidae) Aeshna interrupta ponderosa Karsch Erythrodiplax (Libellulidae) Erythrodiplax
nigrescens Calvert Dythemis (Libellulidae) Dythemis connata
nodisticta Hagen Libellula (Libellulidae) Libellula portoricana Kolbe Diplax (Libellulidae) Erythrodiplax justiniana
notatus Rambur Gomphus (Gomphidae) Stylurus positum Hagen Agrion (Coenagrionidae) Ischnura
novaehispaniae Calvert Enallagma (Coenagrionidae) Enallagma potulentus Needham Gomphus (Gomphidae) Stylurus
praevarum Hagen Agrion (Coenagrionidae) Enallagma
o princeps Hagen Epitheca (Corduliidae) Epitheca
procera Selys Epitheca (Corduliidae) Somatochlora linearis
prognatum Hagen Agrion (Coenagrionidae) Ischnura
obliqua Say Aeschna (Cordulegastridae) Cordulegaster
obscura Rambur Argia (Coenagrionidae) Argia fumipennis
propinqua Scudder Aeshna (Aeshnidae) Aeshna verticalis
obscurus Rambur Diastatomma (Gomphidae) Progomphus
protracta Hagen Cyclophylla (Gomphidae) Aphylla
obsoleta Say Libellula (Corduliidae) Neurocordulia
provocans Calvert Somatochlora (Corduliidae) Somatochlora
obtrusa Hagen Diplax (Libellulidae) Sympetrum
proxima Calvert Leucorrhinia (Libellulidae) Leucorrhinia
occidentale Bartenef Sympetrum (Libellulidae) Sympetrum
pruinosa Kirby Micrathyria (Libellulidae) Micrathyria didyma
occidentalis Walker Aeshna (Aeshnidae) Aeshna umbrosa
pseudamericana Walsh Hetaerina (Calopterygidae) Hetaerina
occidentis Hagen Ophiogomphus (Gomphidae) Ophiogomphus
americana
octoxantha Ris Perithemis (Libellulidae) Perithemis mooma
psilus Calvert Aeshna (Aeshnidae) Aeshna
odiosa Hagen Libellula (Libellulidae) Libellula luctuosa
puella Kirby Aino (Libellulidae) Nannothemis bella
oenea Hagen Argia (Coenagrionidae) Argia
pulchella Burmeister Libellula (Libellulidae) Celithemis amanda
oklahomensis Pritchard Gomphus (Gomphidae) Gomphus
pulchella Drury Libellula (Libellulidae) Libellula
olivaceus Selys Gomphus (Gomphidae) Stylurus
putridum Hagen Agrion (Coenagrionidae) Argia moestum
onusta Hagen Tramea (Libellulidae) Tramea
pygmaea Kirby Nannophya (Libellulidae) Nannothemis bella
opaca Say Calopteryx (Calopterygidae) Calopteryx maculata
ornata Rambur Libellula (Libellulidae) Celithemis
ozarkensis Bird Somatochlora (Corduliidae) Somatochlora q
ozarkensis Westfall Gomphus (Gomphidae) Gomphus
quadratum Selys Acanthagrion (Coenagrionidae) Acanthagrion
p quadricolor Walsh Gomphus (Gomphidae) Gomphus
quadrifida Rambur Gynacantha (Aeshnidae) Gomphaeschna
furcillata
pacifica Hagen Macromia (Corduliidae) Macromia
quadriguttata Burmeister Aeschna (Aeshnidae) Boyeria vinosa
pallens Calvert Argia (Coenagrionidae) Argia
quadrimaculata Linnaeus Libellula (Libellulidae) Libellula
pallidula Calvert Nehalennia (Coenagrionidae) Nehalennia
quadripunctata Fabricius Libellula (Libellulidae) Libellula
pallidum Root Enallagma (Coenagrionidae) Enallagma
quadrimaculata
pallidus Rambur Gomphus (Gomphidae) Arigomphus
quadrupla Say Libellula (Libellulidae) Libellula cyanea
pallipes Hagen Diplax (Libellulidae) Sympetrum
palmata Hagen Aeshna (Aeshnidae) Aeshna
papilionaria Rambur Calopteryx (Calopterygidae) Calopteryx r
maculata
parvidens Currie Gomphus (Gomphidae) Gomphus ramburii Selys Agrion (Coenagrionidae) Ischnura
parvulus Selys Gomphus (Gomphidae) Lanthus rectangularis Say Lestes (Lestidae) Lestes
patricia Walker Leucorrhinia (Libellulidae) Leucorrhinia recurvatum Davis Enallagma (Coenagrionidae) Enallagma
paulina Frster Tramea (Libellulidae) Tramea binotata

ODONATA: SPECIES INDEX


578 NOMINA INSECTA NEARCTICA

regina Hagen Cordulia (Corduliidae) Epitheca princeps socia Rambur Libellula (Libellulidae) Pachydiplax longipennis
resolutum Selys Agrion (Coenagrionidae) Coenagrion solita Kennedy Argia (Coenagrionidae) Argia pallens
reticulata Kirby Crocothemis (Libellulidae) Crocothemis servilia sordidus Hagen Gomphus (Gomphidae) Gomphus lividus
rhoadsi Calvert Argia (Coenagrionidae) Argia soror Rambur Libellula (Libellulidae) Crocothemis servilia
richardsoni Hagen Cordulia (Corduliidae) Somatochlora specularis Hagen Erythemis (Libellulidae) Idiataphe cubensis
septentrionalis specularis Hagen Neogomphus (Gomphidae) Octogomphus
rickeri Walker Macromia (Corduliidae) Macromia magnifica spegazzinii Navas Argia (Coenagrionidae) Argia plana
rita Kennedy Argia (Coenagrionidae) Argia munda spicatus Hagen Gomphus (Gomphidae) Gomphus
robustum Selys Enallagma (Coenagrionidae) Enallagma spiniceps Walsh Macrogomphus (Gomphidae) Stylurus
cyathigerum spiniferus Rambur Anax (Aeshnidae) Anax junia
rochai Navas Macrothemis (Libellulidae) Macrothemis inacuta spinigera Selys Epitheca (Corduliidae) Epitheca
rogersi Gloyd Gomphus (Gomphidae) Gomphus spinosa Hagen Cordulia (Corduliidae) Epitheca
rubicundula Say Libellula (Libellulidae) Sympetrum spinosus Selys Dromogomphus (Gomphidae) Dromogomphus
rupamnensis Walsh Hetaerina (Calopterygidae) Hetaerina titia spoliatus Hagen Gomphus (Gomphidae) Dromogomphus
rupinsulensis Walsh Herpetogomphus (Gomphidae) Ophiogomphus spumaria Hagen Lestes (Lestidae) Lestes
rupinsulensis Walsh Hetaerina (Calopterygidae) Hetaerina titia stella Williamson Epitheca (Corduliidae) Epitheca
ruralis Burmeister Libellula (Libellulidae) Erythrodiplax umbrata stigmata Say Aeschna (Gomphidae) Phyllogomphoides
russata Calvert Dythemis (Libellulidae) Paltothemis lineatipes striata Selys Lestes (Lestidae) Lestes forficula
stulta Hagen Lestes (Lestidae) Lestes
subapicalis Williamson Gomphus (Gomphidae) Arigomphus lentulus
s subarctica Walker Aeshna (Aeshnidae) Aeshna
subbinotata Brauer Tramea (Libellulidae) Tramea binotata
saalasi Valle Argia (Coenagrionidae) Argia nahuana subfasciata Burmeister Libellula (Libellulidae) Erythrodiplax
sabino Garrison Argia (Coenagrionidae) Argia umbrata
sahlbergi Trybom Somatochlora (Corduliidae) Somatochlora sublata Hagen Lestes (Lestidae) Lestes alacer
sallaei Selys Libellula (Libellulidae) Brechmorhoga pertinax submedianus Williamson Gomphus (Gomphidae) Arigomphus
salvum Hagen Agrion (Coenagrionidae) Telebasis subornata Hagen Plathemis (Libellulidae) Libellula
sandrius Tennessen Gomphus (Gomphidae) Gomphus sulcatum Williamson Enallagma (Coenagrionidae) Enallagma
saturata Hagen Cordulia (Corduliidae) Somatochlora elongata superba Hagen Celithemis (Libellulidae) Pseudoleon
saturata Uhler Libellula (Libellulidae) Libellula susbehcha Vogt and Smith Ophiogomphus (Gomphidae)
saucium Burmeister Agrion (Coenagrionidae) Amphiagrion Ophiogomphus
sayi Selys Cordulegaster (Cordulegastridae) Cordulegaster
sclerata Walsh Hetaerina (Calopterygidae) Hetaerina americana
scudderi Selys Gomphus (Gomphidae) Stylurus t
sedulum Hagen Agrion (Coenagrionidae) Argia
segregans Needham Stylurus (Gomphidae) Stylurus spiniceps taeniolata Rambur Macromia (Corduliidae) Macromia
selysii Hagen Cordulia (Corduliidae) Helocordulia tarascana Calvert Argia (Coenagrionidae) Argia
semiaquea Burmeister Libellula (Corduliidae) Epitheca tenebrica Hagen Cordulia (Corduliidae) Somatochlora tenebrosa
semicincta Say Libellula (Libellulidae) Sympetrum tenebrosa Say Libellula (Corduliidae) Somatochlora
semicirculare Selys Enallagma (Coenagrionidae) Enallagma tenera Say Libellula (Libellulidae) Perithemis
semicircularis Selys Epitheca (Corduliidae) Somatochlora tenuata Rambur Lestes (Lestidae) Lestes
semifasciata Burmeister Libellula (Libellulidae) Libellula tenuicincta Say Libellula (Libellulidae) Perithemis tenera
seminole Calvert Perithemis (Libellulidae) Perithemis tenera terminalis Burmeister Libellula (Libellulidae) Pantala flavescens
sepia Gloyd Epitheca (Corduliidae) Epitheca ternaria Say Libellula (Libellulidae) Libellula quadrimaculata
septentrionalis Burmeister Aeshna (Aeshnidae) Aeshna texana Walsh Hetaerina (Calopterygidae) Hetaerina americana
septentrionalis Hagen Cordulia (Corduliidae) Somatochlora tezpi Calvert Argia (Coenagrionidae) Argia
septentrionalis Selys Hetaerina (Calopterygidae) Hetaerina titia thoreyi Hagen Uropetala (Petaluridae) Tachopteryx
septima Selys Gynacantha (Aeshnidae) Triacanthagyna tibialis Rambur Platycnemis (Coenagrionidae) Argia
septima Selys Micrathyria (Libellulidae) Micrathyria aequalis titia Drury Libellula (Calopterygidae) Hetaerina
septima Westfall Gomphus (Gomphidae) Gomphus tomentosa Fabricius Libellula (Corduliidae) Epitheca cynosura
tonto Calvert Argia (Coenagrionidae) Argia
sequoiarum Butler Ophiogomphus (Gomphidae) Ophiogomphus townesi Gloyd Stylurus (Gomphidae) Stylurus
bison translata Hagen Argia (Coenagrionidae) Argia
serva Fabricius Libellula (Libellulidae) Libellula lydia transversa Say Libellula (Corduliidae) Didymops
servilia Drury Libellula (Libellulidae) Crocothemis traviatum Selys Enallagma (Coenagrionidae) Enallagma
servillei Rambur Didymops (Corduliidae) Didymops transversa tricolor Burmeister Calopteryx (Calopterygidae) Hetaerina titia
servum Selys Agrion (Coenagrionidae) Coenagrion resolutum trimaculata De Geer Libellula (Libellulidae) Libellula lydia
severus Hagen Ophiogomphus (Gomphidae) Ophiogomphus tripartita Burmeister Libellula (Libellulidae) Erythrodiplax umbrata
shurtleffi Scudder Cordulia (Corduliidae) Cordulia tuberculata Selys Ischnura (Coenagrionidae) Ischnura ramburii
signatum Hagen Agrion (Coenagrionidae) Enallagma tuberculifera Walker Aeshna (Aeshnidae) Aeshna
signiferum Cannings and Garrison Sympetrum (Libellulidae) tyleri Kirby Trithemis (Libellulidae) Erythrodiplax funerea
Sympetrum
simile Selys Enallagma (Coenagrionidae) Enallagma civile
simplicicollis Say Libellula (Libellulidae) Erythemis u
simulans Muttkowski Epitheca (Corduliidae) Epitheca cynosura
sitchensis Hagen Aeshna (Aeshnidae) Aeshna uhleri Selys Cordulia (Corduliidae) Helocordulia
smithii Kirby Cannacria (Libellulidae) Brachymesia furcata umbrata Linnaeus Libellula (Libellulidae) Erythrodiplax
sobrinus Selys Gomphus (Gomphidae) Gomphus kurilis umbratus Needham Gomphus (Gomphidae) Gomphus lividus

ODONATA: SPECIES INDEX


NOMINA INSECTA NEARCTICA 579

umbrosa Walker Aeshna (Aeshnidae) Aeshna williamsoni Walker Somatochlora (Corduliidae) Somatochlora
uncatus Kirby Lestes (Lestidae) Lestes dryas
unguiculata Hagen Lestes (Lestidae) Lestes
unifasciata De Geer Libellula (Libellulidae) Erythrodiplax umbrata x
uniformis Kirby Belonia (Libellulidae) Libellula croceipennis
utahensis Muttkowski Ischnura (Coenagrionidae) Ischnura barberi xanthosoma Williamson Platycordulia (Corduliidae) Neurocordulia

v y
vacua Hagen Diplax (Corduliidae) Williamsonia fletcheri yakima Hagen Calopteryx (Calopterygidae) Calopteryx aequabilis
validus Hagen Anax (Aeshnidae) Anax walsinghami yamaskanensis Provancher Aeshna (Corduliidae) Neurocordulia
vastus Walsh Gomphus (Gomphidae) Gomphus ypsilon Needham Enallagma (Coenagrionidae) Enallagma
velox Hagen Dythemis (Libellulidae) Dythemis novaehispaniae
venerinotata Haldeman Agrion (Coenagrionidae) Ischnura hastata
ventricosus Walsh Gomphus (Gomphidae) Gomphus
verbenata Hagen Lepthemis (Libellulidae) Erythemis plebeja
verna Pritchard Celithemis (Libellulidae) Celithemis
vernale Gloyd Enallagma (Coenagrionidae) Enallagma cyathigerum
vernalis Carle Lanthus (Gomphidae) Lanthus
versicolor Fabricius Libellula (Libellulidae) Libellula pulchella
verticalis Hagen Aeshna (Aeshnidae) Aeshna
verticalis Say Agrion (Coenagrionidae) Ischnura
verum Bartenef Sympetrum (Libellulidae) Sympetrum danae
vesiculosa Fabricius Libellula (Libellulidae) Erythemis
vesperum Calvert Enallagma (Coenagrionidae) Enallagma
vibrans Fabricius Libellula (Libellulidae) Libellula
vicina Hagen Diplax (Libellulidae) Sympetrum
vidua Hagen Lestes (Lestidae) Lestes
vigilax Hagen Lestes (Lestidae) Lestes
villosipes Selys Gomphus (Gomphidae) Arigomphus
vinosa Say Aeschna (Aeshnidae) Boyeria
violaceum Hagen Agrion (Coenagrionidae) Argia fumipennis
virens Rambur Aeschna (Aeshnidae) Coryphaeschna viriditas
virginica Selys Calopteryx (Calopterygidae) Calopteryx aequabilis
virginiensis Davis Neurocordulia (Corduliidae) Neurocordulia
virgo Calvert Lestes (Lestidae) Lestes inaequalis
viridifrons Hine Gomphus (Gomphidae) Gomphus
viriditas Calvert Coryphaeschna (Aeshnidae) Coryphaeschna
viridula Beauvois Libellula (Libellulidae) Pantala flavescens
vivax Calvert Brechmorhoga (Libellulidae) Brechmorhoga
vivida Hagen Argia (Coenagrionidae) Argia
vulgipes Calvert Macrothemis (Libellulidae) Macrothemis
inequiunguis
vulnerata Hagen Hetaerina (Calopterygidae) Hetaerina

w
walkeri Kennedy Aeshna (Aeshnidae) Aeshna
walkeri Kennedy Somatochlora (Corduliidae) Somatochlora
sahlbergi
walkeri Muttkowski Enallagma (Coenagrionidae) Coenagrion
resolutum
walkeri Whitehouse Tramea (Libellulidae) Tramea binotata
walshii Kellicott Gomphus (Gomphidae) Gomphus crassus
walshii Scudder Cordulia (Corduliidae) Somatochlora
walsinghami McLachlan Anax (Aeshnidae) Anax
weewa Byers Enallagma (Coenagrionidae) Enallagma
westfalli Carle and May Gomphus (Gomphidae) Gomphus
westfalli Cook and Daigle Ophiogomphus (Gomphidae)
Ophiogomphus
westfalli Donnelly Enallagma (Coenagrionidae) Enallagma
traviatum
whedoni Muttkowski Gomphus (Gomphidae) Arigomphus cornutus
whitehousei Walker Somatochlora (Corduliidae) Somatochlora
williamsoni Gloyd Gomphoides (Gomphidae) Aphylla
williamsoni Muttkowski Epitheca (Corduliidae) Epitheca costalis

ODONATA: SPECIES INDEX

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