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An ethnobotanical study of the plant


biodiversity of Mt. Mayon, Bicol Peninsula,
Albay, Philippines

Article · January 2009

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Journal of Nature Studies 8(1): 1-10
ISSN: 1655-3179

A ETHOBOTAICAL STUDY OF THE PLAT BIODIVERSITY OF


MT. MAYO, BICOL PEISULA, ALBAY, PHILIPPIES

Inocencio E. Buot Jr.


Institute of Biological Sciences, College of Arts and Sciences
and School of Environmental Science and Management,
University of the Philippines Los Baños
College, Laguna, Philippines
email: ibuotjr@yahoo.com

ABSTRACT - A total of 71 woody species belonging to 49 genera and 33 families are recorded along the
altitudinal range of 500 -2200 meters above sea level on the northwestern side of Mt. Mayon, Bicol Peninsula,
Albay, Philippines. Ethnobotanical uses are diverse ranging from being medicinal, food, timber, firewood,
ornamental, sources of fibers, industrial compounds to being poisonous. Thirty-two percent of the species are
endemics while the rest are indigenous showing greatest development in the Malesian phytogeographical region.
Endemism is very high despite volcanic activity and extensive anthropogenic disturbances such as slash and
burn farming and overharvesting of plant biota for sale to domestic tourists. There is an urgent need to
determine the biochemical basis for the identified uses of Mt. Mayon plants as this may pave the way to drug
discovery and hence, economic development of the locality. An effective domestication program of Mt. Mayon
plant biodiversity involving the surrounding local communities could be practical and convenient for everyone
and favors protection of the entire Mt Mayon ecosystem and any other forests for that matter.

Key words: ethnoecology, man-environment interaction, Mayon Volcano, plant diversity, economic botany.

ITRODUCTIO
Ethnoecological studies deal with the interaction world for its beautiful cone. The vegetation zones
between people and the nature (Cunningham 1996, and the dominants had been reported by Buot (2005,
Martin 1995). It becomes more ethnobotanical if it is 2008). He identified three altitudinal vegetation
the interaction between people and plants. In the zones of woody species on the northwestern side of
Philippines, ethnobotanical works on the biodiversity of the volcanic mountain ecosystem. Each zone was
mountains are very few despite the rich ethnodiversity named after the most dominant species determined
and the impressive megadiversity of flora and fauna following the method of Ohsawa (1984). Zone I was
(Mittermeier et al., 1997) in the country. There is a Erythrina-Ficus-Glochidion zone (500-800 m asl),
need to document the folk uses of biodiversity which Zone II was Astronia-Cyathea-Weinmannia zone
could range from being a food, medicinal, ornamental (900-1500 m asl) while Zone III was Eurya-Clethra-
or a poison plant, to that of an important constituent in *eonauclea-Fagraea-Vaccinium zone (1600-2200 m
traditional cultural rituals or maybe charms or amulets asl). Detailed description and analyses of these
of rural communities. Such studies are very important altitudinal vegetation zones can be found in Buot
to determine the real economic value or essence of our (2008). Mt. Mayon is also the home of wild pigs,
megadiversity. The megadiversity is nothing unless we wild cats and lizards, too.
know its purpose and its worth and practical relevance Settlers are found along the immediate boundary
to our survival. of Mt. Mayon. Illegal collection of Mt. Mayon
Assessment of biodiversity and ethnobotany along resources has been reported. Habitat destruction and
altitudinal gradient is critical in determining the wealth species extinction are unavoidable on Mt. Mayon.
of resources people can benefit from and that people Hence, regular biodiversity assessment and
need to protect. Results would be very relevant in land ethnobotanical studies along altitudinal zones of the
use planning and zonation of mountain landscapes volcano are imperative just like the works of Maundu
which are vital to species and habitat conservation and et al 2001, Cunningham et al., 2002, Martin et al.,
satisfaction of people residing in the vicinities. 2002, Koizumi and Momose 2007).
Many mountains in Bicol region are left unattended The present study aimed to 1) study the
with respect to biodiversity and ethnobotanical studies ethnobotany of the biodiversity along the altitudinal
along altitudinal gradients despite the threatening gradients reported by Buot (2008) on Mt. Mayon
destruction and extinction of biota either through (e.g. wild food plant, medicinal plant, poison plant or
natural or anthropogenic causes. plant used in traditional cultural rituals), and 2)
Mt. Mayon, the world’s most perfect cone located classify plant biota used by local folks into
in Albay, is one of these. Mt. Mayon (2462 m above indigenous, endemic or exotic species.
sea level) is an active volcano renowned throughout the

Journal of
ature Studies 8(1) January – April 2009
Buot Jr. 2

METHODOLOGY
Describing the Study Area. The study area was Mt. is very pronounced during the month of December to
Mayon National Park (13016’ N latitude and 1230 42’ E January and a dry summer season from March to
longitude) located in Albay, Bicol Peninsula, southern May (PAG-ASA, Albay 2002). Average yearly
Luzon, Philippines (Fig. 1). Mt. Mayon (Fig. 2) is a rainfall is approximately 4330 millimeters. The
national protected area under the National Integrated average relative humidity is about 83%.
Protected Area System (NIPAS). It is a favorite tourist The National Statistical Coordination Board
destination (Buot and Tapay 2003, Tapay and Buot (2002) indicated the land use on Mt. Mayon. Evident
2003) owing to its near perfect cone (Fig. 2) and its are patches of slash and burn clearing (kaingin),
awesome and breathtaking scenery during eruption (Fig cultivated and planted to coconuts, fruit trees and
3). It covers an area of 250 sq. km or roughly 10% of abacas. Some of these kaingin areas are still being
Albay province. Its base circumference is 63 sq. km planted to vegetables, camote, and other seasonal
and its slope angle steepens gradually until it reaches root crops. These cultivated areas were made by
the maximum of 35 degrees (Mines and Geosciences transient migrants.
Bureau – PHIVOLCS and Earth Remote Sensing Documenting the Economic uses of Mt. Mayon
Analysis Center 1998). The topography is characterized Plant Biodiversity. Uses of plants by local people
by scattered fertile plains and valleys, consisting of were documented by non-structured interviews with
rough ridges and deep ravines and seven major river plant collectors and other local residents. (Sample
systems. Mt. Mayon is considered to be the one of the guide questions:…What is the local name of the
most active volcanoes in the Philippines (Paterno plant?..., What is the local use?..., Is it always
1992). It periodically erupts every eight (8) or ten (10) collected by the local residents?..., Why?..., Do you
years for the last forty (40) years, having had at least have other sources of this plant aside from Mt.
forty-seven (47) eruptions since 1616 (Ramos-Villarta Mayon?..., Where?). Plant collectors and local
et al 1985). residents interviewed were selected based on the
recommendation by the local DENR office.
Two plant collectors (Jose Bon Jr. of Mt.
Mayon, Tabaco City and Alfredo Orendain of
Lidong, Sto. Domingo, Albay) and four local
residents (Alfredo Brutas of Canaway, Malilipot,
Albay, Juan Berces of Calbayog, Malilipot, Albay,
Teresa Buban of Buang, Tabaco, Albay, and
Bonifacio Odiver of Matnog, Daraga, Albay were
interviewed.
Most of the information on uses, however, were
taken from books (Rojo 1999, Merrill 1923-1926)
since local residents know more of the firewood,
timber, and ornamental plants only. Taxon
information whether it is endemic, indigenous or
exotic and region of greatest distribution and
development were noted from Philippine taxonomic
literature.

omenclature and biometrics. All woody
species having at least 0.5 m in height measured by
Buot (2008) in his vegetation zonation study using a
combined quadrat and point centered quarter
methods (Mueller-Dombois and Ellenberg 1974)
from 500-2200 meters above sea level (m asl) were
considered samples representing the Mt. Mayon plant
biodiversity. Nomenclature followed that of Merrill
(1923-1926) but were updated by Dr. J.F. Veldkamp
Figure 1. The Philippine map indicating the study area of the National Herbarium of Netherlands.
(bold arrow).
Mt. Mayon is classified as strata volcano or RESULTS AD DISCUSSIO
composite cone. It consists of deposit materials, lava Plant Biodiversity along Altitudinal Gradients.
flows, air fall deposition, pyroclastic flows and lahar A total of 71 woody species belonging to 49 genera
flows triggered by rainfall (Mines and Geosciences and 33 families were recorded by Buot (2008) along
Bureau – PHIVOLCS and Earth Remote Sensing the altitudinal range of 500 -2200 m asl on the
Analysis Center, 1998). This condition leads to the northwestern side of Mt. Mayon (Table 1). The
consequent formation of gravelly sandy loam soil with physical and structural characteristics of the various
extreme acidity and very low natural fertility. sampling sites are presented in Table 2.
The watershed falls in Type II climate where rainfall The flora is dominated expectedly by Malesian

Journal of
ature Studies 8(1) January – April 2009
An Ethnobotanical Study Of The Plant Biodiversity Of Mt. Mayon 3

Figure 2. Mt. Mayon showing its perfect cone and a cloud forming a belt at the montane zone (photo by IE
Buot, Jr.).

Figure 3. A breathtaking view of Mt. Mayon during the eruption in 2000 (photo bought from vendors).

Journal of
ature Studies 8(1) January – April 2009
Buot Jr. 4

elements like families Euphorbiaceae, Moraceae, indigenous and endemic species on Mt. Mayon (Table
Urticaceae and Verbenaceae. This is true to many 1). We have to understand that the Mt. Mayon biota
other mountains here just like Mt. Pangasugan in are already strained due to frequent eruption of the
Leyte (Belonias and Aguilar 2004) and Mt Makiling volcano. When anthropogenic stressors will intensify
in Los Banos, Laguna (Aragones 1991, Payawal and and aggravate the present predicament of the
Markgraf 1981, Luna et al., 1999, Brown 1919, ecosystem, we do not even know yet how long Mt.
Trelease and McLean 1919) or generally in any other Mayon palnt biota could be resilient enough to cope
tropical mountain (Richards 1996). In contrast, only with the extreme disturbance.
Lauraceae which is well developed in the northern Conservation Strategies. Surprisingly, 23 or so out
floristic regions (Buot and Okitsu 1997a, 1997b, 1998, of 71 woody species on Mt. Mayon are endemic to the
1999, Gonzalez-Salcedo 2001, Buot 2007) is Philippines (Table 1) following Merrill (1923-1926)
represented with a number of component species and Rojo (1999). This is a very high figure already in
comparable to the aforementioned Malesian families. an area with active volcanic activity and frequent
Ethnobotany of the Biodiversity on Mt. Mayon. eruption. The rest are either indigenous or yet
Uses of the plant biota along the altitudinal slopes of uncertain as to the distribution of the species/taxon.
Mt. Mayon (Table 1) are diverse ranging from being Table 1 further shows that the region of greatest
medicinal, food, timber, firewood, ornamental, development of Mt. Mayon taxa is the Malesian
sources of fibers, industrial compounds (such as phytogeographical region (Merril 1923-1926, Rojo
tannin, etc.) to being poisonous (Rojo 1999, Merrill 1999). Two each were determined as Pantropic
1923-1926). Many people had not known the uses of (Calophyllum inophyllum and Symplocos
these woody plants yet, other than plants for firewood cochichinensis) and Australasian (Litsea perrottetii
and timber purposes. This implies that much work and Premna obtusifolia) while one had its greatest
have to be done to examine the biochemical basis on development in India (Erythrina variegata).
the potential uses of these plants in industries and We have to understand that the endemics (32%) on
pharmacy. Once realized, this could pave the way to Mt. Mayon are part of the global heritage being found
economic development in the locality benefiting local only in the Philippines. Hence, it is indeed miserable
residents. to lose anyone of them under whatever circumstance.
At present, most plant collectors (Fig. 4) and local However people need to survive as well. Therefore
people know only the uses of the herbaceous plants there should be a just and fair solution to this issue
like ferns and their allies, grasses like Miscanthus (Fig having in mind not only the ecosystem but the people
5A), spice plants like Begonia aequata (Fig. 5B), and as well (Kaplan et al., 1998).
others. They are regularly interacting with these plant Declaring an area as protected had been thought of
groups most of the times. In fact, many residents were as a very good strategy in conserving the plant
already observed utilizing these resources extensively. biodiversity. Indeed, it is enough if there were no
Abuse in the utilization was evident in places near Mt. intervening issues/problems as social and economic
Mayon probably because of the pressure of increasing complexities. But there are! Hence the need for an
population. additional strategy.
Driven by the need to survive, local residents A participatory approach in the conservation of Mt.
around the vicinities, engaged in whatever form of job Mayon resources may be the most appropriate.
to sustain life. Observing the promising resources on Consultation with local residents and collectors can
Mt. Mayon, the people tried to explore possibilities. help design a conservation scheme that will be both
The most rampant around and in Mt. Mayon forests, ecologically sustainable and socially acceptable. One
were agriculture, tourism and collecting of biota, possible strategy would be scheduled and managed
either for family and/or commercial use (Fig. 4-5). harvesting of resources from the various slopes by
Trees from the various slopes of Mt. Mayon were different groups of collectors. However, though this
cut for timber, firewood and for other purposes such appears quite promising, yet the mere fact that Mt.
as expansion of agricultural farms. The influx of Mayon is a national protected area, this may not be the
tourists on Mt. Mayon who were appreciative of the best alternative. It depends on the DENR.
local plant biodiversity gave local residents the hint to Another possibility would be gardening of Mt.
start collecting plants for sale to daily visitors. Most of Mayon plants on a larger scale. People will then have
these plants are ferns and their allies, other herbs and to domesticate and farm Mt. Mayon plants and once
dwarf plants collected from Zone III of Buot (2008). ready and marketable, sell them to interested tourists
Most plants from Zone III are very short (Table 2) visiting the area. This could be a good alternative to
because of stressful environmental conditions in the solve the issue on biodiversity conservation on Mt.
higher altitudes (2100-2200 m asl) (Buot 2001, Buot Mayon.
and Okitsu 1998, 1999, Ohsawa 1984). Dwarf plants Plant domestication as a way of conserving
are easily sold as bonsai. These activities indeed indigenous fruit species is widely practiced in Africa
started to reshape the forest landscape of Mt. Mayon. (Ejiofor et al., 1988, Okafor and Lamb 1994, Okafor
This is alarming if continued over a prolonged 1978, 1980, 1991). Conservation of economically
period of time. This may result to extinction of many important species through cultivation was also
suggested by Buot (1999, 2004).

Journal of
ature Studies 8(1) January – April 2009
An Ethnobotanical Study Of The Plant Biodiversity Of Mt. Mayon 5

Table 1. Plant biodiversity along the altitudinal gradient on the northwestern slope of Mt. Mayon. The family
names are arranged alphabetically under which the scientific names are also in alphabetical order. The
common name, uses, nature of species and region of greatest development are noted.

REGIO OF
FAMILY AME/ ATURE OF
COMMO AME* USES** GREATEST
SCIETIFIC AME SPECIES***
DEVELOPMET***
ANACARDIACEAE
Semecarpus cuneiformis Blanco Ligas (Tag.) Medicinal & poisonous Endemic -
Semecarpus philippinensis Engl. - Medicinal & poisonous Endemic -
CHRYSOBALANACEAE
Chrysobalanos sp. - - - Malesia
CLETHRACEAE
Firewood species and
Clethra lancefolia Turcz. Malaklak Endemic -
hardwood for construction
CRYPTERONIACEAE
Crypteronia cumingii (Planch.) Endl Tigauon Timber Indigenous Malesia
CUNONIACEAE
Weinmannia hutchinsonii Merr. Hutchinson kalilan Timber Endemic -

Perching materials for


CYATHEACEAE
ornamental epiphytes
Cyathea sp. 1 - - Malesia
Perching materials for
Cyathea sp. 2 - - Malesia
ornamental epiphytes
Cyathea sp. 3 - - Malesia
Perching materials for
ornamental epiphytes
DILLENIACEAE
Dillenia sp. - Medicinal - Malesia
ERICACEAE
Vaccinium myrtoides (Blume) Miq. Alimani (Ilk.) Edible fruits Indigenous Malesia
Vaccinium sp. - Firewood - -
EUPHORBIACEAE
Timber and medicinal
Glochidion breynioides C.B. Rob. Matang-buiu (Bik.) Indigenous Malesia
Firewood
Glochidion rubrum Blume - - -
For cough
Glochidion sp. Matang-hipon (Tag.) Indigenous Malesia
Dye and tannin-producing
Macaranga tanarius (L.) Muell.-Arg. Binunga (Tag.) Indigenous -
plants
FABACEAE
Erythrina variegata L. Mottled leaf Dapdap (Tag.) Medicinal Indigenous Indian
FLACOURTIACEAE
Hydnocarpus alcale C.DC. Dudua (Tag.) Medicinal Endemic -
GESERIACEAEA
Cyrtandra ferruginea Merr. - Timber Endemic -
GUTTIFERAE
Calophyllum inophyllum L. Bitaog (Tag.) Timber - Pantropic
Callophyllum sp. - Timber Indigenous Malesia
LAURACEAE
Cinnamomum mercadoi Vidal Kalingag (Tag.) Medicinal Endemic -
Litsea luzonica (Blume) F.-Vill. Dungoi (Tag.) Firewood Endemic Malesia
Litsea perrottetii (Blume) F.-Vill. Marang (Tag.) Firewood Indigenous Australasia
*eolitsea sp. - Firewood - -
*eonauclea sp. - Firewood - Malesia
LEEACEAE
Leea sp. - Firewood - -
Leea philippinensis Merr. Kaliantan (Tag.) Firewood Endemic -
LOGAIACEAE
Fagraea ceilanica Thunb. Kabal (Tag.) Timber and medicinal Indigenous -
MELASTOMATACEAE
Astronia rolfei Vidal Dungau-pula (Tag.) Firewood Endemic -
Medinilla sp. - Ornamental - Malesia
Melastoma malabathricum L. - Medicinal Indigenous -
MELIACEAE
Toona calantas Merr. & Rolfe Kalantas (Tag.) Timber Indigenous Malesia
Toona ciliata M. Roem. Lanipga (Bik.) Timber Indigenous Malesia
MIMOSACEAE
Pithecellobium clypearia (Jack) Benth. - Timber - Malesia
MORACEAE
Broussonetia sp. - Firewood - Malesia
Ficus benguetensis Merr. Tabul (Tag.) Firewood Endemic -
Ficus gul Laut. Et K.. Schum Butli (Tag.) Firewood Indigenous Malesia
Ficus guyeri Elmer Agupili (Tag.) Firewood Endemic -
Ficus minahassae (Teijsm. & de Vr.) Miq. Hagimit (Tag.) Fiber plants Indigenous Malesia
The soft & fleshy fruits are
Ficus nota (Blanco) Merr. Tibig (Tag.) Endemic -
eaten raw.

Journal of
ature Studies 8(1) January – April 2009
Buot Jr. 6

Table 1 (Continuation). Plant biodiversity along the altitudinal gradient on the northwestern slope of Mt.
Mayon. The family names are arranged alphabetically under which the scientific names are also in alphabetical
order. The common name, uses, nature of species and region of greatest development are noted.

REGIO OF
FAMILY AME/ ATURE OF GREATEST
COMMO AME* USES**
SCIETIFIC AME SPECIES*** DEVELOPMET*
**
MORACEAE
Ficus septica (Burm.f. Medicinal, mainly to cover & cure wounds,
Hawili Indigenous Malesia
boils & sores
Ficus sp. 1 - Firewood - Malesia
Ficus sp. 2 - Firewood - Malesia
Ficus sp. 3 - Firewood - Malesia
Ficus ulmifolia Lam. Is-is Fruits are edible Endemic -
Ficus variegata Blume
Tangisang bayawak Plants producing exudates Indigenous Malesia
MYRISTICACEAE
Knema glomerata (Blanco) Merr. Tambalau Medicinal seed-oils & locally used timbers Endemic -
MYRSIACEAE
Ardisia sp. - Medicinal - Malesia
MYRTACEAE
Syzygium sp. - Timber - Malesia
RUBIACEAE
Coptasapelta olaciformis (Merr.) Elm. Medicinal Endemic Malesia
Mussaenda sp. -
- - Malesia
*eonauclea sp. -
- - Malesia
SAPIDACEAE
Guioa falcata Radlk. - Endemic -
SAURAUIACEAE
Saurauia latibractea Merr.,En. Philip. Kolalabang (Tag.) Timber Endemic -
Saurauia sorsogonensis Merr. - Timber Endemic -
SAXIFRAGACEAE
Itea macrophylla Wall. Kodai (Ig.) Ornamental evergreen trees Indigenous Malesia
STERCULIACEAE
Pterocymbium tinctorium (Blanco.)
Merr. Taluto (Tag.) Timber Indigenous Malesia
SYMPLOCACEAE
Symplocos cochichinensis (Lour.) S.
Moore Agosip (Tag.) For stomachache - Pantropic
THEACEAE
Eurya sp. - Firewood and medicinal - Malesia
ULMACEAE
Trema orientalis (L.) Blume Diuretic, febrifuge and for
Anabiong (Tag.) Indigenous -
stomachache
URTICACEAE
Cypholophos molluccanus (Bl.)
Miq. - Firewood
Indigenous Malesia
Dendrocnide meyeniana (Walp.) Lipang kalabaw Analgesic
Endemic -
Chew (Tag.) Firewood
Endemic -
Laportea luzonensis Radlk. Lipa (Tag.) Used to treat phthisis, cough, headache
Indigenous Malesia
Leucosyke capitellata (Poir.) Alagasi (Tag.) & gastralgia
Indigenous Malesia
Wedd. Alilaua (Neg.) Firewood
Villebrunea trinervis Wedd.
VERBEACEAE
Callicarpa sp. - Medicinal, fish-poison etc. - Malesia
Callicarpa candicans (Burm.f.) For abdominal troubles & amenorrhea.
- - -
Hochr Used as a fish poison also
Clerodendrum brachyanthum Given as a tonic for stomach problems
Schauer Mangha and in large doses is considered Endemic
abortive
Clerodendrum cumingiana
Salumget For stomachache Endemic Malesia
Schauer
Premna obtusifolia R.Br. Diuretic, febrifuge and for
Alagau-dagat (Tag.) Indigenous Australasia
stomachache
Wenlandia sp. - Firewood - Malesia
* Rojo (1999) since after all, Tagalog terms were used as common names by people (e.g. dapdap for Erythrina). Bik = Bicolano.
Ilk. = Ilocano, Neg. = Negros Bisaya, Tag. = Tagalog. Other names have no indication of the dialect used. Still others, have
unknown common names.
** All other uses except for firewood, timber and ornamental, are after Rojo (1999).
*** As indicated by Merrill (1923-1926) & Rojo (1999); - no clear indication of the region of greatest development.

Journal of
ature Studies 8(1) January – April 2009
An Ethnobotanical Study Of The Plant Biodiversity Of Mt. Mayon 7

Table 2. Physical and structural characteristics of the various sampling sites along the altitudinal gradients on
Mt. Mayon.
SAMPLIG SITE
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15
Altitude(m) 500 700 800 900 1000 1100 1200 1300 1400 1500 1600 1700 1900 2100 2200
S 150 S 850 S 100 S 300 S 420 S 30 S 800 S 450 N 450 S 150 S 360 S 180 N 600 N 500 N 500
Exposure E E E E E W E E E W W W E E E
Slope (degree) 60 60 40 18 20 18 30 35 45 45 45 50 45 60 60
Plot size (m2)/Number
100 100 100 sq
of points (point 20 20 20 20 20 20 20 20 20 20 20 20
sq m sq m m
centered quarter)
Number of species 18 19 33 26 22 19 25 18 16 16 11 6 4 4 3
Maximum DBH (cm) 25.46 95.49 79.58 111.4 127.32 111.4 127.32 79.58 65.57 57.3 63.66 57.3 13.69 6.05 3.18
Maximum height (m) 8 10 40 25 18 10 10 8 7 8 8 5 7 3 2

Figure 4. A plant collector climbing on to the upper slopes of Mt. Mayon (photo by IE Buot, Jr.).

Figure 5. A. Miscanthus stems used as wall materials for farm houses and as garden trellis. B. Begonia aequata
(local, maasim) collected from Zone I of Mt. Mayon and used as spice by local residents (photo by
IEBuot, Jr.).

Journal of
ature Studies 8(1) January – April 2009
Buot Jr. 8

SUMMARY AD COCLUSIO


The documented economic uses of the plant Belonias, B. and .O. Aguilar. 2004. Altitudinal
biodiversity on Mt. Mayon are very encouraging. The zonation of dicots on Mt. Pangasugan, Leyte Island,
results should push biologists, pharmacists, chemists Philippines. Journal of Environmental Science and
and biochemists together to determine the biochemical Management 7: 47-66.
basis of the indications. We have to explore our Brown WH. 1919. The Vegetation of Philippine
endemic and indigenous megadiversity for the benefit Mountains. Publication Number 13, Bureau of
of the human society. However, there should be a Science, Manila, 434 p.
regulated exploration especially in the case of Mt. Buot, I.E. Jr. 2008. Vertical Distribution and
Mayon because human activity could worsen the Zonation Pattern of Woody Vegetation on the
stress due to frequent volcanic activity. An effective northwestern slope of Mt. Mayon, Philippines. Asia
domestication program of Mt. Mayon plant Life Sciences 17: 189- 205.
biodiversity involving the surrounding local Buot, I.E. Jr. 2007. Vegetation zonation of Mt. Akiki,
communities could be practical and convenient for Cordillera mountain range, Benguet Province,
everyone and favors protection of the entire Mt Philippines. Asia Life Sciences 16(1): 53-70.
Mayon ecosystem and any other forests for that Buot, I.E. Jr. 2005. Dominant woody species of
matter. Mount Mayon forest landscape, Luzon,Philippines.
Flora Malesiana Bulletin 13: 364..368.
ACKOWLEDGEMETS Buot IE Jr. 2004. Human activity endangers tiger
The author is thankful to the following: Bicol orchid (Grammatophyllum sp., Orchidaceae) in the
Studies Program (BSP) of the University of the satoyama forests of Mt Mayon, Albay, southern
Philippines Los Banos (UPLB) for funding the study; Luzon. USM Research & Development Journal
Forester Rodolfo M. Matusalem [Provincial 12: 49-53.
Environment and Natural Resources Officer, Buot, IE Jr. 2001. Latitudinal comparison of the
Department of Environment and Natural Resources vertical vegetation zonation on Mt. Pulog
(DENR), Legaspi City] for his approval of the (Philippines) with other southeast and east Asian
research study in Mt. Mayon National Park; Forester mountains. Philippine Scientist 38: 9-25.
Pepito Asis, Ms. Alleth Agripa, Mr. Alfredo Orendain Buot, I.E. Jr. 1999. Studies on the forest architecture
and all the staff members of the Protected Area along altitudinal gradients on Mt. Pulag, Luzon Is.,
Office-DENR for their generous assistance in the Philippines. University of Southern Mindanao
field; Dr.J.F. Veldkamp, (Taxonomist and Editor-in- Research and Development Journal 7: 200-212.
Chief of the Flora Malesiana Bulletin, National Buot IE Jr. 1999. Pteridophytes frequently sold at
Herbarium of Netherlands) for his valuable Carbon Market, Cebu: implications to urban
corrections and suggestions on nomenclature; Mr horticulture and nature conservation. The Philippine
Danilo N. Tandang (Philippine National Herbarium, Scientist 36: 148-153.
National Museum, Manila), Mr. Nino Vidal (formerly Buot, I.E. Jr. and .E. Tapay. 2003. Impact of
of the Center for Integrative and Development ecotourism on biodiversity of Mt. Mayon and the
Studies, UP System) and Ms. Ma. Celeste N. local residents in the vicinity, pp. 39-45. In: Buot,
Banaticla (Plant Genetic Resources Center, I.E. Jr. and R.T. Bagarinao (Eds.). Ecotourism and
International Rice Research Institute, Los Biodiversity Conservation: Subjects. Theories and
Banos,Laguna) for their help in the field work and External Pressures. Philippine Society for the Study
data analysis. of Nature Inc., Laguna, Philippines.
The author had presented parts of the paper in Buot, I.E. Jr. and S. Okitsu. 1997a.Woody species
various symposia, namely: Philippine Society for the composition in the altitudinal zones of the mossy
Study of Nature Scientific Meeting in Cebu City, forest of Mt. Pulag, Luzon, Philippines. Flora
Philippines (May 2003) and Tagbilaran City, Malesiana Bulletin 12: 6-11.
Philippines (May 2004), and the East Asian Buot, I.E. Jr. and S. Okitsu. 1997b. Changes in
Federation of Ecological Societies Meeting in Mokpo, woody species composition and dominants along
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ature Studies 8(1) January – April 2009
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