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Chapter 30
Homeostasis
Homeostasis refers to maintaining internal stability within an organism and returning to a particular stable state after a fluctuation.
Homeostasis
Homeostasis
Systems within an organism function in an integrated way to maintain a constant internal environment around a setpoint.
Small deviations in pH, temperature, osmotic pressure, glucose levels, & oxygen levels activate physiological mechanisms to return that variable to its setpoint.
Negative feedback
Osmosis
Cells require a balance between osmotic gain and loss of water. Water uptake and loss are balanced by various mechanisms of osmoregulation in different environments.
Osmosis
If two solutions that are separated by a membrane differ in their osmolarity, water will cross the membrane to bring the osmolarity into balance (equal solute concentrations on both sides).
Osmotic Challenges
Osmoconformers, which are only marine animals, are isoosmotic with their surroundings and do not regulate their osmolarity. Osmoregulators expend energy to control water uptake and loss in a hyperosmotic or hypoosmotic environment.
Osmotic Regulation
Most marine invertebrates are osmotic conformers their bodies have the same salt concentration as the seawater.
The sea is highly stable, so most marine invertebrates are not exposed to osmotic fluctuations. These organisms are restricted to a narrow range of salinity stenohaline.
Osmotic Regulation
Conditions along the coasts and in estuaries are often more variable than the open ocean.
Animals must be able to handle large, often abrupt changes in salinity. Euryhaline animals can survive a wide range of salinity changes by using osmotic regulation.
Osmotic Regulation
The problem of dilution is solved by pumping out the excess water as dilute urine. The problem of salt loss is compensated for by salt secreting cells in the gills the actively remove ions from the water and move them into the blood.
Requires energy.
Freshwater animals face an even more extreme osmotic difference than those that inhabit estuaries.
Freshwater fishes have skin covered with scales and mucous to keep excess water out. Water that enters the body is pumped out by the kidney as very dilute urine. Salt absorbing cells in the gills transport salt ions into the blood.
Invertebrates and amphibians also solve these problems in a similar way. Amphibians actively absorb salt from the water through their skin.
Maintain salt concentration at 1/3 that of seawater. Marine fishes drink seawater to replace water lost by diffusion. Excess salt is carried to the gills where salt-secreting cells transport it out to the sea.
Terrestrial animals lose water by evaporation from respiratory and body surfaces, excretion (urine), and elimination (feces). Water is replaced by drinking water, water in food, and retaining metabolic water.
Fishes can excrete ammonia directly because there is plenty of water to wash it away.
Marine birds and turtles have a salt gland capable of excreting highly concentrated salt solution.
Excretory Processes
Most excretory systems produce urine by refining a filtrate derived from body fluids (blood, hemolymph, or coelomic fluid).
Excretory Processes
The simplest arrangement is the protonephridium of acoelomates and some pseudocoelomates. Fluid enters through flame cells, moves through the tubules, water and metabolites are recovered and wastes are excreted through pores that open along the body surface.
The metanephridium is an open system found in annelids, molluscs, and some smaller phyla.
The metanephridium is surrounded by blood vessels that assist in reclaiming water and valuable solutes.
No open nephrostomes, hydrostatic pressure of the blood forms an ultrafiltrate in the end sac. In the tubule, selective resorption of some salts and active secretion of others occurs.
Insects and spiders have Malpighian tubules that are closed and lack an arterial supply. Salts (especially potassium) are secreted into the tubules from the hemolymph (blood).
Water & other solutes (including uric acid) follow. Water & potassium are reabsorbed. Uric acid is expelled in feces.
Vertebrate Kidneys
Kidneys, the excretory organs of vertebrates, function in both excretion and osmoregulation.
Vertebrate Kidneys
Nephrons and associated blood vessels are the functional unit of the mammalian kidney. The mammalian excretory system centers on paired kidneys which are also the principal site of water balance and salt regulation.
Vertebrate Kidneys
Each kidney is supplied with blood by a renal artery and drained by a renal vein.
Vertebrate Kidneys
Urine exits each kidney through a duct called the ureter. Both ureters drain into a common urinary bladder.
The nephron, the functional unit of the vertebrate kidney consists of a single long tubule and a ball of capillaries called the glomerulus.
Filtration occurs as blood pressure forces fluid from the blood in the glomerulus into the lumen of Bowmans capsule.
From Bowmans capsule, the filtrate passes through three regions of the nephron:
Filtrate becomes urine as it flows through the mammalian nephron and collecting duct.
The composition of the filtrate is modified through tubular reabsorption and secretion. Changes in the total osmotic concentration of urine through regulation of water excretion.
Secretion and reabsorption in the proximal tubule substantially alter the volume and composition of filtrate. Reabsorption of water continues as the filtrate moves into the descending limb of the loop of Henle.
As filtrate travels through the ascending limb of the loop of Henle salt diffuses out of the permeable tubule into the interstitial fluid. The distal tubule plays a key role in regulating the K+ and NaCl concentration of body fluids. The collecting duct carries the filtrate through the medulla to the renal pelvis and reabsorbs NaCl.
Conserving Water
The mammalian kidneys ability to conserve water is a key terrestrial adaptation. The mammalian kidney can produce urine much more concentrated than body fluids, thus conserving water.
In a mammalian kidney, the cooperative action and precise arrangement of the loops of Henle and the collecting ducts are largely responsible for the osmotic gradient that concentrates the urine.
The collecting duct, permeable to water but not salt conducts the filtrate through the kidneys osmolarity gradient, and more water exits the filtrate by osmosis.
The osmolarity of the urine is regulated by nervous and hormonal control of water and salt reabsorption in the kidneys.
Antidiuretic hormone (ADH) increases water reabsorption in the distal tubules and collecting ducts of the kidney.
Temperature Regulation
Animals must keep their bodies within a range of temperatures that allows for normal cell function. Each enzyme has an optimum temperature.
Too low and metabolism slows. Too high and metabolic reactions become unbalanced. Enzymes may be destroyed.
Temperature Regulation
Poikilothermic animals body temperatures fluctuate with environmental temperatures. Homeothermic animals body temperatures are constant.
Temperature Regulation
All animals produce heat from cellular metabolism, but in most this heat is lost quickly.
Ectotherms lose metabolic heat quickly, so body temperature is determined by the environment.
Endotherms retain metabolic heat and can maintain a constant internal body temperature.
Most ectotherms can also adjust their metabolic rates to the environmental temperature.
Constant temperature in endotherms is maintained by a delicate balance between heat production and heat loss.
Heat is produced by the animals metabolism. Producing heat requires energy supplied by food.
If an animal is too cool, it can generate heat by increasing muscular activity (exercise or shivering). Heat is retained through insulation. If an animal is too warm it decreases heat production and increases heat loss.
Adaptations to derive water from metabolism and produce concentrated urine & dry feces.
Glossy, pallid color reflects sunlight. Fat tissue is concentrated in a hump, rather than being evenly distributed in an insulating layer. Sweating and panting are ways of dumping heat.
In cold environments, mammals reduce heat loss by having a thick insulating layer of fat, fur, or both. Heat production is increased. Extremities are allowed to cool.
Many avoid direct exposure to the cold by living in tunnels under the snow.
Subnivean environment. This is where food is located.
Adaptive Hypothermia
Adaptive Hypothermia
Some very small mammals & birds (bats or hummingbirds) maintain high body temperatures when active, but allow temperatures to drop when sleeping.
Daily torpor
Adaptive Hypothermia
Hibernation is a way to solve the problem of low temperatures and the scarcity of food.
Metabolism & body slows to a fraction of normal. Body temperature decreases. Shivering helps increase temperatures when they are waking up.
Adaptive Hypothermia
Other mammals, such as bears, badgers, raccoons and opossums enter a state of prolonged sleep, but body temperature does not decrease.
Adaptive Hypothermia