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ODONATA
ROSSER W. GARRISON
RESEARCH ASSOCIATE
NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY
900 EXPOSITION BOULEVARD
LOS ANGELES, CALIFORNIA 90007
552 NOMINA INSECTA NEARCTICA
NOMINA INSECTA NEARCTICA 553
ODONATA: INTRODUCTION
554 NOMINA INSECTA NEARCTICA
STATISTICS
This section contains some raw statistics without comment. A list of the species group
names proposed by decade is listed in the table below followed by a graph of the
numbers. Allowance must be made in interpreting the graph for the truncated decades at
the beginning and end of the graph. The tables on the following page are a break down of
the number of valid species and names by family.
1758 - 1759 3
1760 - 1769 1
1770 - 1779 17
1780 - 1789 1
1790 - 1799 6
1800 - 1809 2
1810 - 1819 0
1820 - 1829 0
1830 - 1839 71
1840 - 1849 53
1850 - 1859 57
1860 - 1869 153
1870 - 1879 72
1880 - 1889 24
1890 - 1899 56
1900 - 1909 56
1910 - 1919 53
1920 - 1929 19
1930 - 1939 18
1940 - 1949 17
1950 - 1959 16
1960 - 1969 6
1970 - 1979 3
1980 - 1989 13
1990 - 10
160
140
120
100
80
60
40
20
0
1758- 1780- 1810- 1840- 1870- 1900- 1930- 1960- 1990
1789 1819 1849 1879 1909 1939 1969 -
ODONATA: STATISTICS
556 NOMINA INSECTA NEARCTICA
ODONATA: STATISTICS
GarrisonNew World Odonata List (NWOL)
CLASSIFICATION: ODONATA
The classification adopted here, with the exception of the Gomphidae, is largely traditional and is based primarily
on: Fraser (1957. A reclassification of the Order Odonata. Royal Zoological Society of New South Wales,
133pp). The classification of the Gomphidae follows: Carle (1986. The classification, phylogeny and biogeography
of the Gomphidae (Anisoptera). I. Classification. Odonatologica 15(3): 275-326). Parts of the classification are
unsatisfactory and phylogenetic studies will alter the classification considerably. Such a new classification has
recently been proposed by Bechly (1996. Morphologische Untersuchungen am Flgelgeder der rezenten Libeeen
und deren Stammgruppenvertreter (insecta; Pterygota; Odonata). Petalura, spec.-vol. 2: 402 pp., 111 figs., 3 tabs.
[revised second edition with an English appendix: "Phylogenetic System and List of Synapomorphies"]). Drastic
changes in the higher classification are advocated. An abstracted English version (The phylogenetic system of fossil
and recent Odonatoptera).is available at the following web site:
http://members.aol.com/odonatadat/phylogeny/bechly.htm
1. Date of publication of Hagen and Se1ys, Monographie des Gomphines: I follow Calvert (1905. Biologia Centrali
Americana: 148), who states in a footnote: "It is usual to quote the "Monographie des Gomphines" with the date
1857, as is done by Kirby [1890] for example, and "November 1857" is, indeed, that which appears on the title
page of the volume; but the original paper cover bore the date "Mai 1858," which has usually not been preserved in
bound copies. Moreover, plates xx-xxii are dated at the bottom as having been drawn in 1858." Cowley (1937. J.
Soc. Bibliography Nat. Hist. 1(3):73-74) opted for 1857, but further evidence of an 1858 publication date was
brought to my attention by P. Machet. In the "appendice" (page 651 [391 separate]) is stated: "Some
circumstances having put off the conclusions of this volume, considerable and rather numerous materials have
reached us during printing [italics mine]. One comes from a trip made by H. Hagen to London in 1857; others
from a shipment of species from Texas, made by the Smithsonian Institution."
2. I give synonyms, nomina nuda, etc., only for New World taxa for Holarctic species such as Enallagma
cyathigerum and Libellula quadrimaculata. The single exception is Crocothemis servilia servilia, an Old World
species recently introduced into peninsular Florida.
Specific Comments:
1. Aeshna clepsydra with synonyms arundinacea Se1ys 1872 and maxima Heikal in Hisinger 1861. The latter
two names have often been included as synonyms of A. clepsydra. My reason for their deletion is based on a
footnote by Walker to Muttkowski (1910. Cat. Odon.: 109):
"Walker in litteris Aug. 14, 1909: 'The synonyms given by you do not belong to the true clepsydra. A.
arundinacea and maxima I have not seen: It is very probable that the first at least is a good species."' And,
2
GarrisonNew World Odonata List (NWOL)
"Walker in litteris, Sept. 24, 1909: 'Some of the references ... do not refer to true clepsydra...I am quite sure
arundinacea is not clepsydra, but as I have not seen the type it is probably best to let it stand as a synonym of
the latter on the authority of Martin'."
Curiously, I can find no mention of these little-used names in Walker's 1912 monograph. The original
description of arundinacea is based on a female from "Irkutzk", and Se1ys states it is similar to Siberian A.
juncea. According to Dr. Matti Hmlinen, University of Helsinki, in a letter to me, A. maxima is a junior
synonym of the Old World Aeshna crenata Hagen. I have found no records of clepsydra occurring anywhere in
the Palearctic, and I strongly suspect that arundinacea refers to a species other than the North American
clepsydra.
2. Triacanthagyna obscuripennis Blanchard. Williamson, in his monograph of this genus (1923. Univ. Mich.
Mus. Zool. Misc. Publ. 9:1-80), believed this to be a possible synonym of T. septima; but P. Machet believes,
after examining the short description and figure by Blanchard, that it may be of the trifida group, and possibly a
synonym of one of the later described species. Should this be the case (if the type is found), I would not be in
favor of using this name, I list obscuripennis as a distinct species, but it should, in my opinion, be considered a
nomen dubium or a nomen obletum, if and when its status is determined.
3. Gomphus complex. The status of the various genera/subgenera of North American entities has not been
satisfactorily resolved. I have not studied the problems, but have largely followed the advice of others. In this
list, I do not recognize the subgenus Stenogomphurus nor do I accept elevation of the subgenus Gomphurus to
generic status since there appears to be questions of their validity and changes in the literature as follows: Carle
(Carle, F. L. 1986. The classification, phylogeny and biogeography of the Gomphidae (Anisoptera). I.
Classification. Odonatologica 15(3): 275-326.) proposed the subgenus Stenogomphurus (type species:
Gomphus consanguis) under the large genus Gomphus. However, G. consanguis has always been included
under the subgenus Gomphurus of Gomphus. Carle (1986) also raises Gomphurus to generic status, but does
not key or diagnose between Gomphus and Gomphurus. He does not explain his reason for removing G.
consanguis from Gomphus and placing it in the new subgenus, Stenogomphurus. It would appear that he
never diagnosed Stenogomphurus from Gomphus. Carle (1986) does not discuss the relationship between
Gomphurus and Gomphus sensu lato, and gives no reason for the elevation of Gomphurus to generic status.
Therefore, I will continue to treat Gomphurus as a subgenus of Gomphus. Phanagomphus is also not
recognized in this list for the following reason: Carle (Carle, F. L. 1986. The classification, phylogeny and
biogeography of the Gomphidae (Anisoptera). I. Classification. Odonatologica 15(3): 275-326.) proposed the
subgenus Phanogomphus (type species: Gomphus minutus Rambur), which Carle & May (Carle, F. L. & M. L.
May. 1987. Gomphus (Phanogomphus) westfalli spec. nov. from the gulf coast of Florida (Anisoptera:
Gomphidae). Odonatologica 16(1): 67-75) later expanded to include other species of Gomphus (Gomphus).
Many Odonatists, including myself, consider Phanogomphus a subjective synonym of the type species of
Gomphus, G. vulgatissimus (Linnaeus). I treat Arigomphus as a separate genus from Gomphus.
Hylogomphus is considered a synonym of Gomphus (Gomphus), following Carle (1986). I realize that this
arrangement will probably not be satisfactory to many, but a final resolution may not be achieved for several
years.
Carle (in. May, M and F. Carle, 1996. An annotated list of the Odonata of New Jersey, with an appendix on
nomenclature in the genus Gomphus. Bulletin of Odonatology 4(1):1-35) has defended his actions regarding the
introduction of the names discussed above. That such an appendix was written and the acknowledgement that
several workers were in doubt about the proper introduction of new taxa and changes illustrates, in my opinion,
the continued confusion by this author's actions.
4. CORDULEGASTRIDAE: In a recent paper, 1992. Revision der Cordulegastridae. 1. Entwurfeiner neuen
Klassifizierung der Familie (Odonata: Anisoptera). Opuscula Zoologica Fluminensia 96:1-18, H. Lohmann has
performed a cladistic analysis on the family, with the result that our eight species are allocated to six genera as
follows:
3
GarrisonNew World Odonata List (NWOL)
While I admire the treatment of this family by Lohmann, I do not think it serves science to split our American
species from one into six genera. Carle, in an earlier paper (1983. Ann. Entomol. Soc. Amer. 76(1):61-68) also
recognized Zoraena and Taeniogaster, but his divisions do not seem to have been followed by Odonatists in this
country. I still consider all of our species in one genus, Cordulegaster. However, I include the above data for
those who wish to make changes.
5. Both Dr. Ken Tennessen and Mr. Carl Cook inform me that Neurocordulia clara Muttkowski is more closely
related to N. molesta, although Cook believes N. clara a good species. I have treated it as a synonym of N.
molesta.
6. Ken Tennessen (in litt.) informs me that he believes Epitheca (Tetragoneuria) williamsoni Muttkowski to be a
probable synonym of E. costalis.
7. Erythrodiplax connata complex. I include some nomenclatoral changes which are not in accord with Borror
1942 (Ohio State Univ. Contr. Zool. Entomol., No. 4). Other Odonatists and I treat E. minuscula and E. fusca
as separate species based on an absence of intergrades with E. connata (despite the fact that Borror reports such
specimens in his monograph)..
8. Libellula bifasciata Blanchard. The type for this old, forgotten name was discovered by Lieftinck to be L.
semifasciata, not L. pulchella, according to Dr. M.J. Westfall, Jr. (pers. comm.). As with Triacanthagyna
obscuripennis, I think it best to consider L. bifasciata a nomen obletum.
4
GarrisonNew World Odonata List (NWOL)
Williamson (1909 Proc. U. S. Nat. Museum 37:369-398.) reports captures of M. wabashensis with Macromia
taeniolata and M. pacifica, which suggests the possibility that M. wabashensis is a hybrid between M. pacifica
and M. taeniolata. Other mostly non-published data suggest the occurrence of both M. pacifica and M.
taeniolata wherever M. wabashensis has been collected. Like Dunkle (1995. Conservation of dragonflies
(Odonata) and their habitats in North America. page 23 in Corbet, P. S., S. W. Dunkle, H. Ubukata, eds.
Proceedings of the International symposium on the conservation of dragonflies and their habitats. Japanese
Society for the Preservation of Birds, Kushiro, xii + 1-70), I am doubtful of according species status to M.
wabashensis. Acquisition of more specimens and the discovery of the female may help to determine its true
status. I suggest that multivariate morphometric analyses of available specimens of M. wabashensis and samples
of M. taeniolata and M. pacifica from zones of sympatry with M. wabashensis may provide evidence about
whether M. wabashensis is a hybrid.
5
562 NOMINA INSECTA NEARCTICA
Calopteryx amata Hagen 1889 (Calopteryx) Argia bipustulata Kirby 1890 Missp.
Calopteryx angustipennis Selys 1853 (Sylphis) Argia cupreum Hagen 1861 (Agrion)
Sylphis elegans Hagen 1853 Syn. Argia emma Kennedy 1915 (Argia)
Calopteryx dimidiata Burmeister 1839 (Calopteryx) Argia extraneum Hagen 1861 (Agrion)
Calopteryx apicalis Burmeister 1839 Syn. Argia mista Navas 1934 Syn.
Calopteryx cognata Rambur 1842 Syn. Argia fumipennis Burmeister 1839 (Agrion)
Calopteryx maculata Beauvois 1805 (Agrion) Argia obscura Rambur 1842 Syn.
Calopteryx holosericea Burmeister 1839 Syn. Agrion violaceum Hagen 1861 Syn. (subspecies)
Calopteryx materna Say 1839 Syn. Argia atra Gloyd 1968 Syn. (subspecies)
Calopteryx opaca Say 1839 Syn. Argia hinei Kennedy 1918 (Argia)
Calopteryx papilionaria Rambur 1842 Syn. Argia immundum Hagen 1861 (Agrion)
Calopteryx floridana Huggins 1927 Syn. Argia lacrimans Hagen 1861 (Agrion)
Argia leonorae Garrison 1994 (Argia)
Hetaerina Hagen 1853 Argia lugens Hagen 1861 (Hyponeura)
Argia moestum Hagen 1861 (Agrion)
Hetaerina americana Fabricius 1798 (Agrion) Agrion putridum Hagen 1861 Syn.
Lestes basalis Say 1839 Syn. Argia intruda Williamson 1912 Syn.
Hetaerina basalis Hagen 1859 Homo. Argia munda Calvert 1902 (Argia)
Hetaerina californica Hagen 1859 Syn. Argia rita Kennedy 1919 Syn.
Hetaerina pseudamericana Walsh 1863 Syn. Argia nahuana Calvert 1902 (Argia)
Hetaerina sclerata Walsh 1863 Syn. Argia saalasi Valle 1942 Syn.
Hetaerina texana Walsh 1863 Syn. Argia oenea Hagen 1865 (Argia)
Hetaerina titia Drury 1773 (Libellula) Argia pallens Calvert 1902 (Argia)
Calopteryx tricolor Burmeister 1839 Syn. Argia solita Kennedy 1918 Syn.
Hetaerina septentrionalis Selys 1853 Syn. Argia pima Garrison 1994 (Argia)
Hetaerina limbata Selys 1853 Syn. Argia plana Calvert 1902 (Argia)
Hetaerina rupamnensis Walsh 1863 Syn. Argia spegazzinii Navas 1934 Syn.
Hetaerina rupinsulensis Walsh 1863 Syn. Argia rhoadsi Calvert 1902 (Argia)
Hetaerina bipartita Selys 1873 Syn. Argia sabino Garrison 1994 (Argia)
Hetaerina vulnerata Hagen 1853 (Hetaerina) Argia sedulum Hagen 1861 (Agrion)
Argia tarascana Calvert 1902 (Argia)
Argia tezpi Calvert 1902 (Argia)
Argia tibialis Rambur 1842 (Platycnemis)
Agrion fontium Hagen 1861 Syn.
COENAGRIONIDAE Agrion binotatum Walsh 1862 Syn.
Argia tonto Calvert 1902 (Argia)
Argia translata Hagen 1865 (Argia)
Argia espinalensis Navas 1935 Syn.
Acanthagrion Selys 1876 Argia vivida Hagen 1865 (Argia)
Myagrion Frster 1914 Syn. Argia kurilis Hagen 1865 Syn.
Cordulia shurtleffi Scudder 1866 (Cordulia) Neurocordulia alabamensis Hodges 1955 (Neurocordulia)
Cordulia bifurcata Hagen 1861 Unav. Neurocordulia molesta Walsh 1863 (Cordulia)
Neurocordulia clara Muttkowski 1910 Syn.
Didymops Rambur 1842 Neurocordulia obsoleta Say 1839 (Libellula)
Libellula polysticta Burmeister 1839 Syn.
Didymops floridensis Davis 1921 (Didymops) Neurocordulia virginiensis Davis 1927 (Neurocordulia)
Didymops transversa Say 1839 (Libellula) Neurocordulia xanthosoma Williamson 1908 (Platycordulia)
Epophthalmia cinnamomea Burmeister 1839 Syn. Neurocordulia yamaskanensis Provancher 1875 (Aeshna)
Didymops servillei Rambur 1842 Syn. Epitheca jamascarensis Hagen 1878 Syn.
ODONATA: LIBELLULIDAE
NOMINA INSECTA NEARCTICA 569
ODONATA: LIBELLULIDAE
570 NOMINA INSECTA NEARCTICA
PETALURIDAE
GENERIC INDEX
Dorocordulia Needham (Corduliidae)
Acanthagrion Selys (Coenagrionidae) Dromogomphus Selys (Gomphidae)
Acanthagyna Kirby (Aeshnidae) Gynacantha Dythemis Hagen (Libellulidae)
Adversaeschna Watson (Aeshnidae) Aeshna Enallagma Charpentier (Coenagrionidae)
Aenallagma Selys (Coenagrionidae) Enallagma Eolibellula Kennedy (Libellulidae) Libellula
Aeshna Fabricius (Aeshnidae) Eoprogomphus Carle (Gomphidae) Progomphus
Africalestes Kennedy (Lestidae) Lestes Ephidatia Kirby (Libellulidae) Idiataphe
Africallagma Kennedy (Coenagrionidae) Enallagma Epiaeschna Hagen (Aeshnidae)
Agrion Fabricius (Calopterygidae) Calopteryx Epicordulia Selys (Corduliidae) Epitheca
Agrion Leach (Coenagrionidae) Coenagrion Epitheca Burmeister (Corduliidae)
Aino Kirby (Libellulidae) Nannothemis Erpetocyclops Carle (Gomphidae) Erpetogomphus
Alloprogomphus Carle (Gomphidae) Progomphus Erpetogomphus Selys (Gomphidae)
Amphiagrion Selys (Coenagrionidae) Erythemis Hagen (Libellulidae)
Amphiallagma Kennedy (Coenagrionidae) Enallagma Erythragrion Selys (Coenagrionidae) Telebasis
Anaciagrion Kennedy (Calopterygidae) Calopteryx Erythrodiplax Brauer (Libellulidae)
Anapates Charpentier (Lestidae) Lestes Eurothemis Kennedy (Libellulidae) Libellula
Anax Leach (Aeshnidae)
Anomalagrion Selys (Coenagrionidae) Ischnura Fonscolombia Selys (Aeshnidae) Boyeria
Anomalura Kennedy (Coenagrionidae) Ischnura
Apanisagrion Kennedy (Coenagrionidae) Gomphaeschna Selys (Aeshnidae)
Aphylla Selys (Gomphidae) Gomphoides Selys (Gomphidae) Progomphus
Archaeoprogomphus Carle (Gomphidae) Progomphus Gomphurus Needham (Gomphidae) Gomphus
Archegaster Lohmann (Cordulegastridae) Cordulegaster Gomphus Leach (Gomphidae)
Archilestes Selys (Lestidae) Gynacantha Rambur (Aeshnidae)
Argia Rambur (Coenagrionidae)
Argiallagma Calvert (Coenagrionidae) Nehalennia Hagenius Selys (Gomphidae)
Argiallagma Selys (Coenagrionidae) Nehalennia Heliargia Kennedy (Coenagrionidae) Argia
Arigomphus Needham (Gomphidae) Helocordulia Needham (Corduliidae)
Austrocoenagrion Kennedy (Coenagrionidae) Coenagrion Herpetogomphus Walsh (Gomphidae) Erpetogomphus
Hesperaeschna Cockerell (Aeshnidae) Aeshna
Basiaeschna Selys (Aeshnidae) Hesperagrion Calvert (Coenagrionidae)
Beblecia Kirby (Libellulidae) Crocothemis Hetaerina Hagen (Calopterygidae)
Bedfordia Mumford (Coenagrionidae) Ischnura Holotania Kirby (Libellulidae) Libellula
Belonia Kennedy (Libellulidae) Libellula Hoplonaeschna Karsch (Aeshnidae) Oplonaeschna
Boyeria McLachlan (Aeshnidae) Hylaeschna Sjstedt (Corduliidae) Macromia
Brachymesia Kirby (Libellulidae) Hylogomphus Needham (Gomphidae) Gomphus
Brechmorhoga Kirby (Libellulidae) Hylogomphus Needham and Westfall (Gomphidae) Gomphus
Hyponeura Selys (Coenagrionidae) Argia
Caenoneura Kirby (Protoneuridae) Neoneura
Calogomphus Carle (Gomphidae) Erpetogomphus Icterolestes Pinhey (Lestidae) Lestes
Calopteryx Leach (Calopterygidae) Idiataphe Cowley (Libellulidae)
Cannacria Kirby (Libellulidae) Brachymesia Ischnallagma Kennedy (Coenagrionidae) Enallagma
Cannaphila Kirby (Libellulidae) Ischnura Charpentier (Coenagrionidae)
Celaenura Kennedy (Coenagrionidae) Ischnura Ischnuridia Donnelly (Coenagrionidae) Ischnura
Celithemis Hagen (Libellulidae)
Cendra Navas (Libellulidae) Macrothemis Kalosympetrum Carle (Libellulidae) Sympetrum
Ceratopyga Nunney (Corduliidae) Macromia Kalyptogaster Lohmann (Cordulegastridae) Cordulegaster
Chalcargia Kennedy (Coenagrionidae) Argia Kuldanagaster Yousuf and Yunus (Cordulegastridae) Cordulegaster
Chalcolestes Kennedy (Lestidae) Lestes
Chloragrion Hartinov and Hartinova (Coenagrionidae) Coenagrion Ladona Needham (Libellulidae) Libellula
Chlorosoma Charpentier (Corduliidae) Somatochlora Lanthus Needham (Gomphidae)
Chromagrion Needham (Coenagrionidae) Lauragaster Lohmann (Cordulegastridae) Cordulegaster
Cnemophila Wallengren (Aeshnidae) Aeshna Leptargia Kennedy (Coenagrionidae) Argia
Coenagrion Kirby (Coenagrionidae) Leptetrum Newman (Libellulidae) Libellula
Coenotiata Buchecker (Libellulidae) Leucorrhinia Lepthemis Hagen (Libellulidae) Erythemis
Cordulegaster Leach (Cordulegastridae) Lestes Leach (Lestidae)
Cordulia Leach (Corduliidae) Leucorrhinia Brittinger (Libellulidae)
Coryphaeschna Williamson (Aeshnidae) Libellula Linnaeus (Libellulidae)
Crocothemis Brauer (Libellulidae)
Cyanargia Kennedy (Coenagrionidae) Argia Macrodiplax Brauer (Libellulidae)
Cyptolestes Williamson (Lestidae) Archilestes Macromia Rambur (Corduliidae)
Macrothemis Hagen (Libellulidae)
Diargia Calvert (Coenagrionidae) Argia Malgassolestes Pinhey (Lestidae) Lestes
Didymops Rambur (Corduliidae) Marmaraeschna Calvert (Aeshnidae) Aeshna
Diplax Burmeister (Libellulidae) Sympetrum Mesothemis Hagen (Libellulidae) Erythemis
Miathyria Kirby (Libellulidae)
SPECIES INDEX
aspersum Hagen Agrion (Coenagrionidae) Enallagma
aspersus Morse Ophiogomphus (Gomphidae) Ophiogomphus
a assimilata Uhler Libellula (Libellulidae) Sympetrum rubicundula
atra Gloyd Argia (Coenagrionidae) Argia fumipennis
aaroni Calvert Neoneura (Protoneuridae) Neoneura atripes Hagen Diplax (Libellulidae) Sympetrum costifera
abboti Hagen Aeschna (Aeshnidae) Coryphaeschna ingens attala Selys Libellula (Libellulidae) Erythemis
abbreviatum Selys Pyrrhosoma (Coenagrionidae) Amphiagrion aurantiaca Hagen Agrion (Coenagrionidae) Ischnura ramburii
abbreviatus Hagen Gomphus (Gomphidae) Gomphus auripennis Burmeister Libellula (Libellulidae) Libellula
abditus Butler Gomphus (Gomphidae) Stylurus amnicola australensis Williamson Macromia (Corduliidae) Macromia
abdominalis Rambur Libellula (Libellulidae) Tramea illinoiensis
acuminatus Carle Ophiogomphus (Gomphidae) Ophiogomphus
acuta Say Libellula (Libellulidae) Erythemis vesiculosa australis Carle Ophiogomphus (Gomphidae) Ophiogomphus
adelphus Selys Gomphus (Gomphidae) Gomphus australis Hagen Tramea (Libellulidae) Tauriphila
adnexa Hagen Aeschna (Aeshnidae) Coryphaeschna australis Kirby Brachymesia (Libellulidae) Brachymesia furcata
aequabilis Say Calopteryx (Calopterygidae) Calopteryx australis Needham Arigomphus (Gomphidae) Gomphus
aequalis Hagen Dythemis (Libellulidae) Micrathyria australis Walker Lestes (Lestidae) Lestes disjuncta
affinis Kirby Neurothemis (Libellulidae) Erythrodiplax funerea axilena Westwood Libellula (Libellulidae) Libellula
agrioides Calvert Argia (Coenagrionidae) Argia azteca Calvert Tauriphila (Libellulidae) Tauriphila
alabamensis Hodges Neurocordulia (Corduliidae) Neurocordulia
alacer Hagen Lestes (Lestidae) Lestes
alachuensis Byers Progomphus (Gomphidae) Progomphus
b
alberta Kennedy Argia (Coenagrionidae) Argia
balteata Hagen Tetragoneuria (Libellulidae) Macrodiplax
albicincta Burmeister Epophthalmia (Corduliidae) Somatochlora
barberi Currie Ischnura (Coenagrionidae) Ischnura
albifrons Charpentier Libellula (Libellulidae) Sympetrum ambigua
barretti Calvert Argia (Coenagrionidae) Argia
albistylus Hagen Gomphus (Gomphidae) Stylogomphus
basalis Burmeister Libellula (Libellulidae) Tramea abdominalis
albrighti Needham Gomphoides (Gomphidae) Phyllogomphoides
basalis Hagen Hetaerina (Calopterygidae) Hetaerina americana
aliasignata Muttkowski Libellula (Libellulidae) Libellula
basalis Say Lestes (Calopterygidae) Hetaerina americana
croceipennis
basalis Say Libellula (Libellulidae) Libellula luctuosa
alleghaniensis Carle Ophiogomphus (Gomphidae) Ophiogomphus
basidens Calvert Enallagma (Coenagrionidae) Enallagma
incurvatus
basifusca Calvert Trithemis (Libellulidae) Erythrodiplax connata
alleghaniensis Williamson Macromia (Corduliidae) Macromia
basiguttata Selys Cordulia (Corduliidae) Epitheca cynosura
alleni Howe Gomphus (Gomphidae) Gomphus quadricolor
batesi Kirby Cannacria (Libellulidae) Brachymesia herbida
amanda Hagen Diplax (Libellulidae) Celithemis
bella Uhler Nannophya (Libellulidae) Nannothemis
amata Hagen Calopteryx (Calopterygidae) Calopteryx
bellei Knopf and Tennessen Progomphus (Gomphidae) Progomphus
amazili Burmeister Aeschna (Aeshnidae) Anax
berenice Drury Libellula (Libellulidae) Erythrodiplax
ambigua Rambur Libellula (Libellulidae) Sympetrum
bertha Williamson Celithemis (Libellulidae) Celithemis
ambigua Selys Gomphoides (Gomphidae) Aphylla protracta
bifasciata Fabricius Libellula (Libellulidae) Libellula semifasciata
ambusta Hagen Diplax (Libellulidae) Erythrodiplax justiniana
bifurcata Hagen Cordulia (Corduliidae) Cordulia shurtleffi
americana Bartenef Aeshna (Aeshnidae) Aeshna juncea
bilineata Carle Zoraena (Cordulegastridae) Cordulegaster
americana Fabricius Agrion (Calopterygidae) Hetaerina
binotata Rambur Libellula (Libellulidae) Tramea
amnicola Walsh Gomphus (Gomphidae) Stylurus
binotatum Walsh Agrion (Coenagrionidae) Argia tibialis
amphion Selys Amphiagrion (Coenagrionidae) Amphiagrion
bipartita Selys Hetaerina (Calopterygidae) Hetaerina titia
saucium
bipunctulatum Hagen Agrion (Coenagrionidae) Argia
analis Burmeister Libellula (Libellulidae) Pantala flavescens
bipustulata Kirby Argia (Coenagrionidae) Argia bipunctulatum
angulatum Walker Coenagrion (Coenagrionidae) Coenagrion
bison Selys Ophiogomphus (Gomphidae) Ophiogomphus
angustifolia Garrison Aphylla (Gomphidae) Aphylla
bistigma Uhler Libellula (Libellulidae) Libellula cyanea
angustipennis Selys Sylphis (Calopterygidae) Calopteryx
boreale Selys Enallagma (Coenagrionidae) Enallagma
anna Williamson Enallagma (Coenagrionidae) Enallagma
borealis Hagen Leucorrhinia (Libellulidae) Leucorrhinia
annexum Hagen Agrion (Coenagrionidae) Enallagma cyathigerum
borealis McLachlan Progomphus (Gomphidae) Progomphus
annulata Hagen Macromia (Corduliidae) Macromia
borealis Needham Gomphus (Gomphidae) Gomphus
annulata Rambur Libellula (Libellulidae) Erythemis attala
boucardi Selys Telebasis (Coenagrionidae) Telebasis salvum
annulosa Selys Libellula (Libellulidae) Erythemis attala
bouchardi Louton Ophiogomphus (Gomphidae) Ophiogomphus
anomalum Rambur Agrion (Coenagrionidae) Ischnura hastata
acuminatus
anomalus Harvey Ophiogomphus (Gomphidae) Ophiogomphus
brasiliana Brauer Tramea (Libellulidae) Tramea binotata
antennatum Say Agrion (Coenagrionidae) Enallagma
brevicincta Robert Somatochlora (Corduliidae) Somatochlora
antilope Hagen Aeschna (Aeshnidae) Gomphaeschna
brevis Hagen Gomphus (Gomphidae) Gomphus adelphus
apicale Say Agrion (Coenagrionidae) Argia
brevistylus Selys Hagenius (Gomphidae) Hagenius
apicalis Burmeister Calopteryx (Calopterygidae) Calopteryx
brimleyi Muttkowski Gomphus (Gomphidae) Gomphus cavillaris
dimidiata
byersi Westfall Telebasis (Coenagrionidae) Telebasis
apomyius Donnelly Gomphus (Gomphidae) Gomphus
argus Needham Gomphus (Gomphidae) Gomphus descriptus
arida Kennedy Aeshna (Aeshnidae) Aeshna palmata c
arizonicus Kennedy Ophiogomphus (Gomphidae) Ophiogomphus
armata Hagen Aeschna (Aeshnidae) Oplonaeschna caderita Needham Macromia (Corduliidae) Macromia annulata
armatus Selys Dromogomphus (Gomphidae) Dromogomphus caerulans Rambur Libellula (Libellulidae) Erythemis simplicicollis
californica Calvert Aeshna (Aeshnidae) Aeshna constricta Say Aeshna (Aeshnidae) Aeshna
californica Hagen Hetaerina (Calopterygidae) Hetaerina americana cornutus Tough Gomphus (Gomphidae) Arigomphus
californica Kennedy Calopteryx (Calopterygidae) Calopteryx corrupta Hagen Mesothemis (Libellulidae) Sympetrum
aequabilis costalis Rambur Libellula (Libellulidae) Libellula auripennis
californica Martin Perithemis (Libellulidae) Perithemis intensa costalis Selys Cordulia (Corduliidae) Epitheca
californica McLachlan Archilestes (Lestidae) Archilestes costifera Hagen Diplax (Libellulidae) Sympetrum
californicum Walker Sympetrum (Libellulidae) Sympetrum crassus Hagen Gomphus (Gomphidae) Gomphus
occidentale credulum Hagen Agrion (Coenagrionidae) Ischnura ramburii
californicus Kennedy Ophiogomphus (Gomphidae) Ophiogomphus croceipennis Selys Libellula (Libellulidae) Libellula
occidentis crotalinus Hagen Ophiogomphus (Gomphidae) Erpetogomphus
calverti Morse Enallagma (Coenagrionidae) Enallagma boreale cubensis Scudder Macromia (Libellulidae) Idiataphe
calverti Muttkowski Epitheca (Corduliidae) Epitheca semiaquea culicinorum Byers Enallagma (Coenagrionidae) Enallagma anna
calverti Muttkowski Tramea (Libellulidae) Tramea cultellatum Selys Enallagma (Coenagrionidae) Neoerythromma
calverti Williamson and Gloyd Somatochlora (Corduliidae) cupreum Hagen Agrion (Coenagrionidae) Argia
Somatochlora cyanea Fabricius Libellula (Libellulidae) Libellula
camilla Rambur Libellula (Libellulidae) Celithemis eponina cyanifrons Hagen Aeschna (Aeshnidae) Coryphaeschna adnexa
canadense Provancher Agrion (Coenagrionidae) Ischnura verticalis cyathigerum Charpentier Agrion (Coenagrionidae) Enallagma
canadensis Walker Aeshna (Aeshnidae) Aeshna cynosura Say Libellula (Corduliidae) Epitheca
canis McLachlan Epitheca (Corduliidae) Epitheca
cara Calvert Protoneura (Protoneuridae) Protoneura
cardenium Hagen Enallagma (Coenagrionidae) Enallagma coecum d
carolina Linnaeus Libellula (Libellulidae) Tramea
carolinus Carle Gomphus (Gomphidae) Gomphus parvidens daeckii Calvert Telagrion (Coenagrionidae) Enallagma
carolinus Hagen Ophiogomphus (Gomphidae) Ophiogomphus damula Calvert Ischnura (Coenagrionidae) Ischnura
rupinsulensis danae Sulzer Libellula (Libellulidae) Sympetrum
davisi Westfall Enallagma (Coenagrionidae) Enallagma
carolott Butler Nehalennia (Coenagrionidae) Nehalennia irene decisa Hagen Diplax (Libellulidae) Sympetrum obtrusa
carolus Needham Ophiogomphus (Gomphidae) Ophiogomphus defixum Hagen Agrion (Coenagrionidae) Ischnura ramburii
carunculatum Morse Enallagma (Coenagrionidae) Enallagma demorsum Hagen Agrion (Coenagrionidae) Ischnura
cavillaris Needham Gomphus (Gomphidae) Gomphus denticollis Burmeister Agrion (Coenagrionidae) Ischnura
cervula Selys Ischnura (Coenagrionidae) Ischnura dentiferum Walsh Agrion (Coenagrionidae) Enallagma signatum
chalybea Hagen Cordulia (Corduliidae) Somatochlora forcipata deplanata Rambur Libellula (Libellulidae) Libellula
charadraea Williamson Somatochlora (Corduliidae) Somatochlora descriptus Banks Gomphus (Gomphidae) Gomphus
ensigera
chlora Rambur Libellula (Libellulidae) Perithemis tenera deserticola Cruden Cordulegaster (Cordulegastridae)
chloropleura Brauer Libellula (Libellulidae) Erythrodiplax connata Cordulegaster dorsalis
chrysoptera Selys Diplax (Libellulidae) Sympetrum madida designatus Hagen Erpetogomphus (Gomphidae) Erpetogomphus
cingulata Selys Epitheca (Corduliidae) Somatochlora diadema Selys Cordulegaster (Cordulegastridae) Cordulegaster
cinnamomea Burmeister Epophthalmia (Corduliidae) Didymops diadophis Calvert Erpetogomphus (Gomphidae) Erpetogomphus
transversa eutainia
citrina Hagen Tholymis (Libellulidae) Tholymis diastatops Selys Thecaphora (Cordulegastridae) Cordulegaster
civile Hagen Agrion (Coenagrionidae) Enallagma dicrota Hagen Dythemis (Libellulidae) Micrathyria didyma
clara Muttkowski Neurocordulia (Corduliidae) Neurocordulia didyma Selys Libellula (Libellulidae) Micrathyria
molesta diffinis Hagen Epitheca (Corduliidae) Epitheca semiaquea
clausum Morse Enallagma (Coenagrionidae) Enallagma diffinis Selys Cordulia (Corduliidae) Epitheca cynosura
clepsydra Say Aeshna (Aeshnidae) Aeshna dilatatus Rambur Gomphus (Gomphidae) Gomphus
cloe Hagen Perithemis (Libellulidae) Perithemis mooma dimidiata Burmeister Calopteryx (Calopterygidae) Calopteryx
coecum Hagen Enallagma (Coenagrionidae) Enallagma diminutus Needham Gomphus (Gomphidae) Gomphus
cognata Rambur Calopteryx (Calopterygidae) Calopteryx dimidiata discolor Burmeister Agrion (Coenagrionidae) Amphiagrion saucium
collocata Hagen Mesothemis (Libellulidae) Erythemis discolor Burmeister Libellula (Libellulidae) Orthemis ferruginea
coloradica Cockerell Calopteryx (Calopterygidae) Calopteryx disjuncta Selys Lestes (Lestidae) Lestes
aequabilis divagans Selys Enallagma (Coenagrionidae) Enallagma
coluber Williamson and Williamson Erpetogomphus (Gomphidae) domina Calvert Palaemnema (Platystictidae) Palaemnema
Erpetogomphus compositus dominicana Hagen Aeshna (Aeshnidae) Aeshna psilus
colubrinus Selys Ophiogomphus (Gomphidae) Ophiogomphus domitia Drury Libellula (Libellulidae) Perithemis
comanche Calvert Libellula (Libellulidae) Libellula donneri Kennedy Gomphus (Gomphidae) Gomphus kurilis
communis Rambur Libellula (Libellulidae) Erythrodiplax connata dorsalis Hagen Cordulegaster (Cordulegastridae) Cordulegaster
complanata Rambur Cordulia (Corduliidae) Epitheca cynosura doubledayi Selys Agrion (Coenagrionidae) Enallagma
composita Hagen Mesothemis (Libellulidae) Libellula dryas Kirby Lestes (Lestidae) Lestes
compositus Hagen Erpetogomphus (Gomphidae) Erpetogomphus dubium Root Enallagma (Coenagrionidae) Enallagma
concisum Williamson Enallagma (Coenagrionidae) Enallagma dugesi Calvert Aeshna (Aeshnidae) Aeshna
conditum Selys Erythromma (Coenagrionidae) Chromagrion durum Hagen Agrion (Coenagrionidae) Enallagma
confraternus Selys Gomphus (Gomphidae) Gomphus kurilis
confusa Uhler Libellula (Libellulidae) Libellula pulchella
congener Hagen Lestes (Lestidae) Lestes
e
connata Burmeister Libellula (Libellulidae) Erythrodiplax
ebrium Hagen Agrion (Coenagrionidae) Enallagma
consanguis Selys Gomphus (Gomphidae) Gomphus
edmundo Needham Ophiogomphus (Gomphidae) Ophiogomphus
consobrinus Walsh Gomphus (Gomphidae) Gomphus externus
elegans Hagen Sylphis (Calopterygidae) Calopteryx angustipennis furcata Hagen Erythemis (Libellulidae) Brachymesia
elisa Hagen Diplax (Libellulidae) Celithemis furcifer Hagen Gomphus (Gomphidae) Arigomphus
elongata Scudder Cordulia (Corduliidae) Somatochlora furcifera Karsch Aeshna (Aeshnidae) Aeshna multicolor
elongatus Selys Gomphus (Gomphidae) Stylurus plagiatus furcillata Say Aeschna (Aeshnidae) Gomphaeschna
emma Kennedy Argia (Coenagrionidae) Argia fusca Rambur Libellula (Libellulidae) Erythrodiplax
ensigera Martin Somatochlora (Corduliidae) Somatochlora fuscofasciata Blanchard Libellula (Libellulidae) Erythrodiplax
eponina Drury Libellula (Libellulidae) Celithemis umbrata
eremita Scudder Aeshna (Aeshnidae) Aeshna
eremita Scudder Cordulia (Corduliidae) Somatochlora albicincta
erratica Calvert Ischnura (Coenagrionidae) Ischnura g
erroneus Hagen Cordulegaster (Cordulegastridae) Cordulegaster
espinalensis Navas Argia (Coenagrionidae) Argia translata gemina Kennedy Celaenura (Coenagrionidae) Ischnura
eurinus Say Lestes (Lestidae) Lestes geminatum Kellicott Enallagma (Coenagrionidae) Enallagma
eutainia Calvert Erpetogomphus (Gomphidae) Erpetogomphus geminatus Carle Gomphus (Gomphidae) Gomphus
exclamationis Selys Agrion (Coenagrionidae) Zoniagrion georgiana Walker Somatochlora (Corduliidae) Somatochlora
exilis Selys Gomphus (Gomphidae) Gomphus georgina Selys Epophthalmia (Corduliidae) Macromia illinoiensis
expertum Hagen Agrion (Coenagrionidae) Ischnura ramburii glacialis Hagen Leucorrhinia (Libellulidae) Leucorrhinia
exstriata Calvert Ischnura (Coenagrionidae) Ischnura denticollis gonzalezi Dunkle Gomphus (Gomphidae) Gomphus
exsulans Hagen Agrion (Coenagrionidae) Enallagma gracilis Morse Nehalennia (Coenagrionidae) Nehalennia
externus Hagen Gomphus (Gomphidae) Gomphus grafiana Williamson Boyeria (Aeshnidae) Boyeria
extraneum Hagen Agrion (Coenagrionidae) Argia grandis Rambur Lestes (Lestidae) Archilestes
exusta Say Libellula (Libellulidae) Libellula graslinellus Walsh Gomphus (Gomphidae) Gomphus
gravida Calvert Lepthemis (Libellulidae) Brachymesia
gundlachi Scudder Mesothemis (Libellulidae) Erythemis
f simplicicollis
guyanensis Machet Coryphaeschna (Aeshnidae) Coryphaeschna
fallax Burmeister Libellula (Libellulidae) Erythrodiplax umbrata adnexa
fasciata Kirby Celithemis (Libellulidae) Celithemis
fasciatum Walker Sympetrum (Libellulidae) Sympetrum occidentale
fasciatus Rambur Cordulegaster (Cordulegastridae) Cordulegaster h
obliqua
fastigiatus Donnelly Ophiogomphus (Gomphidae) Ophiogomphus hageni Calvert Leucorrhinia (Libellulidae) Leucorrhinia hudsonica
mainensis hageni Selys Tachopteryx (Petaluridae) Tanypteryx
ferruginea Fabricius Libellula (Libellulidae) Crocothemis servilia hageni Walsh Agrion (Coenagrionidae) Enallagma
ferruginea Fabricius Libellula (Libellulidae) Orthemis hagenii Kirby Micrathyria (Libellulidae) Micrathyria
filosa Hagen Cordulia (Corduliidae) Somatochlora hamata Hagen Lestes (Lestidae) Lestes forcipata
fischeri Kellicott Enallagma (Coenagrionidae) Enallagma hastata Say Agrion (Coenagrionidae) Ischnura
antennatum hastulatum Stephens Agrion (Coenagrionidae) Enallagma
flavescens Fabricius Libellula (Libellulidae) Pantala cyathigerum
flavescens Selys Agrion (Coenagrionidae) Hesperagrion herbida Gundlach Libellula (Libellulidae) Brachymesia
heterodoxum heros Fabricius Aeschna (Aeshnidae) Epiaeschna
flavicans Rambur Libellula (Libellulidae) Erythrodiplax umbrata hersilia Blanchard Libellula (Libellulidae) Libellula semifasciata
flavicosta Hagen Diplax (Libellulidae) Sympetrum madida heterodon Garrison Erpetogomphus (Gomphidae) Erpetogomphus
flavida Hagen Libellula (Libellulidae) Libellula comanche heterodoxum Selys Amphiagrion (Coenagrionidae) Hesperagrion
flavida Rambur Libellula (Libellulidae) Libellula hineana Williamson Somatochlora (Corduliidae) Somatochlora
flavipennis Walsh Macromia (Corduliidae) Macromia pacifica hinei Kennedy Argia (Coenagrionidae) Argia
flavocaudatus Walker Gomphus (Gomphidae) Gomphus exilis histrio Burmeister Libellula (Libellulidae) Erythrodiplax berenice
fletcheri Williamson Williamsonia (Corduliidae) Williamsonia hodgesi Needham Gomphus (Gomphidae) Gomphus
florida Hagen Aeschna (Aeshnidae) Remartinia luteipennis holosericea Burmeister Calopteryx (Calopterygidae) Calopteryx
floridana Huggins Calopteryx (Calopterygidae) Calopteryx maculata
maculata howei Bromley Ophiogomphus (Gomphidae) Ophiogomphus
floridensis Davis Didymops (Corduliidae) Didymops huanacina Frster Tramea (Libellulidae) Pantala hymenaea
fluvialis Walsh Gomphus (Gomphidae) Stylurus notatus hudsonica Hagen Calopteryx (Calopterygidae) Calopteryx
foliata Kirby Belonia (Libellulidae) Libellula aequabilis
fontium Hagen Agrion (Coenagrionidae) Argia tibialis hudsonica Hagen Epitheca (Corduliidae) Somatochlora
forcipata Rambur Lestes (Lestidae) Lestes hudsonica Selys Aeshna (Aeshnidae) Aeshna eremita
forcipata Scudder Cordulia (Corduliidae) Somatochlora hudsonica Selys Libellula (Libellulidae) Leucorrhinia
forensis Hagen Libellula (Libellulidae) Libellula hybridus Williamson Gomphus (Gomphidae) Gomphus
forficula Rambur Lestes (Lestidae) Lestes hymenaea Say Libellula (Libellulidae) Pantala
franklini Selys Epitheca (Corduliidae) Somatochlora
fraterna Hagen Diplax (Libellulidae) Erythrodiplax connata
fraterna Say Aeshna (Gomphidae) Gomphus
i
frigida Hagen Leucorrhinia (Libellulidae) Leucorrhinia
fugax Hagen Dythemis (Libellulidae) Dythemis
illinoiensis Walsh Macromia (Corduliidae) Macromia
fumipennis Burmeister Agrion (Coenagrionidae) Argia
illota Hagen Mesothemis (Libellulidae) Sympetrum
fumipennis Currie Cannacria (Libellulidae) Brachymesia herbida
imbuta Say Libellula (Libellulidae) Erythemis simplicicollis
funerea Carpenter Misagria (Libellulidae) Cannaphila insularis
imitans Karsch Macrothemis (Libellulidae) Macrothemis
funerea Hagen Libellula (Libellulidae) Erythrodiplax
immundum Hagen Agrion (Coenagrionidae) Argia
inacuta Calvert Macrothemis (Libellulidae) Macrothemis leontina Brauer Libellula (Libellulidae) Erythrodiplax connata
inaequalis Walsh Lestes (Lestidae) Lestes leonura Westfall Celithemis (Libellulidae) Celithemis bertha
incesta Hagen Libellula (Libellulidae) Libellula lepida Hagen Cordulia (Corduliidae) Dorocordulia
incompta Rambur Libellula (Libellulidae) Erythrodiplax fusca leucozona Ris Macrothemis (Libellulidae) Macrothemis imitans
incurvata Walker Somatochlora (Corduliidae) Somatochlora libera Selys Cordulia (Corduliidae) Dorocordulia
incurvatus Carle Ophiogomphus (Gomphidae) Ophiogomphus limbata Selys Hetaerina (Calopterygidae) Hetaerina titia
indistincta Morse Epitheca (Corduliidae) Epitheca spinigera linearis Hagen Cordulia (Corduliidae) Somatochlora
inequiunguis Calvert Macrothemis (Libellulidae) Macrothemis lineata Walker Aeshna (Aeshnidae) Aeshna interrupta
iners Hagen Agrion (Coenagrionidae) Ischnura ramburii
ingens Rambur Aeschna (Aeshnidae) Coryphaeschna lineatifrons Calvert Gomphus (Gomphidae) Gomphus
insularis Hagen Tramea (Libellulidae) Tramea lineatipes Karsch Paltothemis (Libellulidae) Paltothemis
insularis Kirby Cannaphila (Libellulidae) Cannaphila lintneri Hagen Cordulia (Corduliidae) Williamsonia
intacta Hagen Diplax (Libellulidae) Leucorrhinia lividus Selys Gomphus (Gomphidae) Gomphus
integricollis Calvert Nehalennia (Coenagrionidae) Nehalennia longicauda Brauer Tramea (Libellulidae) Tramea binotata
intensa Kirby Perithemis (Libellulidae) Perithemis longipennis Burmeister Libellula (Libellulidae) Pachydiplax
interna Walker Aeshna (Aeshnidae) Aeshna interrupta longipes Hagen Anax (Aeshnidae) Anax
internum Montgomery Sympetrum (Libellulidae) Sympetrum lucilla Rambur Libellula (Libellulidae) Celithemis eponina
interrogatum Selys Agrion (Coenagrionidae) Coenagrion luctuosa Burmeister Libellula (Libellulidae) Libellula
interrupta Walker Aeshna (Aeshnidae) Aeshna lugens Hagen Hyponeura (Coenagrionidae) Argia
intricatus Hagen Gomphus (Gomphidae) Stylurus luteipennis Burmeister Aeschna (Aeshnidae) Remartinia
intruda Williamson Argia (Coenagrionidae) Argia moestum lydia Drury Libellula (Libellulidae) Libellula
iphigenia Hagen Tramea (Libellulidae) Tauriphila australis lynnae Paulson Gomphus (Gomphidae) Gomphus
irene Hagen Agrion (Coenagrionidae) Nehalennia
iris Hagen Perithemis (Libellulidae) Perithemis domitia
ivae Williamson Stylurus (Gomphidae) Stylurus m
macromia Brauer Aeschna (Aeshnidae) Coryphaeschna adnexa
j macrostigma Rambur Libellula (Libellulidae) Orthemis ferruginea
macrotona Williamson Somatochlora (Corduliidae) Somatochlora
jamascarensis Hagen Epitheca (Corduliidae) Neurocordulia franklini
yamaskanensis maculata Beauvois Agrion (Calopterygidae) Calopteryx
janata Say Aeschna (Aeshnidae) Basiaeschna maculata Rambur Libellula (Libellulidae) Libellula semifasciata
janeae Carle Sympetrum (Libellulidae) Sympetrum maculatus Rambur Anax (Aeshnidae) Anax amazili
jesseana Williamson Libellula (Libellulidae) Libellula maculatus Selys Cordulegaster (Cordulegastridae) Cordulegaster
johannus Needham Ophiogomphus (Gomphidae) Ophiogomphus maculiventris Rambur Libellula (Libellulidae) Erythemis
mainensis simplicicollis
jucundus Needham Gomphus (Gomphidae) Stylurus notatus madida Hagen Diplax (Libellulidae) Sympetrum
julia Uhler Libellula (Libellulidae) Libellula magnifica McLachlan Macromia (Corduliidae) Macromia
juncea Linnaeus Libellula (Aeshnidae) Aeshna mainensis Packard Ophiogomphus (Gomphidae) Ophiogomphus
junia Drury Libellula (Aeshnidae) Anax manitobanus Walker Gomphus (Gomphidae) Gomphus fraterna
justiniana Selys Libellula (Libellulidae) Erythrodiplax marcella Selys Libellula (Libellulidae) Miathyria
margarita Donnelly Somatochlora (Corduliidae) Somatochlora
margarita Westfall Macromia (Corduliidae) Macromia
k maritimum Provancher Anax (Aeshnidae) Aeshna canadensis
martha Williamson Celithemis (Libellulidae) Celithemis
kellicotti Williamson Ischnura (Coenagrionidae) Ischnura massettensis Whitehouse Somatochlora (Corduliidae) Somatochlora
kennedyi Walker Somatochlora (Corduliidae) Somatochlora albicincta
krugii Kolbe Enallagma (Coenagrionidae) Enallagma coecum materna Say Calopteryx (Calopterygidae) Calopteryx maculata
kurilis Hagen Argia (Coenagrionidae) Argia vivida maxwelli Ferguson Gomphus (Gomphidae) Arigomphus
kurilis Hagen Gomphus (Gomphidae) Gomphus maya Calvert Dythemis (Libellulidae) Dythemis
mendax Hagen Dythemis (Libellulidae) Brechmorhoga
l meridionalis Hagen Progomphus (Gomphidae) Progomphus
borealis
metella Selys Libellula (Libellulidae) Perithemis domitia
lacerata Hagen Tramea (Libellulidae) Tramea
militaris Hagen Gomphus (Gomphidae) Gomphus
lacrimans Hagen Agrion (Coenagrionidae) Argia
minor Calvert Somatochlora (Corduliidae) Somatochlora
lais Brauer Nehalennia (Coenagrionidae) Apanisagrion
minor Rambur Aeschna (Aeshnidae) Basiaeschna janata
lampropeltis Kennedy Erpetogomphus (Gomphidae) Erpetogomphus
minuscula Rambur Libellula (Libellulidae) Erythrodiplax
laterale Morse Enallagma (Coenagrionidae) Enallagma
minusculum Morse Enallagma (Coenagrionidae) Enallagma
lateralis Burmeister Epophthalmia (Corduliidae) Epitheca cynosura
minutus Rambur Gomphus (Gomphidae) Gomphus
lateralis Needham Somatochlora (Corduliidae) Somatochlora
mista Navas Argia (Coenagrionidae) Argia extraneum
linearis
mithra Selys Libellula (Libellulidae) Erythemis attala
lateralis Scudder Cordulegaster (Cordulegastridae) Cordulegaster
modestus Needham Gomphus (Gomphidae) Gomphus
diastatops
moestum Hagen Agrion (Coenagrionidae) Argia
laurae Williamson Stylurus (Gomphidae) Stylurus
molesta Walsh Cordulia (Corduliidae) Neurocordulia
laurenti Calvert Enallagma (Coenagrionidae) Enallagma vesperum
monomelaena Williamson Celithemis (Libellulidae) Celithemis
leda Say Libellula (Libellulidae) Libellula axilena
fasciata
lentulus Needham Gomphus (Gomphidae) Arigomphus
montanus Selys Ophiogomphus (Gomphidae) Ophiogomphus severus
leonorae Garrison Argia (Coenagrionidae) Argia
montezuma Calvert Trithemis (Libellulidae) Erythrodiplax umbrata pentacantha Rambur Aeschna (Aeshnidae) Nasiaeschna
mooma Kirby Perithemis (Libellulidae) Perithemis perparva Selys Ischnura (Coenagrionidae) Ischnura
morrisoni Ris Sympetrum (Libellulidae) Sympetrum pallipes persephone Donnelly Aeshna (Aeshnidae) Aeshna
morrisoni Selys Ophiogomphus (Gomphidae) Ophiogomphus pertinax Hagen Dythemis (Libellulidae) Brechmorhoga
mortimer Needham Gomphus (Gomphidae) Gomphus descriptus petechialis Muttkowski Epitheca (Corduliidae) Epitheca
multicincta Say Aeschna (Aeshnidae) Epiaeschna heros phaleratus Needham Ophiogomphus (Gomphidae) Ophiogomphus
multicolor Hagen Aeshna (Aeshnidae) Aeshna occidentis
munda Calvert Argia (Coenagrionidae) Argia pictum Morse Enallagma (Coenagrionidae) Enallagma
mutata Hagen Aeshna (Aeshnidae) Aeshna pictus Needham Herpetogomphus (Gomphidae) Ophiogomphus
rupinsulensis
pilipes Hagen Gomphus (Gomphidae) Arigomphus pallidus
n pima Garrison Argia (Coenagrionidae) Argia
piscinarium Williamson Enallagma (Coenagrionidae) Enallagma
naeva Hagen Dythemis (Libellulidae) Erythrodiplax berenice geminatum
naevius Hagen Gomphus (Gomphidae) Stylogomphus albistylus plagiatus Selys Gomphus (Gomphidae) Stylurus
nahuana Calvert Argia (Coenagrionidae) Argia plana Calvert Argia (Coenagrionidae) Argia
nasalis Selys Epitheca (Corduliidae) Somatochlora semicircularis plebeja Burmeister Libellula (Libellulidae) Erythemis
natrix Williamson and Williamson Erpetogomphus (Gomphidae) plebium Selys Enallagma (Coenagrionidae) Enallagma civile
Erpetogomphus lampropeltis plumbea Uhler Libellula (Libellulidae) Libellula flavida
needhami Westfall Libellula (Libellulidae) Libellula pocahontas Kirby Perithemis (Libellulidae) Perithemis domitia
nervosa Rambur Gynacantha (Aeshnidae) Gynacantha poeyi Scudder Mesothemis (Libellulidae) Micrathyria didyma
polernista Navas Oplonaeschna (Aeshnidae) Oplonaeschna armata
nevadensis Kennedy Gomphus (Gomphidae) Stylurus olivaceus pollutum Hagen Agrion (Coenagrionidae) Enallagma
nevadensis Kennedy Ophiogomphus (Gomphidae) Ophiogomphus polysticta Burmeister Libellula (Corduliidae) Neurocordulia
morrisoni obsoleta
nevadensis Walker Aeshna (Aeshnidae) Aeshna interrupta ponderosa Karsch Erythrodiplax (Libellulidae) Erythrodiplax
nigrescens Calvert Dythemis (Libellulidae) Dythemis connata
nodisticta Hagen Libellula (Libellulidae) Libellula portoricana Kolbe Diplax (Libellulidae) Erythrodiplax justiniana
notatus Rambur Gomphus (Gomphidae) Stylurus positum Hagen Agrion (Coenagrionidae) Ischnura
novaehispaniae Calvert Enallagma (Coenagrionidae) Enallagma potulentus Needham Gomphus (Gomphidae) Stylurus
praevarum Hagen Agrion (Coenagrionidae) Enallagma
o princeps Hagen Epitheca (Corduliidae) Epitheca
procera Selys Epitheca (Corduliidae) Somatochlora linearis
prognatum Hagen Agrion (Coenagrionidae) Ischnura
obliqua Say Aeschna (Cordulegastridae) Cordulegaster
obscura Rambur Argia (Coenagrionidae) Argia fumipennis
propinqua Scudder Aeshna (Aeshnidae) Aeshna verticalis
obscurus Rambur Diastatomma (Gomphidae) Progomphus
protracta Hagen Cyclophylla (Gomphidae) Aphylla
obsoleta Say Libellula (Corduliidae) Neurocordulia
provocans Calvert Somatochlora (Corduliidae) Somatochlora
obtrusa Hagen Diplax (Libellulidae) Sympetrum
proxima Calvert Leucorrhinia (Libellulidae) Leucorrhinia
occidentale Bartenef Sympetrum (Libellulidae) Sympetrum
pruinosa Kirby Micrathyria (Libellulidae) Micrathyria didyma
occidentalis Walker Aeshna (Aeshnidae) Aeshna umbrosa
pseudamericana Walsh Hetaerina (Calopterygidae) Hetaerina
occidentis Hagen Ophiogomphus (Gomphidae) Ophiogomphus
americana
octoxantha Ris Perithemis (Libellulidae) Perithemis mooma
psilus Calvert Aeshna (Aeshnidae) Aeshna
odiosa Hagen Libellula (Libellulidae) Libellula luctuosa
puella Kirby Aino (Libellulidae) Nannothemis bella
oenea Hagen Argia (Coenagrionidae) Argia
pulchella Burmeister Libellula (Libellulidae) Celithemis amanda
oklahomensis Pritchard Gomphus (Gomphidae) Gomphus
pulchella Drury Libellula (Libellulidae) Libellula
olivaceus Selys Gomphus (Gomphidae) Stylurus
putridum Hagen Agrion (Coenagrionidae) Argia moestum
onusta Hagen Tramea (Libellulidae) Tramea
pygmaea Kirby Nannophya (Libellulidae) Nannothemis bella
opaca Say Calopteryx (Calopterygidae) Calopteryx maculata
ornata Rambur Libellula (Libellulidae) Celithemis
ozarkensis Bird Somatochlora (Corduliidae) Somatochlora q
ozarkensis Westfall Gomphus (Gomphidae) Gomphus
quadratum Selys Acanthagrion (Coenagrionidae) Acanthagrion
p quadricolor Walsh Gomphus (Gomphidae) Gomphus
quadrifida Rambur Gynacantha (Aeshnidae) Gomphaeschna
furcillata
pacifica Hagen Macromia (Corduliidae) Macromia
quadriguttata Burmeister Aeschna (Aeshnidae) Boyeria vinosa
pallens Calvert Argia (Coenagrionidae) Argia
quadrimaculata Linnaeus Libellula (Libellulidae) Libellula
pallidula Calvert Nehalennia (Coenagrionidae) Nehalennia
quadripunctata Fabricius Libellula (Libellulidae) Libellula
pallidum Root Enallagma (Coenagrionidae) Enallagma
quadrimaculata
pallidus Rambur Gomphus (Gomphidae) Arigomphus
quadrupla Say Libellula (Libellulidae) Libellula cyanea
pallipes Hagen Diplax (Libellulidae) Sympetrum
palmata Hagen Aeshna (Aeshnidae) Aeshna
papilionaria Rambur Calopteryx (Calopterygidae) Calopteryx r
maculata
parvidens Currie Gomphus (Gomphidae) Gomphus ramburii Selys Agrion (Coenagrionidae) Ischnura
parvulus Selys Gomphus (Gomphidae) Lanthus rectangularis Say Lestes (Lestidae) Lestes
patricia Walker Leucorrhinia (Libellulidae) Leucorrhinia recurvatum Davis Enallagma (Coenagrionidae) Enallagma
paulina Frster Tramea (Libellulidae) Tramea binotata
regina Hagen Cordulia (Corduliidae) Epitheca princeps socia Rambur Libellula (Libellulidae) Pachydiplax longipennis
resolutum Selys Agrion (Coenagrionidae) Coenagrion solita Kennedy Argia (Coenagrionidae) Argia pallens
reticulata Kirby Crocothemis (Libellulidae) Crocothemis servilia sordidus Hagen Gomphus (Gomphidae) Gomphus lividus
rhoadsi Calvert Argia (Coenagrionidae) Argia soror Rambur Libellula (Libellulidae) Crocothemis servilia
richardsoni Hagen Cordulia (Corduliidae) Somatochlora specularis Hagen Erythemis (Libellulidae) Idiataphe cubensis
septentrionalis specularis Hagen Neogomphus (Gomphidae) Octogomphus
rickeri Walker Macromia (Corduliidae) Macromia magnifica spegazzinii Navas Argia (Coenagrionidae) Argia plana
rita Kennedy Argia (Coenagrionidae) Argia munda spicatus Hagen Gomphus (Gomphidae) Gomphus
robustum Selys Enallagma (Coenagrionidae) Enallagma spiniceps Walsh Macrogomphus (Gomphidae) Stylurus
cyathigerum spiniferus Rambur Anax (Aeshnidae) Anax junia
rochai Navas Macrothemis (Libellulidae) Macrothemis inacuta spinigera Selys Epitheca (Corduliidae) Epitheca
rogersi Gloyd Gomphus (Gomphidae) Gomphus spinosa Hagen Cordulia (Corduliidae) Epitheca
rubicundula Say Libellula (Libellulidae) Sympetrum spinosus Selys Dromogomphus (Gomphidae) Dromogomphus
rupamnensis Walsh Hetaerina (Calopterygidae) Hetaerina titia spoliatus Hagen Gomphus (Gomphidae) Dromogomphus
rupinsulensis Walsh Herpetogomphus (Gomphidae) Ophiogomphus spumaria Hagen Lestes (Lestidae) Lestes
rupinsulensis Walsh Hetaerina (Calopterygidae) Hetaerina titia stella Williamson Epitheca (Corduliidae) Epitheca
ruralis Burmeister Libellula (Libellulidae) Erythrodiplax umbrata stigmata Say Aeschna (Gomphidae) Phyllogomphoides
russata Calvert Dythemis (Libellulidae) Paltothemis lineatipes striata Selys Lestes (Lestidae) Lestes forficula
stulta Hagen Lestes (Lestidae) Lestes
subapicalis Williamson Gomphus (Gomphidae) Arigomphus lentulus
s subarctica Walker Aeshna (Aeshnidae) Aeshna
subbinotata Brauer Tramea (Libellulidae) Tramea binotata
saalasi Valle Argia (Coenagrionidae) Argia nahuana subfasciata Burmeister Libellula (Libellulidae) Erythrodiplax
sabino Garrison Argia (Coenagrionidae) Argia umbrata
sahlbergi Trybom Somatochlora (Corduliidae) Somatochlora sublata Hagen Lestes (Lestidae) Lestes alacer
sallaei Selys Libellula (Libellulidae) Brechmorhoga pertinax submedianus Williamson Gomphus (Gomphidae) Arigomphus
salvum Hagen Agrion (Coenagrionidae) Telebasis subornata Hagen Plathemis (Libellulidae) Libellula
sandrius Tennessen Gomphus (Gomphidae) Gomphus sulcatum Williamson Enallagma (Coenagrionidae) Enallagma
saturata Hagen Cordulia (Corduliidae) Somatochlora elongata superba Hagen Celithemis (Libellulidae) Pseudoleon
saturata Uhler Libellula (Libellulidae) Libellula susbehcha Vogt and Smith Ophiogomphus (Gomphidae)
saucium Burmeister Agrion (Coenagrionidae) Amphiagrion Ophiogomphus
sayi Selys Cordulegaster (Cordulegastridae) Cordulegaster
sclerata Walsh Hetaerina (Calopterygidae) Hetaerina americana
scudderi Selys Gomphus (Gomphidae) Stylurus t
sedulum Hagen Agrion (Coenagrionidae) Argia
segregans Needham Stylurus (Gomphidae) Stylurus spiniceps taeniolata Rambur Macromia (Corduliidae) Macromia
selysii Hagen Cordulia (Corduliidae) Helocordulia tarascana Calvert Argia (Coenagrionidae) Argia
semiaquea Burmeister Libellula (Corduliidae) Epitheca tenebrica Hagen Cordulia (Corduliidae) Somatochlora tenebrosa
semicincta Say Libellula (Libellulidae) Sympetrum tenebrosa Say Libellula (Corduliidae) Somatochlora
semicirculare Selys Enallagma (Coenagrionidae) Enallagma tenera Say Libellula (Libellulidae) Perithemis
semicircularis Selys Epitheca (Corduliidae) Somatochlora tenuata Rambur Lestes (Lestidae) Lestes
semifasciata Burmeister Libellula (Libellulidae) Libellula tenuicincta Say Libellula (Libellulidae) Perithemis tenera
seminole Calvert Perithemis (Libellulidae) Perithemis tenera terminalis Burmeister Libellula (Libellulidae) Pantala flavescens
sepia Gloyd Epitheca (Corduliidae) Epitheca ternaria Say Libellula (Libellulidae) Libellula quadrimaculata
septentrionalis Burmeister Aeshna (Aeshnidae) Aeshna texana Walsh Hetaerina (Calopterygidae) Hetaerina americana
septentrionalis Hagen Cordulia (Corduliidae) Somatochlora tezpi Calvert Argia (Coenagrionidae) Argia
septentrionalis Selys Hetaerina (Calopterygidae) Hetaerina titia thoreyi Hagen Uropetala (Petaluridae) Tachopteryx
septima Selys Gynacantha (Aeshnidae) Triacanthagyna tibialis Rambur Platycnemis (Coenagrionidae) Argia
septima Selys Micrathyria (Libellulidae) Micrathyria aequalis titia Drury Libellula (Calopterygidae) Hetaerina
septima Westfall Gomphus (Gomphidae) Gomphus tomentosa Fabricius Libellula (Corduliidae) Epitheca cynosura
tonto Calvert Argia (Coenagrionidae) Argia
sequoiarum Butler Ophiogomphus (Gomphidae) Ophiogomphus townesi Gloyd Stylurus (Gomphidae) Stylurus
bison translata Hagen Argia (Coenagrionidae) Argia
serva Fabricius Libellula (Libellulidae) Libellula lydia transversa Say Libellula (Corduliidae) Didymops
servilia Drury Libellula (Libellulidae) Crocothemis traviatum Selys Enallagma (Coenagrionidae) Enallagma
servillei Rambur Didymops (Corduliidae) Didymops transversa tricolor Burmeister Calopteryx (Calopterygidae) Hetaerina titia
servum Selys Agrion (Coenagrionidae) Coenagrion resolutum trimaculata De Geer Libellula (Libellulidae) Libellula lydia
severus Hagen Ophiogomphus (Gomphidae) Ophiogomphus tripartita Burmeister Libellula (Libellulidae) Erythrodiplax umbrata
shurtleffi Scudder Cordulia (Corduliidae) Cordulia tuberculata Selys Ischnura (Coenagrionidae) Ischnura ramburii
signatum Hagen Agrion (Coenagrionidae) Enallagma tuberculifera Walker Aeshna (Aeshnidae) Aeshna
signiferum Cannings and Garrison Sympetrum (Libellulidae) tyleri Kirby Trithemis (Libellulidae) Erythrodiplax funerea
Sympetrum
simile Selys Enallagma (Coenagrionidae) Enallagma civile
simplicicollis Say Libellula (Libellulidae) Erythemis u
simulans Muttkowski Epitheca (Corduliidae) Epitheca cynosura
sitchensis Hagen Aeshna (Aeshnidae) Aeshna uhleri Selys Cordulia (Corduliidae) Helocordulia
smithii Kirby Cannacria (Libellulidae) Brachymesia furcata umbrata Linnaeus Libellula (Libellulidae) Erythrodiplax
sobrinus Selys Gomphus (Gomphidae) Gomphus kurilis umbratus Needham Gomphus (Gomphidae) Gomphus lividus
umbrosa Walker Aeshna (Aeshnidae) Aeshna williamsoni Walker Somatochlora (Corduliidae) Somatochlora
uncatus Kirby Lestes (Lestidae) Lestes dryas
unguiculata Hagen Lestes (Lestidae) Lestes
unifasciata De Geer Libellula (Libellulidae) Erythrodiplax umbrata x
uniformis Kirby Belonia (Libellulidae) Libellula croceipennis
utahensis Muttkowski Ischnura (Coenagrionidae) Ischnura barberi xanthosoma Williamson Platycordulia (Corduliidae) Neurocordulia
v y
vacua Hagen Diplax (Corduliidae) Williamsonia fletcheri yakima Hagen Calopteryx (Calopterygidae) Calopteryx aequabilis
validus Hagen Anax (Aeshnidae) Anax walsinghami yamaskanensis Provancher Aeshna (Corduliidae) Neurocordulia
vastus Walsh Gomphus (Gomphidae) Gomphus ypsilon Needham Enallagma (Coenagrionidae) Enallagma
velox Hagen Dythemis (Libellulidae) Dythemis novaehispaniae
venerinotata Haldeman Agrion (Coenagrionidae) Ischnura hastata
ventricosus Walsh Gomphus (Gomphidae) Gomphus
verbenata Hagen Lepthemis (Libellulidae) Erythemis plebeja
verna Pritchard Celithemis (Libellulidae) Celithemis
vernale Gloyd Enallagma (Coenagrionidae) Enallagma cyathigerum
vernalis Carle Lanthus (Gomphidae) Lanthus
versicolor Fabricius Libellula (Libellulidae) Libellula pulchella
verticalis Hagen Aeshna (Aeshnidae) Aeshna
verticalis Say Agrion (Coenagrionidae) Ischnura
verum Bartenef Sympetrum (Libellulidae) Sympetrum danae
vesiculosa Fabricius Libellula (Libellulidae) Erythemis
vesperum Calvert Enallagma (Coenagrionidae) Enallagma
vibrans Fabricius Libellula (Libellulidae) Libellula
vicina Hagen Diplax (Libellulidae) Sympetrum
vidua Hagen Lestes (Lestidae) Lestes
vigilax Hagen Lestes (Lestidae) Lestes
villosipes Selys Gomphus (Gomphidae) Arigomphus
vinosa Say Aeschna (Aeshnidae) Boyeria
violaceum Hagen Agrion (Coenagrionidae) Argia fumipennis
virens Rambur Aeschna (Aeshnidae) Coryphaeschna viriditas
virginica Selys Calopteryx (Calopterygidae) Calopteryx aequabilis
virginiensis Davis Neurocordulia (Corduliidae) Neurocordulia
virgo Calvert Lestes (Lestidae) Lestes inaequalis
viridifrons Hine Gomphus (Gomphidae) Gomphus
viriditas Calvert Coryphaeschna (Aeshnidae) Coryphaeschna
viridula Beauvois Libellula (Libellulidae) Pantala flavescens
vivax Calvert Brechmorhoga (Libellulidae) Brechmorhoga
vivida Hagen Argia (Coenagrionidae) Argia
vulgipes Calvert Macrothemis (Libellulidae) Macrothemis
inequiunguis
vulnerata Hagen Hetaerina (Calopterygidae) Hetaerina
w
walkeri Kennedy Aeshna (Aeshnidae) Aeshna
walkeri Kennedy Somatochlora (Corduliidae) Somatochlora
sahlbergi
walkeri Muttkowski Enallagma (Coenagrionidae) Coenagrion
resolutum
walkeri Whitehouse Tramea (Libellulidae) Tramea binotata
walshii Kellicott Gomphus (Gomphidae) Gomphus crassus
walshii Scudder Cordulia (Corduliidae) Somatochlora
walsinghami McLachlan Anax (Aeshnidae) Anax
weewa Byers Enallagma (Coenagrionidae) Enallagma
westfalli Carle and May Gomphus (Gomphidae) Gomphus
westfalli Cook and Daigle Ophiogomphus (Gomphidae)
Ophiogomphus
westfalli Donnelly Enallagma (Coenagrionidae) Enallagma
traviatum
whedoni Muttkowski Gomphus (Gomphidae) Arigomphus cornutus
whitehousei Walker Somatochlora (Corduliidae) Somatochlora
williamsoni Gloyd Gomphoides (Gomphidae) Aphylla
williamsoni Muttkowski Epitheca (Corduliidae) Epitheca costalis