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The beginning of growth, as of a seed, spore, or bud. The germination of most seeds and spores occurs
in response to warmth and water.
A Closer Look Dormant seeds are very dry and require the absorption of water to initiate the metabolic
processes of respiration and begin to digest their stored food. Respiration requires the presence of
oxygen, which must be sufficiently available in the soil for germination to proceed, so the soil must be
wet but not so waterlogged as to make oxygen inaccessible. Temperatures must be above freezing (zero
degrees Celsius) but not excessively hot (not more than about 45 degrees Celsius). If conditions are right,
a radicle (an embryonic root) emerges from the seed coat, anchoring the seed; it then grows and puts
out lateral roots. In most eudicots, a part of the developing stem, either the epicotyl (the stem above the
cotyledons) or the hypocotyl (the stem below the cotyledons) elongates, forming a hook and gradually
pulling the seed coat and the delicate shoot tip above the soil surface. Germination of eudicot seeds is
normally divided into two types, designated epigeous and hypogeous. In epigeous germination, the
cotyledons emerge above the soil surface, and wither and drop off after their food stores have been
used up; in hypogeous germination, the cotyledons remain below the surface and decompose after their
food stores have been used up. In most monocots, food is stored in the seed's endosperm (rather than
the cotyledon), and it is the single tubular cotyledon that elongates and draws the seed coat out of the
soil. The cotyledon conducts photosynthesis, making more food, while the shoot grows up inside the
tube.
panicle (pn-kl)
A branched indeterminate inflorescence in which the branches are racemes, so that each flower has its
own stalk (called a pedicel) attached to the branch. Oats and sorghum have panicles. See illustration at
inflorescence.
grain (grn)
n.
1.
a. A small, dry, one-seeded fruit of a cereal grass, having the fruit and the seed walls united: a single
grain of wheat; gleaned the grains from the ground one at a time. Also called caryopsis.
b. The fruits of cereal grasses especially after having been harvested, considered as a group: The grain
was stored in a silo.
2.
5. Abbr. gr. A unit of weight in the U.S. Customary System, an avoirdupois unit equal to 0.002285 ounce
(0.065 gram). See Table at measurement.
7.
a. The side of a hide or piece of leather from which the hair or fur has been removed.
9. The relative size of the particles composing a substance or pattern: a coarse grain.
10. A painted, stamped, or printed design that imitates the pattern found in wood, leather, or stone.
13.
v.tr.
2. To paint, stamp, or print with a design imitating the grain of wood, leather, or stone.
v.intr.
To form grains.
Idioms:
[Middle English, from Old French graine, from Latin grnum; see g-no- in Indo-European roots.]
grainer n.