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Undigested
Grains Triglycerides Bacterial Bacterial bacterial
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Phospholipids Phospholipids Lipids
Phospholipids
Forage Glycolipids
Saturated Saturated
Free Fatty acids Free Fatty acids
Sugar
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Triglyceride rich INTESTINAL
VFA Free Fatty Lipoprotein CELLS
RUMINAL acids
WALL (TG-rich LP)
TG-r LYMPH
ich L
P VESSEL
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PORTAL
BLOOD
(To the liver)
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carbohydrate Energy
metabolism
Glucose Triglyceride
LIVER Ketones storage
Glycerol TG-rich LP
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Fatty Acids
TG-rich LP
BLOOD
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(General
circulation)
Gly
cer Fatty Acids Glycerol TG-rich LP
ol TG-rich LP Glycerol
Mobilisation
University of Wisconsin-Madison 15
Dairy Essentials – Nutrition and Feeding
UTILIZATION OF DIETARY LIPIDS The fat deposited in the liver contributes to
BY THE UDDER development of metabolic disorders (e.g.,
About half the fat in the milk is derived ketosis and fatty liver) in early lactation.
from the uptake of fatty acids by the
ADDED LIPIDS IN DAIRY RATIONS
mammary gland. These fatty acids come
primarily from the triglyceride-rich Lipids contain about 2.25 times more
lipoproteins formed during the intestinal energy than carbohydrates. Also, lipids are
absorption of lipids. An increase in long sometimes referred to as a "cold" nutrients
chain fatty acids (i.e., acids made of more because during digestion and utilization by
than 16 carbons) in the diet increases their the body they produce less heat than
secretion in milk, but it also inhibits the carbohydrates and proteins. Thus,
synthesis of short- and medium-chain fatty increasing lipids in dairy cow rations may
acids in the mammary tissue. Thus, the have several potential benefits:
marked depression in fat secretion when • Increase the caloric (energy) density of
cows are fed low fiber diets can be the ration, especially when intake may
compensated only partially by increasing be limited as in high forage diet;
fat in the diet. • Limit the need for carbohydrate-rich
concentrates which are usually
THE ROLE OF LIVER AND FAT required in early lactation when a cow
MOBILIZATION is in negative energy balance;
During periods of under feeding or in • In hot weather, lipids may help to
early lactation, cows meet their energy reduce the heat stress of a lactating
demand by mobilizing fat from adipose cow.
tissues to obtain energy in addition to that Feed intake and milk production responses
provided by the diet. Fatty acids coming vary greatly according to the type of lipids
from the triglycerides stored in the adipose added in a diet. Cows should not be fed
tissues (located primarily beneath the hide, more than about 1.5 kg/day of lipids in
in the abdomen and over the kidneys) are addition to the lipids present in feedstuffs.
released into the blood. Mobilized fatty This amount of lipids translates into a total
acids are taken up by the liver where they of about 6 to 8% lipids in the diet before
can be used as an energy source or be negative effects become evident. Milk
converted to ketones that may be released production is maximized when lipids
in the blood and used as an energy source comprise 5% of the dietary dry matter.
by many tissues. The liver does not have a Added dietary fat usually decreases milk
high capacity to form and to export TG-rich protein by about 0.1%. In addition, excess
LP and the excess mobilized fatty acids are lipids may depress feed intake, milk
stored as triglycerides within the liver cells. production and milk fat composition.