Because these characteristics show much variation between individuals, most
ofand the lion belong to the same sister group, which diverged between 1.95 and 3.10 million years ago. Hybridisation between lion and snow leopard ancestors, however, may have continued until about 2.1 million years ago these forms were probably not true subspecies The lion's closest relatives are the other species of the genus Panthera; the tiger, snow leopard, jaguar, and leopard. Results of phylogenetic studies published in 2006 and 2009 indicate that the jaguar and the lion belong to one sister group that diverged about 2.06 million years ago. Results of later studies published in 2010 and 2011 indicate that the leopard . The genus name Panthera was coined by German naturalist Lorenz Oken in 1816.[11] Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished on the basis of appearance, size, especially because they were often based upon museum material with "striking, but abnormal" morphological characteristics.[12] . Felis leo was the scientific name used by Carl Linnaeus in 1758, who described the lion in his Based on the morphology of 58 lion skulls in three European museums, the subspecies krugeri, nubica, persica and senegalensis were assessed distinct but bleyenberghi overlapped with senegalensis and krugeri. The Asiatic lion persica was the most distinctive and the Cape lion had characteristics allying it more with persica than the other sub- Saharan lionswork Systema Naturae