A Checklist of Weevils of The Tribe Mecysolobini (Coleoptera Curculionidae Molytinae) PDF

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A checklist of weevils of the

tribe Mecysolobini (Coleoptera:
Curculionidae: Molytinae)
a a
J. P. Andrew & V. V. Ramamurthy
a
Division of Entomology, Indian Agricultural Research
Institute, New Delhi, 110012, India
To cite this article: J. P. Andrew & V. V. Ramamurthy (2010) A checklist of weevils of the tribe
Mecysolobini (Coleoptera: Curculionidae: Molytinae), Oriental Insects, 44:1, 271-336, DOI:
10.1080/00305316.2010.10417619
To link to this article: http://dx.doi.org/10.1080/00305316.2010.10417619
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Oriental Insects, Vol. 44: 271-336, 2010
A CHECKLIST OF WEEVILS OF THE TRIBE MECYSOLOBINI

(COLEOPTERA: CURCULIONIDAE: MOLYTINAE)
1
J. P. ANDREW & V.V.RAMAMURTHY
Division of Entomology
Indian Agricultural Research Institute, New Delhi 110012 India
1
Email: jon666jon @yahoo.co.in
ABSTRACT: An annotated checklist of the weevil tribe Mecysolobini (Molytinae: Curculionidae:

Coleoptera) comprising 597 species under 8 genera along with their up to date nomenclature, synonyms
and distribution is given. The analysis indicates that the tribe is almost exclusively tropical in distribution
Downloaded by [University of Windsor] at 10:20 24 July 2013
with 542 species known from Oriental and Ethiopian regions; of these 360 species are Oriental and 78
occur in India and the adjacent countries.
Key words: Mecysolobini, checklist, genera, subgenera, species, distribution
Introduction
The genus Alcidodes Marshall is one of the largest, economically important weevils
of which the taxonomy needs to be worked out. The review of literature as a prelude
to undertaking the revisionary studies on this and allied genera revealed that there are
a large number of species known, especially from the Oriental region. The taxonomy
of these species has been poorly known and the problem is compounded with
changing concepts at the subfamily, tribes and the generic levels. Hence, efforts were
made to prepare an annotated checklist of the species known so far under the tribe
Mecysolobini Reitter as a result of which 597 species had been enlisted. While doing
so a review of the changes in the subfamily, tribe and generic concepts so far had also
been catalogued. The sole purpose of this checklist is to annotate the available details
so as to serve as a prelude to an impending revisionary study. Hence, it refrains from
assessing and commenting on the nomenclatural problems but presents them on as is
where basis. All the efforts have been made to confer with original publications and
brief review of these details presented herein.
The generic nomenclature in the molytine tribe Mecysolobini is confused, reflecting

both the large amount of convergence and parallelism that has undoubtedly occurred
in the group, and the occasionally weak grasp of the Rules of Zoological
Nomenclature of some of the taxonomists who have studied this tribe (Lyal and
Curran, 2000). There are twentytwo available genus group names in the tribe, their
present usage had been summarized (Fig.l).
The most recent revision on this group under the name Alcidodinae (Pajni and Dhir,
1987) has been observed to be erronaeus in many respects. A review of the literature,
both taxonomic and applied reveal that currently a vast majority use incorrect generic
names. Thus, the challenge for the taxonomist is to preserve nomenclatural stability
as far as possible, while trying to reflect some sort of evolutionary hypothesis and
conform with the rules (Lyal and Curran, 2000).
Some workers, particularly in applied fields, still incorrectly employ Alcides

Schoenherr (1825), but it is a junior homonym (and synonym) of Alcides Sahlberg
272 Oriental Insects Vol. 44
(1823), and a junior homonym of a lepidopteran Alcides Hubner (1822). This was
brought out by Marshall (1939), who proposed the replacement name Alcidodes.
Heller (1918) who divided the same into six subgenera viz. Cylindralcides Heller,
Sternuchopsis Heller, Robustalcides Heller, Granosalcides Heller, Ornatalcides
Heller and Metallalcides Heller, subsequently expressed the inadequacy of these.
Later, Voss (1953; 1956b; 1957; 1958; 1962) put fourth conflicting views over a
decade creating more confusion which led Haaf (1961d; 1964b) to ignore these
views. Pajni and Dhir (1987) while attempting to revise its species from India
proposed deploying generic names namely Brahmalcidodes Pajni and Dhir,
Mecysolobus Reitter, Neomecyslobus Pajni and Dhir, Tuberculomecyslobus Pajni and
Dhir, Cylindralcidodes Pajni and Dhir, Mesalcidodes Voss and Alcidodes. Likewise
the works of Voss (1953) and Haaf (1960-1964) led to more confusion than clarity as
brought out by Lyal and Curran (2000). Thus, the Mecysolobini needs a thorough
revision, as reliable limits to many of the genera remain uncertain. That is why until a
cladistic analysis is done to determine relationships, the most stable approach

suggested is to the species groups retained in Alcidodes as generic names (Lyal and
Curran, 2000).
An analysis of the fauna of the tribe Mecysolobini indicates that most of its species
were known before 1970 and the descriptions are scattered, and the classification
unrealistic. An analysis of the faunistics reveal that, of the 597 species most had been
described between 1891-1950 and 1951-1970, with about 263 and 161 species
respectively. Contributions of Faust, Heller, Marshall, Fairmaire, Haaf and Voss had
been more significant. Except for a few isolated descriptions, no comprehensive
studies have been undertaken recently indicating that the revisionary studies are
imminent. This also reveals the fact that zoogeographically it is predominantly
tropical with 542 species known from Oriental and Ethiopian regions; of these 360
species are Oriental and 78 occur in India and the adjacent countries (Table 1).
Tribe Mecysolobini
1. Alcidodoplesius Voss
Alcidoplesius Voss, 1960c: 314; Voss, 1960d: 138
Type Alcidodoplesius tarsalis Voss, 1960d: 138
1. tarsalis Voss, 1960d: 138 Madagascar
2. parvulusVoss, 1961:111 Philippines
2. Brahmalcidodes Pajni and Dhir, 1987: 29
Type Alcidodes micronychus Pascoe, 1872: 483
1. micronychus (Pascoe) Myanmar, India,
Alcides micronychus Pascoe, 1872: 483; Faust, Thailand, Vietnam
1894a: 242, 258; Marshall, 1939: 570; Pajni and
Dhir, 1987: 30
2. nigromicronychus Pajni and Dhir, 1987: 30 India
3. Cylindralcides Heller
Cylindralcides Heller, 1918: 211; Pajni and Dhir,
1987: 32 (Cylindralcidodes); Alonzo-Zarazaga and
Lyal, 1999: 205
1. aemulus (Faust) Myanmar, India
Alcides aemulus Faust, 1894a: 231, 239; Marshall,
2010 Andrew & Ramamurthy: Checklist, Molytinae 273
1939: 570; Pajni and Dhir, 1987: 32

2. affaber (Aurivillius) Laos, India
Alcides affaber Aurivillius, 1892: 218; Heller,
1911:314; Marshall, 1939: 570; Pajni and Dhir,
1987: 32
3. ashwanae Pajni and Dhir, 1987: 32 India
4. barbicus (Haaf) India, Myanmar
Alcidodes barbicus Haaf, 1964b: 162; Pajni and
Dhir, 1987: 32
4a barbicus sub spfalsus Haaf, 1964b: 159 India, Myanmar
5. bicolorous Pajni and Dhir, 1987: 32 India
6. bubo (Fabricius) Australia, India
Rhynchaenus bubo Fabricius, 1801: 474; Boheman
in Schoenherr, 1836: 612 (Alcides); Marshall,
1939: 570 (Alcidodes); Pajni and Dhir, 1987: 32

Rhynchaenus ferox Olivier,l807:
189;Klima,1934:52
Ale ides terraereginae Blackburn, 1900: 141 ;Klima,
1934: 52
7. castaneipennis (Hustache) Gabon
Alcides castaneipennis Hustache, 1922: 151;
Marshall, 1939: 570; Voss, 1959b: 413
8. corniculus (Faust) Myanmar, India
Alcides corniculus Faust, 1894a: 250, 259;
Marshall, 1939: 570; Pajni and Dhir, 1987:32
9. desertus (Haaf) India
Alcidodes desertus Haaf, 196la: 17; Pajni and Dhir,
1987: 32
10. difficilis Heller, 1918: 220 Philippines
11. fervidus (Haaf) India
Alcidodes fervidus Haaf, 1961 a: 15; Pajni and Dhir,
1987: 32
longirostris Heller, 1918: 211, 221 to
Sternuchopsis: Haaf, 1962: 153
12. ludificator (Faust) Myanmar, India
Alcides ludificator Faust, 1894a: 254, 260;
Marshall, 1939: 570; Pajni and Dhir, 1987: 32
13. major (Haaf) India
Alcidodes major Haaf, 1964b: 167; Pajni and Dhir,
1987:32
14. mali (Marshall) India
Alcides mali Marshall, 1919: 276; 1939: 570; Pajni
and Dhir, 1987: 32
15. miser Heller, 1918: 218 Philippines
16. mustela (Pascoe) Vietnam, Singapore,
Alcides mustela Pascoe, 1882: 446; Heller, 1911: Indonesia,
314; 1918: 219; Marshall, 1939: 570 Philippines
17. mysticus (Faust) Myanmar, India
Alcides mysticus Faust, 1894a: 246, 258; Heller,
1911: 314; Marshall, 1939: 570; Pajni and Dhir,

1987: 32
18. obesus (Faust) Myanmar, India,
Alcides obesus Faust, 1894a: 248, 259; Heller, Cambodia, Laos,
1911: 314; Haaf, 1964b: 155; Pajni and Dhir, 1987: Vietnam, Malaysia,
32 Singapore
Alcides robustus Hustache, 1933: 83; Haaf, 1964b:
155
19. persegnis Heller, 1918: 222 Philippines
20. pius Heller, 1918: 219 Philippines
21. rufescens Pajni and Dhir, 1987: 32 India
22. saundersi (Pascoe) Myanmar, Thailand,
Alcides carbonarius Pascoe, 1888: 412; Klima, Indonesia, India,
1934: 58; Marshall, 1939: 570 Philippines,
Alcides lichen Faust, 1894a: 244, 258; Haaf, 1961e: Cambodia, Laos,
485 Vietnam, Malaysia,
Alcides luxuriosus Faust, 1894a: 244, 258; Heller, Singapore
1922a: 19; Haaf, 1961e: 485
Alcides saundersi Pascoe, 1870b: 459; Faust,
1894a: 244, 258; Marshall, 1930: 575; Haaf, 1961e:
485; Haaf, 1964b: 174; Pajni and Dhir, 1987:32
Alcides umenoi Kono, 1937b: 15; Haaf, 1964b: 174
23. separandus (Haaf) India
Alcidodes separandus Haaf, 1961 a: 16; Pajni and
Dhir, 1987: 32
24. serratidens Heller, 1918: 216 Philippines
25. tutus (Faust) Myanmar, India
Alcides tutus Faust, 1894a: 253, 259; Marshall,
1939: 570; Pajni and Dhir, 1987: 32
26. vafellus (Faust) India
Alcides vafellus Faust, 1898: 308; Marshall, 1939:
570; Pajni and Dhir, 1987: 32
4. Merus Gistel, 1857: 606
Type Lixus fasciatus Redtenbacher, 1848: 548
1. artivittis (Marshall) India, Myanmar
Alcidodes artivittis Marshall, 1948: 428; Pajni and
Dhir, 1987: 31 (Mecyslobus); Alonso-Zarazaga and
Lyal; 1999: 205
2. assamensis (Pajni and Dhir) India
Mecyslobus assamensis Pajni and Dhir, 1987: 30;
Alonso-Zarazaga and Lyal; 1999: 205
3. butuanensis Schultze Philippines
Metallalcides butuanensis Schultze, 1922: 589;
Alonso-Zarazaga and Lyal; 1999: 205
4. delicatulus (Haaf) India
Alcidodes delicatulus Haaf 1961: 23; Pajni and
Lyal; 1999: 205
5. dentitibialis (Voss) Sri Lanka
Mecysolobus dentitibialis Voss, 1957: 103; Alonso-

Zarazaga and Lyal; 1999: 205
6. deremius (Kolbe) Tanzania, Uganda,
Alcides deremius Kolbe, 1898: 266; Marshall, Kenya
1939: 570; Alonso-Zarazaga and Lyal, 1999: 205
7. erro (Pascoe) China, Japan
Alcides erro Pascoe, 1871: 182; Roelofs, 1875:
151; Heller, 1911: 314; Marshall, 1939: 570; Zhou,
1998: 38 (Mecysolobus); Alonso-Zarazaga and
Lyal, 1999: 205
8. fasciatus (Redtenbacher) India
Lixus fasciatus Redtenbacher in Hugel, 1848: 548;
Lacordiare, 1866: 16 (Alcides); Klima, 1934: 43;
Marshall, 1939: 570 (Alcidodes); Pajni and Dhir,
1987: 30 (Mecyslobus); Alonso-Zarazaga and Lyal,

1999:205
9. gubatanus (Schultze) Philippines
Metallalcides gubatanus Schultze, 1922: 590;
Alonso-Zarazaga and Lyal, 1999: 205
10. josephus (Duvivier) Congo
Alcides josephus Duvivier, 1892: 312; Marshall,
1939: 570; Alonzo-Zarazaga and Lyal, 1999: 205
lOa josephus var compressus Hustache, 1934: 355 Congo
11. kilimanus (Aurivillius) Tanzania, Kenya
Alcides kilimanus Aurivillius, 1910: 427; Voss,
1962a: 8 (Alcidodes); Alonso-Zarazaga and Lyal,
1999:205
lla kilimanus sub sp discedens Voss, 1962a: 8 Tanzania
llb kilimanus var. kenyae Hustache, 1829: 508 Kenya
12. lamprocolius (Heller) Philippines
Metallalcides lamprocolius Heller, 1924: 184;
13. leopardus (Olivier) India
Lixus alboguttatus Wiedemann, 182: 162;
Boheman in Schoenherr, 1836: 612 (Alcides);
Klima, 1934: 55; Marshall, 1939: 570 (Alcidodes);
Rhynchaenus leopardus Olivier, 1807: 190, 296;
Bovie, 1908a: 6 (Alcides); Marshall, 1939: 570
(Alcidodes); Pajni and Dhir, 1987: 33
(Mesalcidodes); Alonso-Zarazaga and Lyal, 1999:
205
14. lixoides (Reitter) Japan, China,
Alcidesflavosignatus Roelofs, 1875: 151; Reitter, Russia, Mongolia,
1905: 248; Marshall, 1939: 570; Alonso-Zarazaga India
and Lyal, 1999: 205
Mecysolobus lixoides Reitter, 1905: 248; Pajni and
Lyal, 1999: 205
15. mangyanicus (Schultze) Philippines

Metallalcides mangyanicus Schultze, 1922: 590;
16. mervanoides (Voss) Congo
Mecysolobus mervanoides Voss, 1962b: 276;
17. montanus (Haat) Cambodia, Laos,
Alcidodes montanus Haaf, 1964b: 160; Pajni and Vietnam, Malaysia,
Dhir, 1987: 30 (Mecyslobus); Alonso-Zarazaga and Myanmar, India
Lyal, 1999: 205 Singapore, Thailand
18. obfuscatus (Fabricius) Indonesia, India,
Alcides pavidus Faust, 1894a: 251, 259; Haaf, Myanmar
196lb: 434; Pajni and Dhir, 1987: 30; Alonso-
Zarazaga and Lyal, 1999: 205
Alcides raptorius Boheman in Schoenherr, 1836:

610; 1844: 50; Marshall, 1939: 570; Alonso-
Lixus obfuscatus Fabricius, 1775: 138; 1801: 505;
Pajni and Dhir, 1987: 30 (Mecyslobus); Alonso-
19. obfuscatus Voss Congo
Mecysolobus obfuscatus Voss, 1962b: 275; Alonso-
20. semperi (Pascoe) Philippines
Metallalcides obsidianus Heller, 1918: 242; 1921:
197; Alonso-Zarazaga and Lyal, 1999: 205
Alcides opacimaculatus Heller, 1918: 242;
Marshall, 1939: 570; Alonso-Zarazaga and Lyal,
1999: 205
Alcides semperi Pascoe, 1870b: 462; Roelofs,
1893: 34; Bovie, 1908a: 8; Heller, 1918: 242
(Metallalcides); Marshall, 1939: 570; Alonso-
20a semperi sub sp ocellatus Roelofs Philippines
Alcides ocellatus Roelofs, 1893: 35; Heller, 1918:
216, 243; 1921: 197; Marshall, 1939: 570; Alonso-
21. scenicus (Faust) Japan, Myanmar,
Alcides scenicus Faust, 1894a: 256, 260; Bovie, India, Vietnam
1908a: 8; Marshall, 1939: 570; Pajni and Dhir,
1999:205
21a scenicus sub sp intrusomimus Heller, 1922a: 19 Japan, Myanmar,
India, Vietnam
22. siccus (Haat) India
Alcidodes siccus Haaf, 1964b: 163; Pajni and Dhir,
1999: 205
23. smaragdinus (Roelofs) Philippines
Alcides smaragdinus Roelofs, 1893: 39; Heller,

1918: 216 (Metallalcides); Marshall, 1939: 570
24. tenuisignatus (Voss) Africa
Mecysolobus tenuisignatus Voss, 1956a: 180;
25. trigonophorus (Marshall) India
Alcidodes trigonophorus Marshall, 1922: 394, 396;
1939: 570; Pajni and Dhir, 1987: 30 (Mecyslobus);
26. tshibindanus (Voss) Congo
Mecysolobus tshibindanus Voss, 1962b: 275;
26a tshibindanus sub sp lusingaensis Voss, 1962b: 275 Congo
27. westermanni (Boheman) India
Alcides westermanni Boheman in Schoenherr,

1844: 51; Marshall, 1939: 570; Pajni and Dhir,
1999:205
5 Neomecyslobus Pajni and Dhir, 1987: 31
Type Alcidodes loratus Marshall, 1922: 394, 397
1. audax (Heller) India
Alcides audax Heller, 1908b: 164; Marshall, 1939:
2. feae (Faust) Myanmar, India
Alcides feae Faust, 1894a: 245,258; Marshall,
1939: 570; Pajni and Dhir, 1987: 31
3. gardneri Pajni and Dhir, 1987: 31 India
4. habenatus (Marshall) India
Alcidodes habenatus Marshall, 1922: 395;
5. loratus (Marshall) India, Myanmar
Alcidodes loratus Marshall, 1922: 394, 397;
6. mellitus (Faust) India
Alcides mellitus Faust, 1898: 308; Marshall, 1939:
7. porrectirostris (Marshall) India
Alcidodes porrectirostris Marshall, 1913: 188;
Marshall, 1939:570; Pajni and Dhir, 1987: 31
8. saturnus (Haaf) India
Alcidodes saturnus Haaf, 1961a: 20; Pajni and
Dhir, 1987: 31
9. squamosus Pajni and Dhir, 1987: 31 India
10. strobilanthesia Pajni and Dhir, 1987: 31 India
11. vinculosus (Heller) India
Alcides vinculosus Heller, 1908b: 160, 161;
6 Ornatalcides Heller, 1918: 214
Type Alcides decoratus Roelofs, 1893: 37
1. anamalaiensis (Pajni and Dhir) India

Mesalcidodes anamalaiensis Pajni and Dhir, 1987:
cuprinus Heller, 1918: 241 to Sternuchopsis: Haaf,
1961e:490
2. decoratus (Roelofs) Philippines
Alcides decoratus Roelofs, 1893: 37; Heller, 1918:
215 (Ornatalcides); Marshall, 1939: 570; Alonso-
3. fuscocupreus Pajni and Dhir India
Mesalcidodesfuscocupreus Pajni and Dhir, 1987:
4. hypocritus (Boheman) Indonesia, India
Alcides hypocritus Boheman in Schoenherr, 1844:
47; Marshall, 1939: 570; Pajni and Dhir, 1987: 33

(Mesalcidodes); Alonso-Zarazaga and Lyal, 1999:
205
5. ignoratus Heller, 1918: 238 Philippines
6. improvidus (Faust) Pajni and Dhir, 1987: 33 Myanmar, India
Alcides improvidus Faust, 1894a: 241, 257; Bovie,
1908a: 6; Pajni and Dhir, 1987: 33 (Mesalcidodes);
7. kalinganus Schultze, 1923: 604 Philippines
leopardus Olivier, 1807: 190, 296 to Merus:
Alonzo-Zarazaga and Lyal, 1999: 205
8. leucospilus (Erichson) Philippines,
Alcides leucospilus Erichson, 1834: 264; Heller, Indonesia
1918: 240 (Ornatalcides); Haaf, 1961e: 493
(Alcidodes); Alonso-Zarazaga and Lyal, 1999: 205
Alcidodes leytensis Haaf, 1961e: 493
Alcides mindanaoensis Schultze, 1918: 271; Haaf,
1961e: 493 (Alcidodes)
8a leucospilus sub sp burmeisteri Boheman in Schoenherr
Alcides burmeisteri Boheman in Schoenherr, 1845:
448; Pascoe, 1885: 242; Heller, 1918: 237; Haaf,
1961e: 493; Alonso-Zarazaga and Lyal, 1999: 205
8b leucospilus var confluens Heller, 1918: 240
8c leucospilus var erichsoni Heller, 1918: 240
8d leucospilus var subaeneus Heller, 1918: 237
8e leucospilus var honestus Heller, 1918: 240
9. nocens (Heller) India
Alcides nocens Heller, 1908b:l62; Marshall, 1939:
570; Pajni and Dhir, 1987: 33 (Mesalcidodes);
10. rutilans (Roelofs) Philippines
Alcides rutilans Roelofs, 1893: 38; Heller, 1918:
215
11. satelles (Faust) Myanmar, India
Alcides satelles Faust, 1894a: 240, 258; Bovie,
1908a:8; Marshall, 1939: 570; Pajni and Dhir,

1987: 33 (Mesalcidodes); Alonso-Zarazaga and
Lyal, 1999: 205
12. septentrionalis Heller, 1918: 243 Philippines
13. serotinus (Haaf) India
Alcidodes serotinus Haaf, 1961a: 48; Pajni and
Dhir, 1987: 33 (Mesalcidodes); Alonso-Zarazaga
and Lyal, 1999: 205
14. trifidus (Pascoe) Taiwan, India,
trifidus sub sp taiwanus Kono, 1930: 139; Haaf, Japan, Korea, China
1964b: 199
Alcides trifidus Pascoe, 1870b: 459; Marshall,
1939: 570 (Alcidodes); Haaf, 1964b: 199; Pajni and
and Lyal, 1999: 205

15. vitalisi (Marshall) Cambodia, Laos,
Alcidodes vitalisi Marshall, 1922: 394, 409; 1939: Vietnam, Malaysia,
570; Pajni and Dhir, 1987: 33 (Mesalcidodes); Myanmar,
Alonso-Zarazaga and Lyal, 1999: 205 Singapore, Thailand,
India
16. waltoni (Boheman) Pajni and Dhir, 1987: 33 Philippines,
Alcides albolineatus Roelofs, 1875: 152; Klima, Cambodia, India,
1934: 44; Marshall, 1939: 470 Japan, Indonesia,
Alcides roelofsi Lewis, 1879: 23; Kono, 1930: Taiwan, Sri Lanka,
138,142; Haaf, 1961b: 412 (Alcidodes) Myanmar
Alcides sexvittatus Faust, 1895a: 243,258; Klima,
1934:44
Alcides waltoni Boheman in Schoenherr, 1844: 58;
Heller, 1918: 212, 223 (Sternuchopsis); Pajni and
and Lyal, 1999: 205
7. Sternuchopsis Heller, 1918: 212
Type Alcides pectoralis Boheman in Schoenherr,
1836: 618
1. abbreviatus (Heller) Papua New Guinea
Alcides abbreviatus Heller, 1942: 211; Marshall,
1939: 570; A1onso-Zarazaga and Lyal, 1999: 206
2 achteni (Hustache) Congo
Alcides achteni Hustache, 1934: 352; Marshall,
2a achteni sub sp sobrinus Haaf, 1961c: 56 Congo
acutus Faust, 1898: 310 to Tuberculomecyslobus:
Pajni and Dhir, 1987: 31
3 adspersus (Boheman) Cape Colony
Alcides adspersus Boheman, 1859: 134; Marshall,
4 adustus (Haaf) Indonesia
Alcidodes adustus Haaf, 1961b: 392; Alonso-
5 adversarius (Schultze) Philippines

Alcides adversarius Schultze, 1919: 338; Marshall,
aemulus Faust, 1894a:231,239 to Cylindralcidodes:
6 aeratus (Heller) Philippines, Samoa
Alcides aeratus Heller, 1922b: 554; Marshall,
6a aeratus sub sp interpostitus Heller, 1924: 185 Samoa, Philippines
affaber Aurivillius, 1892: 218 to Cylindralcides:
Heller, 1911: 314
affinis Fahraeus, 1871: 256 see dentipes (Olivier)
7 agnatus (Haaf) Indonesia,Malaysia
Alcidodes agnatus Haaf, 1964a: 19; Alonso-

8 albocinctus (Blanchard) Indonesia
Alcides albocinctus Blanchard in Hombron and
Jacquinot, 1853: 246; Marshall, 1939: 570; Alonso-
9 alboguttatus (Schoenherr) Congo
Alcides alboguttatus Schoenherr, 1825: 581;
1999:206
alboguttatus Wiedemann, 1823: 162 see leopardus
(Olivier)
10 albolineatus (Boheman) Gabon, Senegal,
Alcides albolineatus Boheman in Schoenherr, Congo, Tanzania,
1836: 612; Marshall, 1939: 570; Alonso-Zarazaga Germany, South
and Lyal, 1999: 206 Africa
albolineatus Roelofs, 1875: 152 see waltoni
11 albolituratus (Blanchard) Indonesia
Alcides albolituratus Blanchard in Hombron and
12 albomaculatus (Kono) Taiwan
Alcides albomaculatus Kono, 1930: 140; Marshall,
13 albopictus (Fairmaire) Madagascar
Alcides albopictus Fairmaire, 1904: 247; Marshall,
14 alfkeni (Heller) Philippines
Alcides alfkeni Heller, 1934: 295; Marshall, 1939:
15 alienus (Haaf) Africa
Alcidodes alienus Haaf, 1961c: 50; Alonso-
amandus Haaf, 1961c: 102 see depressipennis sub
sp amandus
16 ambatoides (Voss) Congo
Alcidodes ambatoides Voss, 1962b: 267; Alonso-

17 ambroensis (Hustache) Madagascar
Alcides ambroensis Hustache, 1933: 82; Marshall,
18 amitinus (Kolbe) Kenya, Tanzania,
Alcides amitinus Kolbe, 1898: 264; Marshall, 1939: Uganda
19 amoenus (Pascoe) Indonesia
Alcides amoenus Pascoe, 1882: 448; Haaf, 1963:
Alcides bellus Pascoe, 1882: 448; Haaf, 1963: 112
20 amplus (Haaf) Philippines
Alcidodes amplus Haaf, 1961e: 479; Alonso-

analogus Heller, 1918: 244 see nitidus (Pascoe)
anceyi Bovie 1908b: 43 see humerosus (Ancey)
21 androycus (Fairmaire) Madagascar
Alcides androycus Fairmaire, 1902a: 367; Marshall,
22 anfractus (Heller) Indonesia
Alcides anfractus Heller, 1933: 101, 103; Marshall,
23 angolanus (Hustache) Angola
Alcides angolanus Hustache, 1932: 66; Marshall,
1939:470
24 angulus (Fabricius) Indonesia, Malaysia,
Rhynchaenus angulus Fabricius, 1801: 464; Cambodia
Boheman in Schoenherr, 1844: 46 (Alcides); Haaf
1961b: 394; Alonso-Zarazaga and Lyal, 1999: 206
Alcides geometricus Pascoe, 1885: 243; Haaf
1961b:394
Alcides lacumsus Boheman in Schoenherr, 1844:
47; Haaf 1961b: 394
25 angulus (Schoenherr) Congo
Alcides angulus Schoenherr, 1825: 581; Marshall,
26 angustus (Haaf) Indonesia
Alcidodes angustus Haaf, 1961b: 397; Alonso-
27 annulatus (Hustache) Senegal
Alcides annulatus Hustache, 1932: 65; Marshall,
28 antenna/is (Faust) Togo, West Africa
Alcides antenna/is Faust, 1894b: 535; Marshall,
29 apicalis (Heller) Papua New Guinea
Alcides apicalis Heller, 1914: 653; Marshall, 1939:
30 arcanus (Haaf) Indonesia

Alcidodes arcanus Haaf, 1964a: 44; Alonso-
31 arcicollis (Marshall) Indonesia
Alcides arcicollis Marshall, 1932: 341; 1939: 570;
arcuatus Schultze, 1919: 559 see schultzei
32 arcuatus (Boheman) Germany, Cape
Alcides arcuatus Boheman in Schoenherr, 1844: Colony, Angola,
46; Haaf, 1961c: 79; Alonso-Zarazaga and Lyal, Africa
1999:206
32a arcuatus var maculiventris Hustache, 1924: 81
32b arcuatus sub sp stolzi Haaf, 1961c: 79
33 arenatus (Schultze) Philippines
Alcides arenatus Schultze, 1919: 559; Marshall,

34 argutor (Pascoe) Srilank:a
Alcides argutor Pascoe, 1883: 126; Marshall, 1939:
35 armiger (Thunberg) Cape Colony
Ale ides armiger Thunberg, 1815: 114; Marshall,
artivittis Marshall, 1948: 428 to Merus: Alonso-
Zarazaga and Lyal; 1999: 205
aschanticus Faust, 1895a: 236 see guessfeldti
(Kolbe) ashinagazo Kono, 1930: 142 see yoshidai
Kono
36 asperatus (Haaf) Indonesia
Alcidodes asperatus Haaf, 1963: 141; Alonso-
aspericollis Heller, 1929: 114,115 see vestitus
(Pascoe)
37 asphaltinus (Pascoe) Indonesia
Alcides asphaltinus Pascoe, 1870b: 461; Marshall,
atrocretosus Fairmaire, 1883b: 37 see elegans
(Guerin-Meneville)
audax Heller, 1908b: 164 to Neomecyslobus: Pajni
andDhir, 1987:31
38 auritus (Pascoe) Vietnam
Alcides auritus Pascoe, 1873: 182; Marshall, 1939:
39 australis (Boisduval) Indonesia, Papua
Alcides australis Boisduval, 1835: 423; Haaf, 1963: New Guinea
Alcides profluens Pascoe, 1885: 239; Klima, 1934:
51
39a australis sub sp rectus Haaf, 1963: 127 Indonesia
australis subsp. ternatensis see ternatensis
40 avidus (Haaf) Cambodia, Laos,

Alcidodes avidus Haaf, 1964b: 159; Alonso- Vietnam, Malaysia,
Zarazaga and Lyal, 1999: 206 Myanmar,
Singapore, Thailand
41 balachowskyi (Hoffmann) West Africa
Alcidodes balachowskyi Hoffmann, 1962: 124;
barbicus Haaf, 1964b: 162 to Cylindralcidodes:
42 basipennis (Marshall) Myanmar
Alcidodes basipennis Marshall, 1948: 429; Alonso-
43 baueri (Haaf) Indonesia
Alcidodes baueri Haaf, 1963: 100; Alonso-

44 beatus (Haaf) Indonesia
Alcidodes beatus Haaf, 1964a: 59; Alonso-
bellus Pascoe, 1882: 448 see amoenus (Pascoe)
biai Duvivier, 1890: 24 see olivaceus (Gerstacker)
45 biangulatus (Marshall) Kenya, Uganda,
Alcidodes biangulatus Marshall, 1928a: 407; 1939: Zimbabwe,
570; Alonso-Zarazaga and Lyal, 1999: 206 Tanzania, Nigeria
46 bicinctus (Faust) Malaysia
Alcides bicinctus Faust, 1895c: 97; Marshall, 1939:
47 bifasciatus (Aurivillius) Kenya
Alcides bifasciatus Aurivillius, 1910: 428;
1999: 206
bifasciatus Hustache, 1922: 152 see bivinculatus
(Heller)
48 bimaculatus (Faust) Myanmar
Alcides bimaculatus. Faust, 1894a: 253, 260;
1999:206
49 bimbiense (Hustache) Angola
Alcides bimbiense Hustache, 1935: 165; Marshall,
bioculatus Heller, 1933: 98 see bivinculatus
(Heller)
50 biornatus (Fairmaire) Madagascar
Alcides biornatus Fairmaire, 1903: 231; Marshall,
51 biplagiatus (Haaf) India
Alcidodes biplagiatus Haaf, 1961 a: 41; Alonso-
52 bipunctatus (Hustache) Zanzibar
Alcidodes bipunctatus Hustache, 1932: 60; Alonso-

53 bisignatus (Pascoe) Indonesia
Alcidodes bisignatus Pascoe, 1882: 447; Alonso-
54 bivinculatus (Heller) Kenya, Congo
Alcides bifasciatus Hustache, 1922: 152; Klima,
1934:45
Alcides bivinculatus Heller, 1933: 98; Haaf, 1961c:
Alcides bioculatus Heller, 1933: 98; Klima, 1934:
45
54 a bivinculatus sub spjueundus Haaf, 1961c: 124
54b bivinculatus sub sp suratus Haaf, 1961c: 123
55 blanchardi (Faust) Togo
Alcides blanchardi Faust, 1895a: 97; Marshall,

blanchardi Faust, 1899a: 45 see sturnus Haaf
56 blandus (Haaf) Indonesia
Alcidodes blandus Haaf, 1963: 116; Alonso-
57 bohemani (Heller) Philippines,
Alcides bohemani Heller, 1918: 233; Haaf, 1961e: Thailand(?)
Alcides catanduanensis Schulze, 1918: 273; Klima,
1934:51
58 bonguensis (Hartmann) Ghana
Alcides bonguensis Hartmann, 1900: 276; Marshall,
59 brevicollis (Pascoe) Papua New Guinea,
Alcides brevicollis Pascoe, 1885: 243; Haaf, 1963: Indonesia
59 a brevicollis sub sp reductus Faust Papua New Guinea,
brevicollis sub sp reductus Faust, 1892b: 208; Indonesia
Haaf, 1963: 116
Alcides reductus Faust, 1892b: 208; Marshall,
1939: 570; Haaf, 1963: 116
60 brevior (Heller) Malaysia
Alcides brevior Heller, 1933: 101,102; Marshall,
61 brevirostris (Boheman) Equatorial Guinea,
Alcides brevirostris Boheman in Schoenherr, 1836: SouthAfrica,
621; Haaf, 1962: 140; Alonso-Zarazaga and Lyal, Tanzania,
1999:206 Madagascar
61a brevirostris sub sp curtirostris Fairmaire
brevirostris sub sp curtirostris Fairmaire, 1871: 50;
Haaf, 1962: 140
Alcides curtirostris Fairmaire, 1871: 50; Marshall,
1939: 570; Haaf, 1962: 140
61b brevirostris var obscurus Aurivillius
brevirostris var p Boheman in Schoenherr, 1844:

62; Aurivillius, 1910: 427
62 brevithorax (Hustache) Madagascar
Alcides brevithorax Hustache, 1920b: 10; Marshall,
63 brunigueli (Roudier) Gabon, Congo,
Alcidodes brunigueli Roudier, 1957: 214; Alonso- Central African
Zarazaga and Lyal, 1999: 206 Republic, Chad
64 bryanti (Marshall) Indonesia
Alcidodes bryanti Marshall, 1922: 394,407; 1939:
65 bubulus (Heller) Philippines
Alcides bubulus Heller, 1918: 213, 227; Marshall,
65a bubulus var. punctiger Heller

bubulus var punctiger Heller, 1918: 213; Klima,
1934:51
bubulus var.an subsp? punctiger Heller, 1918: 228
bubo Fabricius, 1801: 474 to Cylindralcidodes:
66 burgeoni (Hustache) Congo
Alcides burgeoni Hustache, 1934: 349; Marshall,
burmeisteri Boheman in Schoenherr, 1845: 448 see
leucospilus
67 chaudoiri (Chevrolat) Armenia, Iran,
Alcides chaudoiri Chevrolat in Guerin-Meneville, Kazakhistan, Russia,
1844: 154; Marshall, 1939: 570; Alonso-Zarazaga Turkmenistan
and Lyal, 1999: 206
Alcides karelinii Boheman in Schoenherr, 1844: 65;
Klima, 1934: 43; Marshall, 1939: 570
68 camerunus (Hustache) Cameroon
Alcides camerunus Hustache, 1932: 61; Marshall,
69 candidulatus (Haaf) Africa
Alcidodes candidulatus Haaf, 1961c: 62; Alonso-
carbonarius Pascoe, 1888: 412 see saundersi
Pascoe
70 carus (Haaf) Africa
Alcidodes carus Haaf, 1961c: 75; Alonso-Zarazaga
and Lyal, 1999: 206
castaneipennis Hustache, 1922: 151 to
Cylindralcides: Voss, 1959b: 413
71 castus (Haaf) Africa
Alcidodes castus Haaf, 1961c: 64; Alonso-Zarazaga
and Lyal, 1999: 206
catanduanensis Schulze, 1918: 273 see bohemani
Heller
72 caudatus (Hustache) Madagascar

Alcides caudatus Hustache, 1933: 80; Marshall,
73 caviventris Lyal in Lyal and Curran, 2000: 1831 Indonesia
centroguttatus Marshall, 1926b: 357 see
hexamitosus (Heller)
74 censorius (Pascoe) Indonesia
Alcides censorius Pascoe, 1887: 355; Marshall,
75 ceramodelta (Marshall) Indonesia
Alcidodes ceramodelta Marshall, 1918a: 153, 156;
76 cervinus (Haaf) Indonesia, Papua
Alcidodes cervinus Haaf, 1963: 123; Alonso- New Guinea

77 ceylonensis (Heller) Sri Lanka
Alcides ceylonensis Heller, 1901: 344; Marshall,
78 chalcomorphus (Heller) Philippines
Alcides chalcomorphus Heller, 1922b: 553;
1999:206
79 chiliarchus (Boheman) Indonesia
Alcides chiliarchus Boheman in Schoenherr, 1836:
609; Haaf, 1964a: 72; Alonso-Zarazaga and Lyal,
1999:206
79a chiliarchus sub sp effertus Haaf, 1964a: 72 Indonesia
80 cinchonae (Marshall) Indonesia
Alcidodes cinchonae Marshall, 1923: 269; 1939:
81 circulifer (Heller) Philippines
Alcides circulifer Heller, 1924: 186; Marshall, Philippines
Alcides fraudator Heller, 1924: 186; Klima, 1934:
52
82 circumscutellatus (Hustache) Ethiopia
Alcides circumscutellatus Hustache, 1932: 62;
1999:206
clathratus Aurivillius, 1892: 219 seefrenatus Faust
83 clathratus (Bertolini) Mozambique
Alcides clathratus Bertolini, 1849: 426; Marshall,
84 clausus (Walker) Sri Lanka
Alcides clausus Walker, 1859: 264; Marshall, 1939:
570; Haaf, 1960: 663; Alonso-Zarazaga and Lyal,
1999:206
Sternuchopsis keiseri Voss, 1957: 107; Haaf, 1960:
663
85 collaris (Pascoe) India

Alcides collaris Pascoe, 1887: 358; Marshall, 1939:
86 collarti (Hustache) Congo
Alcides collarti Hustache, 1934: 353; Marshall,
87 comiculus (Faust) Myanmar
Alcides comiculus Faust, 1895a: 250; Marshall,
88 comma (Kolbe) Tanzania, Kenya,
Alcides comma Kolbe, 1898: 264; Marshall, 1939: Uganda
89 comparabilis (Bovie) India
Alcides comparabilis Bovie, 1908a: 3; Marshall,

Alcides geminus Heller, 1908b: 159,161; Klima,
1934: 52; Marshall, 1939: 570
90 comptus (Haaf) Sri Lanka
Alcidodes comptus Haaf, 1960: 655; Alonso-
91 congoanus (Faust) Congo
Alcides congoanus Faust, 1899b: 413,414;
1999:206
92 confusus (Haaf) Indonesia
Alcidodes confusus Haaf, 1963: 153; Alonso-
Zarazaga and Lyal, 1999: 206; Lyal in Lyal and
Curran, 2000: 1808
93 connexus (Haaf) Indonesia
Alcidodes connexus Haaf, 1964a: 39; Alonso-
94 conradti (Faust) Equatorial Guinea
Alcides conradti Faust, 1895a: 236; Marshall,
95 contemptus (Voss) China
Alcidodes contemptus Voss, 1937: 226; Alonso-
96 convexus (Olivier) Tanzania, Kenya,
Alcides convexus Olivier, 1790: 507; 1807: 176; Uganda, South
Marshall, 1939: 570; Alonso-Zarazaga and Lyal, Africa, Madagascar,
1999:206 Cameroon, Gabon,
Alcides gibbus Fabricius, 1792: 431; 1801: 471; Zanzibar
Klima, 1934: 45; Marshall, 1939: 570
96a convexus var orienta/is Chevrolat, 1881b: 90
confluens Heller, 1918: 240 see leucospilus Erichson
97 coquereli (Fairmaire) Madagascar
Alcides coquereli Fairmaire, 1871: 48; Marshall,
Alcides lixoides Hustache, 1925a: 346; Klima,
1934: 46; Marshall, 1939: 570

coquereli var lixoidalis Klima, 1934: 46
corniculus Faust, 1894a: 250, 259 to
Cylindralcidodes: Pajni and Dhir, 1987:32
98 costipennis (Fairmare) Mayotte
Alcides costipennis Fairmare, 1871: 49; Marshall,
99 costulatus (Heller) Cambodia
Alcides costulatus Heller, 1922a: 22; Marshall,
100 crassirostris (Thomson) Equatorial Guinea,
Alcides crassirostris Thomson, 1858: 133; Gabon
1999:206
101 crassus (Pascoe) India, Philippines

Alcides crassus Pascoe, 1882: 450; Heller, 1918:
212, 224 (Sternuchopsis); Marshall, 1939: 570;
Pajni and Dhir, 1987:34; Alonso-Zarazaga and
Lyal, 1999: 206
Alcidodes hoplomachus Heller, 1918: 224; Lyal in
Lyal and Curran, 2000: 1788
102 crenatidens (Heller) Indonesia
Alcides crenatidens Heller, 1908b: 168; Marshall,
103 cribrithorax (Kolbe) Tanzania, Kenya,
Alcides cribrithorax Kolbe, 1898: 265; Marshall, Uganda
crinalifer Marshall, 1922: 393 to
Tuberculomecyslobus: Pajni and Dhir, 1987: 31
104 cultirostris (Thomson) Equatorial Guinea
Alcides cultirostris Thomson, 1858: 133; Marshall,
105 cultratus (Haaf) Indonesia
Alcidodes cultratus Haaf, 1964a: 29; Alonso-
cuprinus Heller, 1918: 241 see septemdecimnotatus
(Roelofs)
curialis Pascoe, 1883: 124 see transversus
(Walker)
106 curranae Lyal in Lyal and Curran, 2000: 1794 Indonesia
curtirostris Fairmaire, 1871: 50 see brevirostris
(Boheman)
107 curtus (Haaf) Indonesia
Alcidodes curtus Haaf, 1964a: 26; Alonso-Zarazaga
and Lyal, 1999: 206
108 *curvirostris Lyal in Lyal and Curran, 2000: 1833 Malaysia
109 cylindricus (Kolbe) Tanzania, Kenya,
Alcides cylindricus Kolbe, 1898: 263; Marshall, Uganda
cylindricus Hustache, 1920b: 10 see hustachei

(Heller)
cylindriformis Hustache, 1925a: 338,346 see
hustachei (Heller)
110 daedalus (Pascoe) Indonesia
Alcides daedalus Pascoe, 1882: 449; Marshall,
111 dealbatus (Boisduval) Indonesia, Figi,
Alcides dealbatus Boisduval, 1835: 297; Marshall, Papua New Guinea,
1939: 570; Alonso-Zarazaga and Lyal, 1999: 206 Indonesia
112 daitoanus (Kono) Thailand
Alcidodes daitoanus Kono, 1942: 163; Alonso-
113 decemvittatus (Aurivillius) India, Myanmar,
Alcides decemvittatus Aurivillius, 1892: 220; Cambodia

1999:206
decoratus Roelofs, 1893: 37 to Ornatalcides:
Heller, 1918: 215
114 decorsei (Hustache) Madagascar
Alcides decorsei Hustache, 1920b: 10; Marshall,
decretus Haaf, 1961c: 151 see sparsus (Boheman)
115 decursus (Pascoe) Indonesia
Alcides decursus Pascoe, 1885: 240; Marshall,
116 dejeani (Faust) Indonesia
Alcides dejeani Faust, 1896: 150; Marshall, 1939:
delicatulus Haaf 1961: 23 to Merus: Alonzo-
117 delta (Pascoe) Myanmar,
Alcides delta Pascoe, 1870b: 460; Faust, 1894a: Cambodia, Laos,
243, 258; Marshall, 1918a: 153, 154; Heller, 1933: Indonesia, Sri Lanka
99; Marshall, 1939: 570; Alonso-Zarazaga and
Lyal, 1999: 206
118 densus (Haaf) Indonesia
Alcidodes densus Haaf, 1963: 158; Alonso-
119 denticulatus (Kolbe) Kenya, Tanzania,
Alcides denticulatus Kolbe, 1898: 265; Marshall, Uganda
119a denticulatus sub sp vicinus Haaf, 1961c: 140
120 dentipes (Olivier) Equatorial Guinea,
Alcides affinis Fahraeus, 1871: 256; Marshall, South Africa,
1952: 268 Germany
Curculio dentipes Olivier, 1790: 504; Fabricius,
1792: 428 (Rhynchaenus); Herbst, 1795: 180
(Alcides); Marshall, 1952: 268 (Alcidodes);

121 dentirostris (Voss) China
Alcides dentirostris Voss, 1932a: 296; Marshall,
122 depressipennis (Hustache) Congo
Alcides depressipennis Hustache, 1924: 72;
1999:206
122a depressipennis sub sp amandus Haaf, 1961c: 102
deremius Kolbe, 1898: 266 to Merus: Alonso-
desertus Haaf, 1961a: 17 to Cylindralcidodes: Pajni
and Dhir, 1987: 32
123 detectus (Heller) fudonesia
Alcides detectus Heller, 1925: 231; Marshall, 1939:

Alcides deturbatus Heller, 1925: 231; Marshall,
1939: 570; Haaf, 1964a: 58
124 detentus (Pascoe) fudonesia
Alcides detentus Pascoe, 1885: 240; Marshall,
125 detractus (Heller) Papua New Guinea,
Alcides detractus Heller, 1925: 232; Marshall, fudonesia
deturbatus Heller, 1925: 231 see detectus (Heller)
126 diabolicus (Haaf) fudonesia
Alcidodes diabolicus Haaf, 1961b: 430; Alonso-
127 diadema (Heller) fudia
Alcides diadema Heller, 1908b: 158, 161; Marshall,
128 dictyodes (Heller) Philippines
Alcides dictyodes Heller, 1918: 213, 230; Marshall,
Ale ides tagalicus Schulze, 1918: 273; Klima, 1934:
53
129 didymus (Haaf) Malaysia
Alcidodes didymus Haaf, 1964a: 25; Alonso-
difficilis Heller, 1918: 220 to Cylindralcides:
Heller, 1918: 211
130 dignus (Haaf) Philippines
Alcidodes dignus Haaf, 1961e: 477; Alonso-
131 dipterocarpi (Marshall) fudia
Alcides dipterocarpi Marshall, 1921: 166; 1939:
132 disceedens (Pascoe) Singapore, fudonesia
Alcides disceedens Pascoe, 1870b: 461; Marshall,

133 discrepans (Faust) India
Alcides discrepans Faust, 1898: 309; Marshall,
134 distigma (Pascoe) Indonesia
Alcides distigma Pascoe, 1882: 448; Marshall,
135 distinchus (Haaf) India
Alcidodes distinchus Haaf, 1961a: 27; Alonso-
136 divergens (Pascoe) Singapore, Indonesia
Alcides divergens Pascoe, 1882: 446; Marshall,
137 diversus (Haaf) Africa
Alcidodes diversus Haaf, 1961c: 135; Alonso-

138 drescheri (Haaf) Indonesia
Alcidodes drescheri Haaf, 1964a: 50; Alonso-
139 duyagi (Schultze) Philippines
Alcides duyagi Schultze, 1934: 333; Marshall,
effertus Haaf, 1964a: 72 see chiliarchus sub sp
effertus
140 egregius (Haaf) Africa
Alcidodes egregius Haaf, 196lc: 115; Alonso-
141 elegans (Boisduval) Indonesia
Alcides elegans Boisduval, 1835: 421; Marshall,
142 elegans (Guerin-Meneville) Papua New Guinea
Alcides elegans Guerin-Meneville, 1838: 124; Indonesia, England
1999: 206
142a elegans var albolituratus Blanchard, 1853: 245
142b elegans var atrocretosus Fairmaire, 1883b: 37
142c elegans var blanchardi (Faust)
1952: 268
143 elongatus (Heller) Indonesia
Alcides elongatus Heller, 1929: 114; Marshall,
144 ephippiatus (Marshall) Sri Lanka
Alcidodes ephippiatus Marshall, 1918a: 153, 154;
145 eremitus (Haaf) Indonesia
Alcidodes eremitus Haaf, 1963: 99; Alonso-
erichsoni Heller, 1918: 240 see leucospilus
Erichson
146 erosus (Haaf) Sri Lanka
Alcidodes erosus Haaf, 1960: 651; Alonso-
erro Pascoe, 1871: 182 to Merus: Alonzo-Zarazaga
and Lyal, 1999: 205
147 erroneous (Thomson) Congo, Gabon
Alcides erroneous Thomson, 1858: 132; Marshall,
148 eruditus (Duvivier) Congo
Alcides eruditus Duvivier, 1892: 312; Marshall,
149 erythropterus (Chevrolat) Zanzibar, Kenya,
Alcides erythropterus Chevrolat, 1881b: 90; Tanzania, Uganda,
Marshall, 1952: 268 (Alcidodes); Voss, 1962b: 271; Congo

Alcides otiosus Hartmann, 1899: 24; Marshall,
1952:268
149a erythropterus sub sp paucifasciatus Voss, 1962b: 271 Congo
150 eugeniophilus (Lyal) Malaysia
Alcidodes eugeniophilus Lyal, 1996: 1702;
eupidus Haaf, 1961c: 58 see guttulatus (Thomson)
151 evidens (Heller) Philippines
Alcides evidens Heller, 1918: 214, 231; Marshall,
152 excavatus (Olivier) Madagascar,
Rhynchaenus excavatus Olivier, 1807: 199; Cameroon
Schoenherr, 1836: 625 (Alcides); Marshall, 1939:
153 excellens Haaf, 1961b: 423 Indonesia
154 exilis (Boheman) Cape Colony
Alcides exilis Boheman in Schoenherr, 1836: 613;
1999:206
155 exornatus (Chevrolat) Papua New Guinea
Alcides exornatus Chevrolat, 1880: 333; Marshall,
Alcides gestroi Pascoe, 1885: 242; Klima, 1934: 54
156 expansitarsis (Lyal) Cambodia, India,
Alcidodes expansitarsis Lyal, 1996: 1694; Alonso- Indonesia, Laos,
Zarazaga and Lyal, 1999: 206 Myanmar
157 expansus (Haaf) Indonesia
Alcidodes expansus Haaf, 1964a: 22; Alonso-
158 eypocrite (Heller) Indonesia
Alcides eypocrite Heller, 1929: 114; Marshall,
159 fabricii (Fabricius) India
Curculiofabricii Fabricius, 1792: 169; 1801:471

(Rhynchaenus); Boheman in Schoenherr, 1836: 623
(Alcides); Pajni and Dhir, 1987: 34; Alonso-
159a fabricii varfenestratus Olivier, 1807: 204 India
facetus Haaf, 1961c: 148 see obsoletus sub sp
facetus
160 fairmairei (Gemminger) Madagascar
Alcides fairmairei Gemminger, 1871: 123;
1999:206
Alcides olivaceus Fairmaire, 1871: 50; Klima,
1934:47
161 fallax (Boheman) Madagascar
Alcides fallax Boheman in Schoenherr, 1844: 62;

1999: 206
162 fallerianus (Hustache) Ethiopia
Alcidesfallerianus Hustache, 1936a: 353; Marshall,
falsus Haaf 1964b: 159 see barbicus
163 farinosus (Haaf) Africa
Alcidodesfarinosus Haaf, 1961c: 146; Alonso-
fasciatus Redtenbacher in Hugel, 1848: 548 to
Merus: Alonzo-Zaraza and Lyal, 1999: 205
164 fasciculatus (Boheman) Congo, Germany
Alcidesfasciculatus Boheman in Schoenherr, 1836:
Lyal, 1999: 206
165 fasciger (Faust) Madagascar
Alcidesfasciger Faust, 1889: 101; Marshall, 1939:
166 fastuosus (Pascoe) Indonesia
Alcidesfastuosus Pascoe, 1873: 182; Marshall,
feae Faust, 1894a: 245,258 to Neomecyslobus:
ferox Olivier, 1807: 189 see bubo (Fabricius)
167 ferrugineus (Cristofori and Jan) Germany(?)
Alcides ferrugineus Cristofori and Jan, 1832: 4;
1999: 206
168 ferrugineus (Hartmann) Ghana
Alcidesferrugineus Hartmann, 1900: 274;
1999: 206
fervidus Haaf, 1961a: 15 to Cylindralcidodes: Pajni
and Dhir, 1987: 32
169 figuratus (Schultze) Philippines

Alcides figuratus Schultze, 1919: 55 8; Marshall,
170 firmus (Haaf) Papua New Guinea
Alcidodesfirmus Haaf, 1963: 167; Alonso-
171 flanarantsoaensis (Hustache) Madagascar
Ale ides flanarantsoaensis Hustache, 1933: 84;
1999:206
flavofurcatus Voss, 1957: 107 see suspensus
(Pascoe)
172 flavoguttatus (Marshall) Kenya, Tanzania,
Alcidodesflavoguttatus Marshall, 1939: 570; Uganda

flavo-signatus Roelofs, 1875:151 see lixoides
(Reitter)
173 formicatus (Haaf) Sri Lanka
Alcidodesformicatus Haaf, 1961d: 55; Alonso-
174 fossor (Haaf) Malaysia
Alcidodes foss or Haaf, 1964a: 33; Alonso-Zarazaga
and Lyal, 1999: 206
175 franzi (Hoffmann) Tanzania
Alcidodes franzi Hoffmann, 1965: 405; Alonso-
176 frater (Hartmann) Ghana
Ale ides frater Hartmann, 1900: 276; Marshall,
fraudator Heller, 1924: 186 see circulifer Heller
177 frenatus (Faust) Myanmar
Alcidesfrenatus Faust, 1894a: 249, 259; Marshall,
Alcides clathratus Aurivillius, 1892: 219; Klima,
1934:54
178 freudei (Haaf) Papua New Guinea
Alcidodesfreudei Haaf, 1963: 159; Alonso-
179 frigidus (Haaf) Indonesia
Alcidodesfrigidus Haaf, 1961b: 389; Alonso-
frontalis Pascoe, 1871: 183 to Paipalephorus
mucoreus Jekel: Alonzo-Zarazaga and Lyal, 1999:
206
180 fugitus Lyal in Lyal and Curran, 2000: 1809 Indonesia
181 fulvocinctus (Marshall) Africa
Alcidodesfulvocinctus Marshall, 1958b: 715;
182 funestus (Faust) Myanmar, India
Alcidesfunestus Faust, 1894a: 247, 258; Marshall,

gallarius Pascoe, 1888: 412 see gallus (Pascoe)
183 gallus (Pascoe) Indonesia
Alcides gallus Pascoe, 1887: 354; Marshall, 1939:
Alcides gallarius Pascoe, 1888: 412; Haaf, 1963:
118
184 gmelinae (Marshall) India, Myanmar
Alcides gmelinae Marshall, 1922: 406; 1939: 570;
185 geminatus (Pascoe) Indonesia
Alcides geminatus Pascoe, 1887: 357; Marshall,
geminus Heller, 1908b: 159,161 see comparabilis

Bovie
186 geminus (Faust) Congo, Kenya,
Alcides geminus Faust, 1899b: 411; Marshall, Tanzania, Uganda,
1939: 570; Alonso-Zarazaga and Lyal, 1999: 206 Zanzibar
187 gemmatus (Harold) Angola
Alcides gemmatus Harold, 1879: 150,151;
1999:206
188 generosus (Haaf) Indonesia
Alcidodes generosus Haaf, 1964a: 20; Alonso-
189 geniculatus (Faust) Papua New Guinea
Alcides geniculatus Faust, 1897b: 182; Marshall,
geometricus Pascoe, 1885: 243 see angulus
(Fabricius)
gestroi Pascoe, 1885: 242 see exornatus
(Chevrolat)
190 gibbipennis (Haaf) Indonesia
Alcidodes gibbipennis Haaf, 1961b: 403; Alonso-
gibbus Fabricius, 1792: 431 see convexus (Olivier)
191 glabratus (Haaf) Philippines
Alcidodes glabratus Haaf, 1962: 146; Alonso-
192 glanvillei (Peringuey) Cape Colony, South
Alcides glanvillei Peringuey, 1885: 143; Marshall, Africa
Alcides mouffleti Faust, 1899b: 411; Marshall, 1906:
957
193 gonzoi Lyal in Lyal and Curran, 2000: 1813 Malaysia
194 gossypii (Hustache) Congo
Alcides gossypii Hustache, 1922: 150; Marshall,
195 granicollis (Fairmaire) Madagascar

Alcides granicollis Fairmaire, 1886: 79; Marshall,
graniger Haaf, 1961c: 81 see leucogrammus sub sp
graniger
196 granulipennis (Hustache) Gabon
Alcides granulipennis Hustache, 1932: 64;
Marshall, 1939: 570; Alonso-Zarazaga and Lya1,
1999:206
197 griseolineatus (Kolbe) Africa
Alcides griseolineatus Kolbe, 1883: 34; Marshall,
198 griseoniger (Fairmaire) Madagascar
Alcides griseoniger Fairmaire, 1902b: 238;

1999:206
gubatanus Schultze, 1922: 590 to Merus: Schultze,
1922: 590
199 guessfeldti (Kolbe) Congo, West Africa
Alcides aschanticus Faust, 1895a: 236; Marshall,
1952:268
Alcides guessfeldti Kolbe, 1883: 34; Marshall,
1939, 570; Alonso-Zarazaga and Lyal, 1999: 206
199a guessfeldti sub sp paucus Haaf, 1961c: 72
199b guessfeldti sub sp remotus Haaf, 1961c: 72
guttulatus Pascoe, 1883: 125 see pascoei Bovie
200 guttulatus (Thomson) Gabon
Alcides guttulatus Thomson, 1858: 134; Marshall,
200a guttulatus sub sp eupidus Haaf, 1961c: 58
200b guttulatus sub sp vividus Haaf, 1961c: 58
201 gymnasticus (Lyal) Malaysia, Myanmar
Alcidodes gymnasticus Lyal, 1996: 1696; Alonso-
202 habena (Heller) Philippines
Alcides habena Heller, 1918: 226; Marshall, 1939:
habena sub sp pangayensis Schultze, 1918: 275;
Haaf, 1961e: 478
Alcides pangayensis Schultze, 1918: 275; Marshall,
1939:570
Alcides taylori Schultze, 1918: 275; Klima, 1934:
54
habenatus Marshall, 1922: 395 to Neomecyslobus:
203 haddeni (Heller) Philippines
Alcides haddeni Heller, 1934: 296; Marshall, 1939:
204 haemopterus (Boheman) Ethiopia, Germany,

Alcides haemopterus Boheman in Schoenherr, Cape Colony, South
1836: 624; Marshall, 1939: 570; Alonso-Zarazaga Africa, Tanzania,
and Lyal, 1999: 206 Gabon
204a haemopterus var abyssinicus Hustache, 1932: 67
204b haemopterus var ater Aurivillius, 1910: 427
204c haemopterus sub sp lautus Haaf, 1961c: 91
204d haemopterus sub sp ornatus Haaf, 196lc: 89
204e haemopterus sub sp tectus Haaf, 1961c: 91
205 harmonicus (Haaf) India
Alcidodes harmonicus Haaf, 1961a: 47; Alonso-
206 haruspax (Faust) Myanmar
Alcides haruspax Faust, 1894a: 243,258; Marshall,

207 heilipoides Pascoe Tanzania, Uganda,
Alcides heilipoides Pascoe, 1870d: 211; Marshall, Kenya
hemicyclus Marshall, 1922: 400 see pyrifer sub sp
hemicyclus
208 herteli Haaf, 1963: 140 Papua New Guinea
209 hexamitosus (Heller) Indonesia
Alcides hexamitosus Heller, 1908b: 155; Marshall,
Alcides centroguttatus Marshall, 1926b: 357; Haaf,
1961b:428
210 hispidus (Haaf) Africa
Alcidodes hispidus Haaf, 1961c: 153; Alonso-
211 homeyeri (Harold) Congo
Alcides homeyeri Harold, 1879: 150, 151; Marshall,
212 hopeae (Marshall) Indonesia
Alcides hopeae Marshall, 1926a: 347; 1939: 570;
213 hoplomachus Lyal in Lyal and Curran, 2000: 1788 Philippines
hoplomachus Heller, 1918: 224 see crassus
214 hospitus (Haaf) Myanmar
Alcidodes hospitus Haaf, 1964b: 192; Alonso-
215 hovanus (Hustache) Madagascar
Alcides hovanus Hustache, 1920b: 10; Marshall,
216 humeralis (Heller) India, Malaysia
Alcidodes humeralis Heller, 1940: 93; Alonso-
humerosus Ancey, 1881a: 461 see olivaceus
(Gerstacker)
217 humerosus (Harold) Kenya, Tanzania,
Alcides humerosus Harold, 1880: 265; Marshall, Uganda

218 humatus (Haaf) Philippines
Alcidodes humatus Haaf, 1961e: 482; Alonso-
219 humilis (Hustache) Congo
Alcides humilis Hustache, 1932: 67; Marshall,
220 hustachei (Heller) India, Malaysia,
Alcides hustachei Heller, 1933: 98; Marshall, 1939: Kenya, Tanzania,
570; Alonso-Zarazaga and Lyal, 1999: 206 Uganda, Madagascar
Alcides cylindricus Hustache, 1920b: 10; Heller,
1933:98
Alcides cylindriformis Hustache, 1925a: 338,346;
Heller, 1933: 98
hypocrites Boheman in Schoenherr, 1844: 47 to
Ornatalcides:
221 idenburgi (Heller) Papua New Guinea
Alcidodes idenburgi Heller, 1942: 212; Alonso-
ignoratus Heller, 1918: 238 to Ornatalcides:
Heller, 1918: 214
222 imbellis (Thomson) Gabon
Alcides imbellis Thomson, 1858: 134; Marshall,
223 imitator (Haaf) Indonesia
Alcidodes imitator Haaf, 1964a: 66; Alonso-
224 immixtus (Hustache) Congo
Alcides immixtus Hustache, 1934: 348; Marshall,
225 immutatus Haaf, 196lb: 409 Indonesia
226 impressus (Heller) India
Alcides impressus Heller, 1908b: 160, 161;
1999:206
improvidus Faust, 1894a: 241, 257 to Ornatalcides:
incomptus Marshall, 1915: 516 see tenuistria
(Heller)
227 indigaceus (Pascoe) Indonesia
Alcides indigaceus Pascoe, 1882: 447; Marshall,
Alcides parilis Pascoe, 1882: 448; Marshall, 1918a:
158
228 indubitus (Haaf) Indonesia
Alcidodes indubitus Haaf, 1961b: 425; Alonso-
229 inducens (Heller) Indonesia

Alcides inducens Heller, 1925: 230, 231; Marshall,
230 inermis (Heller) Sri Lanka
Alcides inermis Heller, 1901: 345; Marshall, 1939:
231 inops (Haaf) Tanzania
Alcidodes inops Haaf, 1964c: 291; Alonso-
232 inquietus (Haaf) India
Alcidodes inquietus Haaf, 1961a: 38; Alonso-
233 insularis (Schultze) Philippines
Alcides insularis Schultze, 1918: 271; Marshall,

234 intermedius (Ancey) Kenya, Tanzania,
Alcides intermedius Ancey, 1881a: 485; Marshall, Uganda
interpostitus Heller, 1924: 185 see aeratus sub sp
interpostitus
235 interruptolineatus (Faust) Sierra Leone, West
Alcides interruptolineatus Faust, 1894c: 554; Africa
1999:206
236 interruptus (Boheman) Gabon, Congo,
Alcides interruptus Boheman in Schoenherr, 1836: Central African
622; Marshall, 1939: 570; Alonso-Zarazaga and Republic, Chad,
Lyal, 1999: 206 Senegal, Germany
236a interruptus var. bilineellus Heller, 1904: 186
intrusomimus Heller, 1922a: 19 see scenicus
237 intrusus (Pascoe) Indonesia
Alcides intrusus Pascoe, 1885: 240; Heller, 1918:
Lyal, 1999: 206
238 janetae (Lyal) Indonesia
Alcidodesjanetae Lyal, 1996: 1700; Alonso-
javanodelta Marshall, 1918a: 155 see perturbatus
(Faust)
josephus Duvivier, 1892: 312 to Merus: Alonzo-
jueundus Haaf, 1961c: 124 see bivinculatus sub sp
jueundus
239 jugalis (Marshall) Malaysia
Alcidodes jugalis Marshall, 1922: 394, 401;
240 juglans (Chao and Chen) China
Alcidodes juglans Chao and Chen, 1980: 168;
kalinganus Schultze, 1923: 604 to Ornatalcides:

Heller, 1918: 214
karelinii Boheman in Schoenherr, 1844: 65 see
chaudoiri (Chevrolat)
241 karnyi (Heller) Philippines
Alcides karnyi Heller, 1933: 102,103; Marshall,
kenyae Hustache, 1929: 508 see kilimanus
Aurivillius
kuatunensis Voss, 1957: 104 see nipponicus (Kono)
242 kignonseranus (Hustache) Congo
Alcides kignonseranus Hustache, 1937: 221;
1999:206
kilimanus Aurivillius, 1910: 427 to Merus:

Alonzo-Zarazaga and Lyal, 1999: 205
243 kirschi (Pascoe) Labuan, Malaysia
Alcides kirschi Pascoe, 1882: 449; Marshall, 1939:
244 konoi (Voss) Taiwan
Alcides konoi Voss, 1932b: 296; Marshall, 1939:
245 korotyaevi (Egorov) Russia
Alcidodes korotyaevi Egorov, 1977: 39; Alonso-
246 kuvanguensis (Hustache) Angola
Alcides kuvanguensis Hustache,1938b: 74;
1999:206
lacumsus Boheman in Schoenherr, 1844: 47 see
angulus (Fabricius)
247 lacunosus (Boheman) Indonesia,
Alcides lacunosus Boheman in Schoenherr, 1844: Cambodia, Laos
Lyal, 1999: 206
248 laetus (Heller) Indonesia
Alcides laetus Heller, 1908b: 165; 1918: 227;
1999:206
249 lagunensis (Schultze) Philippines
Alcides lagunensis Schultze, 1934: 335; Marshall,
250 lameerei (Faust) Congo, Benguela (?)
Alcides lameerei Faust, 1899b: 411; Marshall,
Alcides rubrirostris Pape, 1907: 326; Marshall,
1918a: 158
lamprocolius Heller, 1924: 184 to Merus: Heller,
1924: 184
2010 Andrew & Ramamurthy: Checklist, Mo1ytinae 301
251 lamprotrox (Heller) Philippines

Alcides lamprotrox Heller, 1924: 185; Marshall,
252 laniger (Sparrm) Congo, South
Curculio laniger Sparrm, 1785: 43; Boheman in Africa, Cape Colony
Schoenherr, 1836: 611 (Alcides); Marshall, 1939:
253 laqueifer (Heller) Indonesia
Alcides laqueifer Heller, 1925: 232; Marshall,
254 larinoides (Fairmaire) Madagascar
Alcides larinoides Fairmaire in Fleutiaux, 1899: 25;
Marshall, 1939: 570; A1onso-Zarazaga and Lya1,
1999:206
Alcides tomentosus Hustache, 1925a: 339,345;

Klima, 1934: 48
254a larinoides var modestus Klima, 1934: 48 Madagascar
255 lascivus (Haaf) Indonesia
Alcidodes lascivus Haaf, 1963: 144; Alonso-
256 latefasciatus (Chevrolat) Zanzibar
Alcides latefasciatus Chevrolat, 1881b: 89;
1999:206
lautus Haaf, 1961c: 91 see haemopterus sub sp
lautus
257 lemniscatus (Haaf) Papua New Guinea
Alcidodes lemniscatus Haaf, 1964c: 293; Alonso-
258 lepidus (Haaf) Indonesia
Alcidodes lepidus Haaf, 1963: 122; Alonso-
259 leechi (Haaf) Indonesia
Alcidodes leechi Haaf, 1962: 147; Alonso-Zarazaga
and Lyal, 1999: 206
260 leeuweni (Heller) Indonesia
Alcides leeuweni Heller, 1911: 312, 314; Marshall,
leopardus Olivier, 1807: 190, 296 to Ornatalcides:
261 leucogrammus (Erichson) Congo, Angola,
Alcides leucogrammus Erichson, 1842: 260; Uganda
1999: 206
261a leucogrammus sub sp graniger Haaf, 1961c: 81
leucospilus Erichson, 1834: 264 to Ornatalcides
Heller, 1918: 215
262 lewisi (Pascoe) Sri Lanka
Alcides lewisi Pascoe, 1883: 124; Marshall, 1939:
302 Oriental fusects Vol. 44

leytensis Heller, 1918: 239 see leucospilus
lichen Faust, 1894a: 245, 258 see saundersi
(Pascoe)
263 liciatus (Haaf) Philippines
Alcidodes liciatus Haaf, 1962: 149; Alonso-
264 ligatus (Pascoe) Indonesia
Alcides ligatus Pascoe, 1870b: 461; Marshall,
265 lineatus (Klug) Madagascar
Alcides lineatus Klug, 1833: 91; Marshall, 1939:
266 lineellus (Faust) Congo
Alcides lineellus Faust, 1899b: 415; Marshall,

267 lituratus (Hustache) Congo, Tanzania
Alcides lituratus Hustache, 1932: 64; Marshall,
268 lividinasus (Heller) Philippines
Alcides lividinasus Heller, 1918: 225; Marshall,
269 lixiformis (Marshall) Zimbabwe, Congo
Alcides lixiformis Marshall, 1920: 391; 1939: 570;
Alcides maynei Hustache, 1924: 70; Klima, 1934:
48
lixoides Reitter, 1905: 248 to Ornatalcides: Pajni
and Dhir, 1987: 30
lixoides Hustache, 1925a: 346 see coquereli
(Fairmaire)
lobipennis Fairmaire, 1902a: 365 see malgassicus
Bovie
270 lobipennis (Fairmaire) China
Alcides lobipennis Fairmaire, 1900: 634; Marshall,
271 longirostris (Heller) Philippines
Alcides longirostris Heller, 1918: 221; Haaf, 1962:
loratus Marshall, 1922: 394, 397 to
Neomecyslobus: Pajni and Dhir, 1987: 31
ludificator Faust, 1894a: 254, 260 to
Cylindralcidodes: Pajni and Dhir, 1987: 32
272 lugubris (Haaf) Sri Lanka
Alcidodes lugubris Haaf, 1961d: 56; Alonso-
luxuriosus Faust, 1894a: 245, 258 see saundersi
(Pascoe)
273 luzonensis (Schultze) Philippines
Alcides luzonensis Schultze, 1918: 273; Marshall,

274 macellus (Haaf) Philippines
Alcidodes macellus Haaf, 1962: 148; Alonso-
275 maculosus (Fairmaire) Madagascar
Alcides maculosus Fairmaire, 1902a: 366;
1999: 206
276 magicus (Pascoe) Cambodia
Alcides magicus Pascoe, 1870b: 460; Marshall,
277 magister (Pascoe) Papua New Guinea,
Alcides magister Pascoe, 1873: 181; Marshall, Indonesia

278 magnificus (Haaf) Sri Lanka
Alcidodes magnificus Haaf, 1960: 657; Alonso-
major Haaf, 1964b: 167 to Cylindralcidodes: Pajni
and Dhir, 1987: 32
279 malgassicus (Bovie) Madagascar, China
Alcides malgassicus Bovie, 1907: 67; Marshall,
Ale ides lobipennis Fairmaire, 1902a: 365; Klima,
1934:49
mali Marshall, 1919: 276 to Cylindralcidodes:
280 mamilla (Heller) Indonesia
Alcides mamilla Heller, 1937: 60; Marshall, 1939:
mangyanicus Schultze, 1922: 590 to Merus:
Schultze, 1922: 590
281 maynei (Hustache) Congo
Alcides maynei Hustache, 1924: 43; Marshall,
mellitus Faust, 1898: 308 to Neomecyslobus: Pajni
and Dhir, 1987: 31
282 merrilli (Schultze) Philippines
Alcides merrilli Schultze, 1918: 272; Marshall,
283 meruanus (Aurivillius) Kenya
Alcides meruanus Aurivillius, 1910: 428; Marshall,
284 micranthiphilus Lyal in Lyal and Curran, 2000: 1836 Indonesia
micronychus Pascoe, 1872: 483 to Brahmalcidodes:
285 mimus (Heller) Philippines
Alcides mimus Heller, 1934: 296; Marshall, 1939:
mindanaoensis Schultze, 1918: 271 see leucospilus

286 mindorensis (Schultze) Philippines
Alcides mindorensis Schultze, 1918: 272; Marshall,
287 minutepustulatus (Hustache) Africa
Alcides minutepustulatus Hustache, 1936b: 13;
1999:206
288 mirandus (Haaf) Africa
Alcidodes mirandus Haaf, 1961c: 127; Alonso-
miser Heller, 1918: 218 to Cylindralcides: Heller,
1918:218
289 molitor (Heller) India
Alcides molitor Heller, 1908b: 161; Marshall, 1939:

290 monardi (Hustache) Angola
Alcides monardi Hustache, 1935: 165; Marshall,
291 monilifer (Pascoe) Sri Lanka
Alcides monilifer Pascoe, 1882: 450; Marshall,
292 monstratus (Haaf) Thailand
Alcidodes monstratus Haaf, 1964b: 189; Alonso-
montanus Haaf, 1964b: 160 to Ornatalcides: Pajni
and Dhir, 1987: 30
293 montivagus (Heller) Philippines
Alcides montivagus Heller, 1933: 101, 103;
1999:206
294 mario (Heller) India
Alcides mario Heller, 1908b: 171; Marshall, 1939:
295 morosus (Haaf) Malaysia
Alcidodes morosus Haaf, 1964a: 65; Alonso-
296 morulus (Haaf) Africa
Alcidodes morulus Haaf, 1961c: 102; Alonso-
mouffleti Faust, 1899b: 411 see glanvillei
(Peringuey)
297 muiri (Marshall) Indonesia
Alcidodes muiri Marshall, 1918a: 154; 1939: 570;
298 murranus (Marshall) Ethiopia, Sudan
Alcidodes murranus Marshall, 1942: 238; Alonso-
299 musculus (Fairrnaire) Madagascar
Alcides musculus Fairmaire, 1902a: 367; Marshall,

mustela Pascoe, 1882: 446 to Cylindralcides:
Heller, 1918: 211, 219
300 muticus (Haaf) Africa
Alcidodes muticus Haaf, 1961c: 157; Alonso-
mysticus Faust, 1894a: 246, 258 to
Cylindralcidodes: Pajni and Dhir, 1987: 32
301 nanus (Haaf) Indonesia
Alcidodes nanus Haaf, 1964a: 34; Alonso-Zarazaga
and Lyal, 1999: 206
302 naso (Pascoe) Indonesia
Alcides naso Pascoe, 1885: 241; Marshall, 1939:

303 natalensis (Haaf) Africa
Alcidodes natalensis Haaf, 1961c: 82; Alonso-
304 neavei (Marshall) Mosambique
Alcidodes neavei Marshall, 1935: 1; Alonso-
necopinus Faust, 1899b: 415 see obsoletus
(Gerstacker)
305 neglectus (Cristofori and Jan) Germany(?)
Alcides neglectus Cristofori and Jan, 1832: 4;
1999: 206
306 negrosensis (Schultze) Philippines
Alcides negrosensis Schultze, 1934: 333; Marshall,
307 nepalensis (Faust) Nepal
Alcides nepalensis Faust, 1894a: 253, 260; Marshall,
308 nero (Fabricius) Indonesia
Alcides nero Fabricius, 1801: 466; Marshall, 1939:
Alcides patruelis Faust, 1892b: 209; Haaf, 1961b:
388
nicopinus Faust, 1899b: 415 see obsoletus
(Gerstacker)
309 niger (Hustache) Congo
Alcides niger Hustache, 1924: 71; Marshall, 1939:
310 nigricollis (Haaf) Vietnam
Alcidodes nigricollis Haaf, 1964b: 175; Alonso-
311 nigritus (Haaf) Indonesia
Alcidodes nigritus Haaf, 1963: 125; Alonso-
Zarazaga and Lya1, 1999: 206
312 nigrofasciatus (Kono) Japan

Alcides nigrofasciatus Kono, 1928: 175; Marshall,
313 nigrovinculatus (Heller) India, Malaysia
Alcidodes nigrovinculatus Heller, 1940: 92;
314 nipponicus (Kono) Japan
Alcides nipponicus Kono, 1930:138, 140; Marshall,
Mecyslobus kuatunensis Voss, 1957: 104; Haaf,
1964b: 169
315 nitidus (Pascoe) Indonesia
Alcides nitidus Pascoe, 1887: 356; Marshall, 1939:
Alcides analogus Heller, 1918: 244; Haaf, 1963:

161
316 nivosus (Fairmaire) Madagascar
Alcides nivosus Fairmaire, 1899: 542; Marshall,
nocens Heller, 1908b:162 to Ornatalcides: Pajni
and Dhir, 1987: 33
317 notabilis (Haaf) Srilank:a
Alcidodes notabilis Haaf, 1961d: 58; Alonso-
318 notatus (Blanchard) Indonesia
Alcides notatus Blanchard in Hombron and
319 notatus (Klug) Madagascar
Alcides notatus Klug, 1833: 91; Marshall, 1939:
320 novellas (Haaf) Indonesia
Alcidodes novellas Haaf, 1963: 157; Alonso-
321 nubilus (Marshall) Indonesia
Alcidodes nubilus Marshall, 1955: 278; Alonso-
322 nudiusculus (Haaf) Indonesia
Alcidodes nudiusculus Haaf, 1962: 144; Alonso-
323 nyamukubianus (Hustache) Congo
Alcides nyamukubianus Hustache, 1934: 350;
1999:206
324 oberthurii (Pascoe) India
Alcides oberthurii Pascoe, 1887: 357; Marshall,
obesus Faust, 1894a: 248, 259 to Cylindralcidodes:
obfuscatus Fabricius, 1775: 138 to Merus: Alonzo-

325 obliquatus (Thomson) Gabon
Alcides obliquatus Thomson, 1858: 131; Marshall,
326 obliquus (Walker) Sri Lanka
Alcides obliquus Walker, 1859: 264; Marshall,
327 oblongulus (Fairmaire) Madagascar
Alcides oblongulus Fairmaire, 1902a: 368;
1999:206
obsidianus Heller, 1918: 242 to Merus: Heller,
1918:242
328 obsoletus (Gerstacker) Kenya, Congo

Alcides obsoletus Gerstacker, 1871: 74; Marshall,
Alcides necopinus Faust, 1899b: 415; Marshall,
1952: 268
328a obsoletus sub spfacetus Haaf, 1961c: 148
329 obsoletus (Klug) Madagascar
Alcides obsoletus Klug, 1833: 91; Marshall, 1939:
330 obtusus (Pascoe) Indonesia
Alcides obtusus Pascoe, 1888: 412; Marshall, 1939:
ocellatus Roelofs, 1893: 35 see semperi (Pascoe)
331 ochraceus (Haaf) Papua New Guinea
Alcidodes ochraceus Haaf, 1963: 155; Alonso-
332 octoguttatus (Redtenbacher) India
Alcides octoguttatus Redtenbacher in Hugel, 1848:
Lyal, 1999: 206
olivaceus Fairmaire, 1871: 50 seefairmairii
Gemminger
333 olivaceus (Gerstacker) Chad, Tanzania,
Alcides anceyi Bovie, 1908b: 43; Klima, 1934: 49 Congo, Gabon,
Alcides biai Duvivier, 1890: 24; Klima, 1934: 49 Zanzibar,
Alcides humerosus Ancey, 1881a: 461; Klima, Mosambique,
1934: 49 Alcides olivaceus Gerstacker, 1855: 85; Germany
1999:206
Alcides wahlbergi Chevrolat, 1881b: 90; Klima,
1934:49
olivieri Bovie, 1908b: 43 see pectoralis
334 olsufievi (Hustache) Madagascar
Alcides olsufievi Hustache, 1938b: 26; Marshall,
335 omissus (Haaf) Philippines

Alcidodes omissus Haaf, 1963: 160; Alonso-
opacimaculatus Heller, 1918: 242 see obsidianus
Heller
336 opacus (Haaf) Indonesia
Alcidodes opacus Haaf, 1963: 126; Alonso-
337 opulentus (Haaf) Indonesia
Alcidodes opulentus Haaf, 1961b: 419; Alonso-
orientalis Chevrolat, 1881b: 90 see convexus
(Olivier)
ornatus Haaf, 1961c: 89 see haemopterus sub sp
ornatus
338 ostentatus (Haaf) Indonesia
Alcidodes ostentatus Haaf, 1964a: 72; Alonso-
339 otiosus (Hartmann) Chad, Tanzania,
Alcides otiosus Hartmann, 1899: 24; Marshall, Congo, Gabon
340 oxyomus (Fairmaire) Madagascar
Alcides oxyomus Fairmaire, 1902a: 365; Marshall,
341 paetus (Haaf) India
Alcidodes paetus Haaf, 1961: 29; Alonso-Zarazaga
and Lyal, 1999: 206 panayensis Schultze, 1918:
275 see taylori Schultze
342 paradictodes (Haaf) Philippines
Alcidodes paradictodes Haaf, 1961e: 498; Alonso-
343 parcus (Faust) Equatorial Guinea
Alcides parcus Faust, 1895a: 235; Marshall, 1939:
344 parentheticus (Marshall) Papua New Guinea
Alcidodes parentheticus Marshall, 1915: 515; 1939:
parilis Pascoe, 1882: 448 see indigaceus (Pascoe)
345 parnassius (Haaf) Indonesia
Alcidodes parnassius Haaf, 1964a: 67; Alonso-
346 pascoei (Bovie) Sri Lanka
Alcides pascoei Bovie, 1908b: 43; Marshall, 1939:
Alcides guttulatus Pascoe, 1883: 125; Klima, 1934:
57
patruelis Faust, 1892b: 209 see nero (Fabricius)
paucifasciatus Voss, 1962b: 271 see erythropterus
sub sp paucifasciatus
paucus Haaf, 1961c: 72 see guessfeldti sub sp

paucus
347 pauper (Boheman) India, Myanmar
Alcides pauper Boheman in Schoenherr, 1844: 61;
1999:206
348 pauxillus (Haaf) Philippines
Alcidodes pauxillus Haaf, 1962: 154; Alonso-
pavidus Faust, 1894a: 251, 259; Haaf, 1961b: 434
see obfuscatus (Fabricius)
349 paykulli (Boheman) Sierra Leone
Alcides paykulli Boheman in Schoenherr 1836:
Lyal, 1999: 206

350 pectoralis (Boheman) Myanmar, Thailand,
Alcides pectoralis Boheman in Schoenherr, 1836: Indonesia,
618; Marshall, 1939: 570; Alonso-Zarazaga and Philippines
Lyal, 1999: 206
Alcides olivieri Bovie, 1908b: 43; Klima, 1934: 57
Rhynchaenus sulcatulus Olivier, 1807: 203; Klima,
1934: 57
351 pentastictus (Ancey) Australia
Alcides pentastictus Ancey, 1881a: 372; Marshall,
352 perditor (Haaf) Indonesia
Alcidodes perditor Haaf, 1963: 137; Alonso-
353 peregrinus (Haat) Indonesia
Alcidodes peregrinus Haaf, 1961b: 424; Alonso-
354 perinetensis (Hustache) Madagascar
Alcides perinetensis Hustache, 1938a: 27; Marshall,
persegnis Heller, 1918: 222 to Cylindralcides:
Heller, 1918: 222
355 personatus (Haat) Indonesia
Alcidodes personatus Haaf, 1964a: 43; Alonso-
356 perturbatus (Faust) Indonesia
Alcidesjavanodelta Marshall, 1918a: 155; Klima,
1934:57
Alcides perturbatus Faust, 1896: 149; Marshall,
357 pervicax (Faust) Papua New Guinea
Alcides pervicax Faust, 1892b: 207; Marshall,
358 piceus (Roelofs) Japan
Alcides piceus Roelofs, 1875: 152; Marshall, 1939:

359 picturatus (Fairmaire) Madagascar
Alcides picturatus Fairmaire, 1897: 174; Marshall,
360 pictus (Fabricius) India
Alcides pictus Fabricius, 1798: 168; Marshall,
361 pindicus (Haaf) Africa
Alcidodes pindicus Haaf, 1961c: 59; Alonso-
pius Heller, 1918: 219 to Cylindralcides: Heller,
1918:219
362 plagiatus (Schultze) Philippines
Alcides plagiatus Schultze, 1919: 559; Marshall,

363 platysomus (Haaf) Papua New Guinea
Alcidodes platysomus Haaf, 1963: 132; Alonso-
364 plurilineatus (Hustache) Congo
Alcides plurilineatus Hustache, 1934: 356;
1999:206
365 porosus (Faust) Myanmar
Alcides porosus Faust, 1894a: 255, 260; Marshall,
porrectirostris Marshall, 1913: 188 to
366 posticus (Haaf) Indonesia
Alcidodes posticus Haaf, 1961b: 390; Alonso-
367 praeustus (Guerin-Meneville) Indonesia
Alcides praeustus Guerin-Meneville, 1833: 36;
1999:206
368 praeustus (Guerin-Meneville) Papua New Guinea,
Alcides praeustus Guerin-Meneville, 1838: 123; Indonesia
1999:206
369 praevius (Haaf) Papua New Guinea
Alcidodes praevius Haaf, 1963: 156; Alonso-
370 pretiosus (Haaf) Indonesia
Alcidodes pretiosus Haaf, 1963: 162; Alonso-
profluens Pascoe, 1885: 239 see australis
(Boisduval)
371 prolixus (Haaf) Indonesia
Alcidodes prolixus Haaf, 1963: 124; Alonso-
372 prominens (Hustache) Madagascar

Alcides prominens Hustache, 1933: 81; Marshall,
373 proximus (Faust) Equatorial Guinea,
Alcides proximus Faust, 1894c: 552; Marshall, Congo
374 pseudomellitus (Heller) Cambodia
Alcides pseudomellitus Heller, 1922a: 22; Marshall,
375 pubicollis (Fairmaire) Madagascar
Alcides pubicollis Fairmaire, 1902a: 365; Marshall,
376 pulchellus (Haaf) Malaysia, Indonesia
Alcidodes pulchellus Haaf, 1964a: 31; Alonso-

376a pulchellus sub sp sejugatus Haaf, 1964a: 33
377 pullus (Haaf) Indonesia
Alcidodes pullus Haaf, 1961b: 393; Alonso-
378 pumilus (Haaf) Philippines
Alcidodes pumilus Haaf, 1962: 151; Alonso-
379 puncticollis (Hustache) Africa
Alcides puncticollis Hustache, 1935: 165; Marshall,
380 puntaticollis (Voss) Namibia
Alcidodes puntaticollis Voss, 1960b: 5; Alonso-
381 pusillus (Pascoe) Papua New Guinea
Alcides pusillus Pascoe, 1885: 243; Marshall, 1939:
382 pustulatus (Heller) Indonesia
Alcides pustulatus Heller, 1929: 113-115; Marshall,
383 putus (Haaf) Indonesia
Alcidodes putus Haaf, 1964a: 70; Alonso-Zarazaga
and Lyal, 1999: 206
384 pyrifer (Marshall) Indonesia
Alcidodes pyrifer Marshall, 1922: 394, 399;
384a pyrifer sub sp hemicyclus Marshall, 1922: 400 Indonesia
385 quindecimspilotus (Heller) Philippines
Alcides quindecimspilotus Heller, 1923: 9;
1999:206
386 quinquesignatus (Aurivillius) Kenya
Alcides quinquesignatus Aurivillius, 1912: 367;
1999:206
387 ramezei Lyal in Lyal and Curran, 2000: 1797 Thailand

388 ramosus (Haa:f) Papua New Guinea
Alcidodes ramosus Haaf, 1963: 132; Alonso-
Zarazaga and Lya1, 1999: 206
raptorius Boheman in Schoenherr, 1836: 610 see
obfuscatus (Fabricius)
rectus Haaf, 1963: 127 see australis sub sp rectus
reductus Faust, 1892b: 208 see brevicollis (Pascoe)
389 relictus (Haaf) Africa
Alcidodes relictus Haaf, 1961a: 43; Alonso-
remotus Haaf, 1961c: 72 see guessfeldti sub sp
remotus
390 reticulatus (Schoenherr) Philippines
Alcides reticulatus Schoenherr, 1844: 46; Marshall,

391 rhomboidalis (Faust) Congo
Alcides rhomboidalis Faust, 1894c: 553; Marshall,
1939: 570; Alonso-Zarazaga and Lya1, 1999: 206
392 richteri (Faust) Papua New Guinea
Alcides richteri Faust, 1892b: 207; Marshall, 1939:
570; A1onso-Zarazaga and Lya1, 1999: 206
robustus Hustache, 1933: 83 see obesus (Faust)
roelofsi Lewis, 1879: 23 see waltoni
393 rostratus (Pascoe) Papua New Guinea
Alcides rostratus Pascoe, 1885: 241; Marshall,
394 rotundulus (Haaf) Indonesia
Alcidodes rotundulus Haaf, 1961b: 402; Alonso-
395 ruandanus (Hustache) Congo
Alcides ruandanus Hustache, 1934: 351; Marshall,
396 rubiginosus (Hustache) Madagascar
Alcides rubiginosus Hustache, 1920b: 10; Marshall,
397 rubripennis (Ancey) Tanzania
Alcides rubripennis Ancey, 1881a: 461; Marshall,
rubrirostris Marshall, 1918a: 158 see lameerei
(Faust)
398 rufipennis (Montrouzier) Samoa
Alcides rufipennis Montrouzier, 1855: 49;
1999: 206
399 rufus (Boheman) Cape Colony,
Alcides rufus Boheman in Schoenherr, 1836: 615; Germany
1999: 206
400 ruptus (Pascoe) Sri Lanka

Alcides ruptus Pascoe, 1883: 124; Marshall, 1939:
rutilans Roelofs, 1893: 38 to Ornatalcides: Heller,
1918:215
401 ryoichii (Kono) Thailand
Alcides ryoichii Kono, 1942: 162; Marshall, 1939:
402 sagonai (Hustache) Congo
Alcides sagonai Hustache, 1924: 71; Marshall,
403 saitoi (Kono) Korea
Alcides saitoi Kono, 1937a: 6; Marshall, 1939: 570;
404 salebrosus (Haat) Indonesia

Alcidodes salebrosus Haaf, 1961b: 408; Alonso-
405 sanctus (Haat) Africa
Alcidodes sanctus Haaf, 1961c: 108; Alonso-
satelles Faust, 1894a: 240, 258 to Ornatalcides:
saturnus Haaf, 1961a: 20 to Neomecyslobus: Pajni
and Dhir, 1987: 31
saundersi Pascoe, 1870b: 459 to Cylindralcidodes:
406 sauteri (Heller) Taiwan, Cambodia,
Alcides sauteri Heller, 1922a: 19; Marshall, 1939: Vietnam
scenicus Faust, 1894a: 256 to Ornatalcides: Pajni
and Dhir, 1987: 31
407 schereri (Haat) Malaysia,Indonesia
Alcidodes schereri Haaf, 1964b: 210; Alonso-
408 schuetzei (Schultze) Philippines
Alcides schuetzei Schultze, 1918: 274; Marshall,
409 schultzei (Klima) Philippines
Alcides schultzei Klima, 1934: 59; Marshall, 1939:
Alcides arcuatus Schultze, 1919: 559; Klima, 1934:
59
410 scitus (Haat) Indonesia
Alcidodes scitus Haaf, 1963: 147; Alonso-Zarazaga
and Lyal, 1999: 206
411 sedi (Marshall) South Africa,
Alcidodes sedi Marshall, 1938: 194; Alonso- Australia
412 sedulus (Haat) Indonesia
Alcidodes sedulus Haaf, 1964a: 61; Alonso-

413 segnis (Faust) Papua New Guinea,
Alcides segnis Faust, 1897b: 180; Marshall, 1939: Indonesia
segnis var. ternatensis Faust see ternatensis
sejugatus Haaf, 1964a: 33 see pulchellus sub sp
sejugatus
414 sellatus (Faust) Indonesia,
Alcides sellatus Faust, 1892b: 208; Marshall, 1939: Philippines
415 semiochraceus (Faust) Papua New Guinea
Alcidodes semiochraceus Faust, 1897b: 179;
1999: 206
416 semiroseus (Hustache) Madagascar
Alcidodes semiroseus Hustache, 1956: 119;
semperi Pascoe, 1870b: 462 to Merus: Heller,
1918:216,242
417 senex (Sahlberg) Sierra Leone, Congo
Alcides senex Sahlberg, 1823: 47; Marshall, 1939:
418 senex (Schoenherr) Congo
Alcides senex Schoenherr, 1825: 581; Marshall,
separandus Haaf, 1961a: 16 to Cylindralcidodes:
419 septemdecimnotatus (Roelofs) Philippines
Alcides cuprinus Heller, 1918: 241; Haaf, 1961e:
490
Alcides septemdecimnotatus Roelofs, 1893: 36;
1999:206
Alcides septemdecimprolatus Bertkau, 1894: 221;
Klima, 1934: 59
septemdecimprolatus Bertkau, 1894: 221 see
septemdecimnotatus (Roelofs)
420 septemmaculatus (Faust) Papua New Guinea
Alcides septemmaculatus Faust, 1897b: 181;
1999:206
septentrionalis Heller, 1918: 243 to Ornatalcides:
Heller, 1918: 243
421 serius (Haaf) Indonesia
Alcidodes serius Haaf, 1961b: 391; Alonso-
serotinus Haaf, 1961a: 48 to Ornatalcides: Pajni
and Dhir, 1987: 33
sexvittatus Faust, 1895a: 243,258 see waltoni

422 shoreaphilus Lyal in Lyal and Curran, 2000: 1828 Malaysia
423 siamodelta (Marshall) Cambodia, Laos,
Alcidodes siamodelta Marshall, 1918a: 154, 155; Vietnam,Malaysia,
Alonso-Zarazaga and Lyal, 1999: 206 Myanmar ,Singapore,
siccus Haaf, 1964b: 163 to Merus: Alonzo- India, Thailand
424 signatus (Boheman) India, Uganda,
Alcides signatus Boheman in Schoenherr, 1836: Congo(?), South
617; Marshall, 1918a: 158; Alonso-Zarazaga and Africa(?)
Lyal, 1999: 206
Alcides trilineatus Faust, 1891: 284; Klima, 1934:
60
425 signifer (Boheman) West Africa
Alcides signifer Boheman in Schoenherr, 1844: 52;

1999:206
serratidens Heller, 1918: 216 to Cylindralcides:
Heller, 1918: 211
426 simplex (Fahraeus) Germany, Indonesia
Alcides simplex Fahraeus, 1871: 256; Marshall,
426a simplex sub sp vernicatus Haaf, 1961a: 34
427 simus (Chevrolat) Congo, Zanzibar,
Alcides simus Chevrolat, 1881b: 91; Marshall, Tanzania
428 similatus (Boheman) Indonesia
Alcides similatus Boheman in Schoenherr, 1836:
Lyal, 1999: 206
429 simmondsi (Haaf) Fiji Islands
Alcidodessimmondsi Haaf, 1963: 154; Alonso-
smaragdinus Roelofs, 1893: 39 to Merus: Heller,
1918: 216 sobrinus Haaf, 1961c: 56 see achteni sub
sp sobrinus
430 solitarius (Haaf) Indonesia, Malaysia
Alcidodes solitarius Haaf, 1964a: 30; Alonso-
431 sordidus (Haaf) Philippines
Alcidodes sordidus Haaf, 1961e: 476; Alonso-
432 sparsus (Boheman) Cape Colony
Alcides sparsus Boheman in Schoenherr, 1844: 53;
1999:206
432a sparsus subsp. decretus Haaf, 1961c: 151 Cape Colony
433 speciosus (Haaf) Indonesia
Alcidodes speciosus Haaf, 1964a: 27; Alonso-

434 speculator (Haaf) Indonesia
Alcidodes speculator Haaf, 1963: 148; Alonso-
435 XV-spilotus (Heller) Philippines
Alcides XV-spilotus Heller, 1923: 9; Marshall,
436 sterryorum Lyal in Lyal and Curran, 2000: 1815 Indonesia
437 stevensi (Marshall) India
Alcidodes stevensi Marshall, 1922: 405; Alonso-
stolzi Haaf, 1961c: 79 see arcuatus sub sp stolzi
438 sturnus (Haaf)
Alcidodes sturnus Haaf, 1963: 152; Alonso-

1939: 570; Haaf, 1963: 152
439 subcuprinus (Schultze) Philippines
Alcides subcuprinus Schultze, 1934: 334; Marshall,
440 subfasciatus (Boheman) Madagascar
Alcides subfasciatus Boheman in Schoenherr,
1844: 67; Marshall, 1939: 570; Alonso-Zarazaga
and Lyal, 1999: 206
441 subhumerosus (Marshall) India
Alcidodes subhumerosus Marshall, 1922: 402;
442 subsellatus (Marshall) Indonesia
Alcidodes subsellatus Marshall, 1922: 402; Alonso-
443 subtilis (Fahraeus) Germany
Alcides subtilis Fahraeus, 1871: 255; Marshall,
444 subvillosus (Fahraeus) Germany
Alcides subvillosus Fahraeus, 1871: 254; Marshall,
445 subvirens (Heller) Indonesia
Alcides subvirens Heller, 1908b: 169; Marshall,
446 sulcatulus (Fabricius) Myanmar, Indonesia
Rhynchaenus sulcatulus Fabricius, 1801: 474;
Boheman in Schoenherr, 1836: 616 (Alcides);
1999:206
sulcatulus Olivier, 1807: 203 see pectoralis
(Boheman)
447 sulcicollis (Hustache) Congo
Alcides sulcicollis Hustache, 1934: 356; Marshall,
448 sumatranus (Heller) Indonesia

Alcides sumatranus Heller, 1933: 102; Marshall,
449 supernus (Haaf) Indonesia
Alcidodes supernus Haaf, 1964a: 63; Alonso-
suratus Haaf, 1961c: 123 see bivinculatus sub sp
suratus
450 suspensus (Pascoe) Sri Lanka
Alcides suspensus Pascoe, 1883: 125; Marshall,
1939: 570; Haaf, 1960: 649; Alonso-Zarazaga and
Lyal, 1999: 206
Sternuchopsis flavofurcatus Voss, 1957: 107; Haaf,
1960:649
451 sycophanta (Thomson) Gabon

Alcides sycophanta Thomson, 1858: 132; Marshall,
taeniatus Heller, 1908b: 156 to
Tuberculomecyslobus: Pajni and Dhir, 1987: 31
tagalicus Schulze, 1918: 273 see dictyodes Heller
taiwanus Haaf, 1964b: 199 see trifidus (Pascoe)
452 takahashii (Kono) Taiwan
Alcides takahashii Kono, 1930: 140; Marshall,
453 tamsi (Marshall) Equatorial Guinea
Alcidodes tamsi Marshall, 1954: 216; Alonso-
454 tanalaensis (Hustache) Madagascar
Alcides tanalaensis Hustache, 1933: 81; Marshall,
455 tapirus (Heller) Indonesia
Alcides tapirus Heller, 1908b: 167; Marshall, 1939:
456 tarsus (Haaf) Indonesia
Alcidodes tarsus Haaf, 1963: 133; Alonso-Zarazaga
and Lyal, 1999: 206
taylori Schultze, 1918: 275 see habena
457 tchadenses (Hustache) Congo
Alcides tchadenses Hustache, 1922: 154; Marshall,
tectus Haaf, 1961c: 91 see haemopterus sub sp
tectus
458 tenellus (Haaf) Philippines
Alcidodes tenellus Haaf, 1962: 152; Alonso-
459 tentus (Haaf) Indonesia
Alcidodes tentus Haaf, 1961b: 387; Alonso-
460 tenuistria (Heller) Papua New Guinea
Alcides tenuistria Heller 1914: 654; Marshall,

Alcides incomptus Marshall, 1915: 516; Klima,
1934: 60
461 ternatensis (Faust) Indonesia, Papua
Alcides ternatensis Heller, 1929: 116; Marshall, New Guinea
australis sub sp ternatensis Faust, 1897b: 181;
Heller, 1929: 116
segnis var. ternatensis Faust, 1897b: 180; 1899: 46;
Heller, 1929: 116
462 termentosus (Fairmaire) Madagascar
Alcides termentosus Fairmaire, 1902a: 368; Haaf,
terraereginae Blackburn, 1900: 141 see bubo

(Fabricius)
463 terrosus (Haaf) Papua New Guinea
Alcidodes terrosus Haaf, 1963: 149; Alonso-
464 tersus (Haaf) Indonesia
Alcidodes tersus Haaf, 1963: 133; Alonso-Zarazaga
and Lyal, 1999: 206
465 tetanicus (Pascoe) Indonesia
Alcides tetanicus Pascoe, 1887: 355; Marshall,
466 tetragrammus (Chevrolat) Zanzibar
Alcides tetragrammus Chevrolat, 1881b: 90;
1999:206
467 texatus (Haaf) Sri Lanka
Alcidodes texatus Haaf, 1960: 659; Alonso-
468 themus Lyal in Lyal and Curran, 2000: 1792 Malaysia,
Philippines
469 theobromae (Marshall) Congo
Alcidodes theobromae Marshall, 1917: 115;
470 thompsoni (Haaf) South Africa
Alcidodes thompsoni Haaf, 1962: 142; Alonso-
471 tomentosus (Fairmaire) Singapore
Alcides tomentosus Fairmaire, 1902a: 365;
1999: 206
tomentosus Hustache, 1925a: 339,345 see
larinoides (Fairmaire)
472 *toyi Lyal in Lyal and Curran, 2000: 1799 Malaysia
473 transversus (Walker) Sri Lanka
Alcides curialis Pascoe, 1883: 124; Klima, 1934:
60
Alcides transversus Walker, 1859: 264; Marshall,
1918a: 157; 1939: 570; Alonso-Zarazaga and Lyal,
1999: 206
474 transversoguttatus (Voss) Sri Lanka
Alcidodes transversoguttatus Voss, 1957: 106;
475 triangulifer (Kirsch) Myanmar,
Alcides triangulifer Kirsch, 1875: 40; Marshall, Indonesia, India
476 tricolor (Heller) Malaysia, India
Alcidodes tricolor Heller, 1940: 92; Alonso-
477 trifasciculatus (Hustache) Congo
Alcides trifasciculatus Hustache, 1924: 73;

1999:206
trifidus Pascoe, 1870b: 459 to Ornatalcides: Pajni
and Dhir, 1987: 33
trifidus sub sp taiwanus Kono, 1930:139 see
trifidus
trigonophorus Marshall, 1922: 394, 396 to
Ornatalcides: Pajni and Dhir, 1987: 30
trilineatus Faust, 1891: 284 see signatus
(Boheman)
478 trilobus (Fabricius) Congo, Gabon, West
Lixus trilobus Fabricius, 1801: 505; Boheman in Africa, Kenya,
Schoenherr, 1844: 60 (Alcides); Marshall, 1939: Tanzania, South
570; Alonso-Zarazaga and Lyal, 1999: 206 Africa
479 trinotatus (Fischer) Uzbekistan
Cionus trinotatus Fischer, 1821: 14; Motschulsky,
1845: 100 (Alcides); Marshall, 1939: 570; Alonso-
480 tristis (Haaf) China
Alcidodes tristis Haaf, 1974: 177; Alonso-Zarazaga
and Lyal, 1999: 206
481 tshibindanus (Hustache) Congo
Alcides tshibindanus Hustache, 1934: 354;
1999:206
tshibindanus Voss, 1962b: 275 to Merus: Voss,
1962b:275
482 tumidus (Haaf) India
Alcidodes tumidus Haaf, 1961a: 28; Alonso-
483 turpis (Heller) Philippines
Alcides turpis Heller, 1918: 235; Marshall, 1939:
tutus Faust, 1894a: 253, 259 to Cylindralcidodes:

484 typicus (Haaf) Malaysia
Alcidodes typicus Haaf, 1964a: 49; Alonso-
485 umbrifer (Haaf) Indonesia
Alcidodes umbrifer Haaf, 1963: 120; Alonso-
umenoi Kono, 1937b: 15 see saundersi (Pascoe)
486 undulatus (Heller) Philippines
Alcides undulatus Heller, 1918: 223; Marshall,
487 vacillans (Heller) Philippines
Alcides vacillans Heller, 1918: 236; Marshall,
488 vadoni (Hustache) Madagascar

Alcidodes vadoni Hustache, 1956: 120; Alonso-
vafellus Faust, 1898: 308 to Cylindralcidodes:
489 vafer (Marshall) Ruanda
Alcidodes vafer Marshall, 1955: 277; Alonso-
490 variabilis (Heller) Philippines
Alcides variabilis Heller, 1918: 234; Marshall,
491 vaticus Lyal in Lyal and Curran, 2000: 1834 Malaysia
492 venosus (Faust) Myanmar
Alcides venosus Faust, 1894a: 239, 257; Marshall,
493 venustus (Haaf) Indonesia
Alcidodes venustus Haaf, 1964a: 62; Alonso-
vernicatus Haaf, 1961a: 34 see simplex sub sp
vernicatus
494 vestitus (Cristofori and Jan) Germany(?)
Alcides vestitus Cristofori and Jan, 1832: 4;
1999:206
495 vestitus (Pascoe) Ghana
Alcides vestitus Pascoe, 1887: 356; Marshall, 1939:
Alcides aspericollis Heller, 1929: 114,115; Haaf,
1963: 136
496 viator (Heller) Philippines,Taiwan
Alcides viator Heller, 1918: 229; Marshall, 1939:
497 vicarius (Haaf) Indonesia
Alcidodes vicarius Haaf, 1961b: 390; Alonso-
vicinus Haaf, 1961c: 140 see denticulatus sub sp

vicinus
498 vinaceomicans (Heller) Philippines
Alcides vinaceomicans Heller, 1923: 9; Marshall,
vinculosus Heller, 1908b: 160, 161 to
499 virgatus (Haaf) Sri Lanka
Alcidodes virgatus Haaf, 1960: 666; Alonso-
vitalisi Marshall, 1922: 394, 409 to Ornatalcides:
500 vitellus (Haat) Africa
Alcidodes vitellus Haaf, 1961c: 67; Alonso-

501 vitiosus (Haat) Indonesia
Alcidodes vitiosus Haaf, 1964a: 73; Alonso-
vividus Haaf, 1961c: 58 see guttulatus (Thomson)
502 vossi (Haat) Tanzania
Alcidodes vossi Haaf, 1962: 141; Alonso-Zarazaga
and Lyal, 1999: 206
wahlbergi Chevrolat, 1881b: 90 see olivaceus
(Gerstacker)
503 *walliorum Lyal in Lyal and Curran, 2000: 1805 Malaysia, Indonesia
waltoni Boheman in Schoenherr, 1844: 58 to
Ornatalcides:
westermanni Boheman in Schoenherr, 1844: 51 to
Ornatalcides: Pajni and Dhir, 1987: 30
504 willcocksi (Pic) Egypt
Alcides willcocksi Pic, 1931a: 103; Marshall, 1939:
Alcides ab. testaceipes Pic, 1931a: 106; Klima,
1934:44
505 wirthi (Haat) Tanzania
Alcidodes wirthi Haaf, 1964a: 292; Alonso-
506 yoshidai (Kono) Taiwan
Alcides yoshidai Kono, 1930: 138; Marshall, 1939:
Alcides ashinagazo Kono, 1930: 142; Klima, 1934:
61
507 zanzibaricus (Hustache) Zanzibar
Alcides zanzibaricus Hustache, 1932:62; Marshall,
508 zonatus (Pascoe) Indonesia
Alcides zonatus Pascoe, 1885: 242; Marshall, 1939:
8 Tuberculomecyslobus Pajni and Dhir, 1987: 31

Type Alcides acutus Faust, 1898: 310
1. acutus (Faust) India
Alcides acutus Faust, 1898: 310; Marshall, 1939:
570 (Alcidodes); Pajni and Dhir, 1987: 31
2. crinalifer (Marshall) India
Alcidodes crinalifer Marshall, 1922: 393; 1939:
570 (Alcidodes); Pajni and Dhir, 1987: 31
3. oberthuri (Pascoe) India
Alcides oberthuri Pascoe, 1887: 357; Marshall,
4. profanus (Haaf) India
Alcidodes profanus Haaf, 1961a: 43; Marshall,
5. taeniatus (Heller) India

Alcides taeniatus Heller, 1908b: 156, 160;
Marshall, 1939: 570 (Alcidodes); Pajni and Dhir,
1987: 31
Acknowledgments
The help and suggestions of Dr. Christopher H.C. Lyal, The Natural History
Museum, London and the financial assistance of Senior Research Fellowship by the
Post Graduate School, Indian Agricultural Research Institute are gratefully
acknowledged.
References
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Tribe Genus Subgenus

Alctdodoplesius Voss
- Brahmalcidodes Pajni and .Dhir
M--1_§ -----c
- Cylindralcides Heller Cylindralcides Reller
Paramecyslobus Voss
- MentsGistel
Exalcidodes Voss
Indomecyslobus Pajni and Dhir
Kismecyslobus Voss
Metallalcides Heller
Pseudalcidodes Voss
Mecysolobini Reitter
-E
- Neomecyslobus P.ajni and Dhir
- Ornatalcides Heller Ornatalcides Heller
Mesalcidodes Voss
Pseudmesalcidodes Pajni and Dhir
r----
Sternuchopsis Heller Alcidodes Marshall
~ Granosalcides Heller
Robustalctdes Heller
Tuberculomecyslobus Pajni and Dhir
Fig. 1 Hierarchical list oftheMecysolobini (after Alonzo-Zarazaga and Lyal, 1999)
Table 1 Species of Mecysolobini over periods, contributors and distribution

Genera Number of species described
Periods Contributing Coleopterists Distribution
§
- -- -- - -
0 0 0 0 0 00
•vJ
0 l() a. l() r-- 0 Q)
§ u
00
~
00 a. :::::: 0
Sl
..c: ~
"''"' ~ "'
<il ·a ·~
--
Q)
.;, ' ' ' '
'til ~ ~ ;;;
~
0 !:J 0
-
000 a. u Q)
~
:E
:1"'
Q)
r-- l() l() r-- g "' ·.:
r-- a. a.
"' "' ~ <il
Q)
00 00
~ ~ ~ ~ ~ g, Q f!l g,
Alcidodoplesius 2 2 1
Brahmalcidodes 1 1 2
Cylindralcides 2 15 5 3 5 5 7 2 5 24
Merus 3 2 12 8 3 2 2 18 17 8 2
Neomecyslobus 7 3 2 2 3 3 11
Ornatalcides 3 9 2 3 2 1 8 16
Sternuchopsis * 5 45 78 217 143 20 128 59 45 33 35 39 169 284 31 172 21
Tubercul_bus 3 5
Total 6 52 85 263 161 30 139 72 46 46 43 45 206 360 32 182 23
* 15 species considered under Alcidodes in the original publications

A Checklist of Weevils of The Tribe Mecysolobini (Coleoptera Curculionidae Molytinae) PDF

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A Checklist of Weevils of The Tribe Mecysolobini (Coleoptera Curculionidae Molytinae) PDF

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A checklist of weevils of the

To link to this article: http://dx.doi.org/10.1080/00305316.2010.10417619

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A CHECKLIST OF WEEVILS OF THE TRIBE MECYSOLOBINI

ABSTRACT: An annotated checklist of the weevil tribe Mecysolobini (Molytinae: Curculionidae:

Key words: Mecysolobini, checklist, genera, subgenera, species, distribution

The generic nomenclature in the molytine tribe Mecysolobini is confused, reflecting

Some workers, particularly in applied fields, still incorrectly employ Alcides

cladistic analysis is done to determine relationships, the most stable approach

1939: 570; Pajni and Dhir, 1987: 32

1939: 570 (Alcidodes); Pajni and Dhir, 1987: 32

1911: 314; Marshall, 1939: 570; Pajni and Dhir,

Mecysolobus dentitibialis Voss, 1957: 103; Alonso-

1987: 30 (Mecyslobus); Alonso-Zarazaga and Lyal,

15. mangyanicus (Schultze) Philippines

Alcides raptorius Boheman in Schoenherr, 1836:

Alcides smaragdinus Roelofs, 1893: 39; Heller,

Alcides westermanni Boheman in Schoenherr,

1. anamalaiensis (Pajni and Dhir) India

47; Marshall, 1939: 570; Pajni and Dhir, 1987: 33

1908a:8; Marshall, 1939: 570; Pajni and Dhir,

and Lyal, 1999: 205

5 adversarius (Schultze) Philippines

Zarazaga and Lyal, 1999: 206

Alcidodes ambatoides Voss, 1962b: 267; Alonso-

Zarazaga and Lyal, 1999: 206

30 arcanus (Haaf) Indonesia

Alcides arenatus Schultze, 1919: 559; Marshall,

40 avidus (Haaf) Cambodia, Laos,

Zarazaga and Lyal, 1999: 206

Zarazaga and Lyal, 1999: 206

Alcides blanchardi Faust, 1895a: 97; Marshall,

brevirostris var p Boheman in Schoenherr, 1844:

65a bubulus var. punctiger Heller

72 caudatus (Hustache) Madagascar

Zarazaga and Lyal, 1999: 206

85 collaris (Pascoe) India

1939: 570; Alonso-Zarazaga and Lyal, 1999: 206

1934: 46; Marshall, 1939: 570

101 crassus (Pascoe) India, Philippines

cylindricus Hustache, 1920b: 10 see hustachei

Alcides decemvittatus Aurivillius, 1892: 220; Cambodia

Alonso-Zarazaga and Lyal, 1999: 206

Alcides detectus Heller, 1925: 231; Marshall, 1939:

1939: 570; Alonso-Zarazaga and Lyal, 1999: 206

Alcidodes diversus Haaf, 1961c: 135; Alonso-

Marshall, 1952: 268 (Alcidodes); Voss, 1962b: 271; Congo

Curculiofabricii Fabricius, 1792: 169; 1801:471

Alcides fallax Boheman in Schoenherr, 1844: 62;

169 figuratus (Schultze) Philippines

Alonso-Zarazaga and Lyal, 1999: 206

Alcidesfunestus Faust, 1894a: 247, 258; Marshall,

geminus Heller, 1908b: 159,161 see comparabilis

195 granicollis (Fairmaire) Madagascar

Marshall, 1939: 570; Alonso-Zarazaga and Lyal,

204 haemopterus (Boheman) Ethiopia, Germany,

1939: 570; Alonso-Zarazaga and Lyal, 1999: 206

Alcides humerosus Harold, 1880: 265; Marshall, Uganda

229 inducens (Heller) Indonesia

1939: 570; Alonso-Zarazaga and Lyal, 1999: 206

kalinganus Schultze, 1923: 604 to Ornatalcides: